View source for 2021 in paleoentomology

{{Short description|none}} <!-- "none" is preferred when the title is sufficiently descriptive; see [[WP:SDNONE]] -->
{{Year nav topic20 |2021|paleoentomology|paleontology|paleobotany|arthropod paleontology|paleoichthyology|paleomalacology|paleomammalogy|reptile paleontology|archosaur paleontology}}
{{see|2021 in arthropod paleontology|2021 in paleontology}}
This [[paleoentomology]] list records new  [[fossil]] [[insect]] [[taxa]] that were to be [[binomial nomenclature|described]] during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.
{{TOC right}}

==Clade Amphiesmenoptera==
===Lepidoptera===
====New Lepidopteran taxa====
{| class="wikitable sortable" align="center" width="80%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Acanthocorona]]''<ref name=Acanthocorona>{{Cite journal|last1=Mey |first1=W. |last2=Léger |first2=T. |last3=Lien |first3=V. V. |title=New taxa of extant and fossil primitive moths in South-East Asia and their biogeographic significance (Lepidoptera, Micropterigidae, Agathiphagidae, Lophocoronidae) |year=2021 |journal=Nota Lepidopterologica |volume=44 |pages=29–56 |doi=10.3897/nl.44.52350 |doi-access=free }}</ref>
|
Gen. et 7 sp. nov
|
Valid
|
Mey, Léger & Lien
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Lophocoronidae]] moth.<br/> The type species is ''A. skalskii''<br/>The genus also includes ''A. bowangi'', ''A. kuranishii'', ''A. muelleri'', ''A. sattleri'', ''A. spinifera'', & ''A. wichardi''.
|
[[File:Acanthocorona skalskii holotype img1.jpg|thumb|center|upright|''[[Acanthocorona skalskii]]'']]
|-
|
''[[Agathiphagama]]''<ref name=Acanthocorona />
|
Gen. et sp. nov
|
Valid
|
Mey, Léger & Lien
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Agathiphagidae|agathiphagid]] [[aglossata]]n moth.<br/> The type species is ''A. perdita''.
|
[[File:Agathiphagama perdita holotype wetted img1.jpg|thumb|center|upright|''[[Agathiphagama perdita]]'']]
|-
|
''[[Phyllocnistis cretacea]]''<ref name=ZittelianaFischer>{{Cite journal|last=Fischer |first=T. C. |title=In search for the unlikely: Leaf-mining caterpillars (Gracillariidae, Lepidoptera) from Upper Cretaceous and Eocene ambers |year=2021 |journal=Zitteliana |volume=95 |pages=135–145 |doi=10.3897/zitteliana.95.63317 |s2cid=245086588 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Fischer
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''[[Phyllocnistis]]''
|
|-
|
''[[Phyllonorycter inopinata]]''<ref name=ZittelianaFischer />
|
Sp. nov
|
Valid
|
Fischer
|
Lutetian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A species of ''[[Phyllonorycter]]''
|
|-
|}

====Lepidopteran research====
* The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).<ref>{{Cite journal|last1=Haug |first1=J. T. |last2=Haug |first2=C. |title=A 100 million-year-old armoured caterpillar supports the early diversification of moths and butterflies |year=2021 |journal=Gondwana Research |volume=93 |pages=101–105 |doi=10.1016/j.gr.2021.01.009 |bibcode=2021GondR..93..101H |s2cid=233548334 }}</ref>
* Fossil leaf mine produced by a [[Lyonetiidae|lyonetiid]]-like [[Leaf miner|leaf-mining]] moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous ([[Campanian]]) [[Kaiparowits Formation]] ([[Utah]], United States) by Maccracken ''et al.'' (2021), who name a new [[ichnotaxon]] ''[[Leucopteropsa]] spiralae''.<ref>{{Cite journal|last1=Maccracken |first1=S. A. |last2=Sohn |first2=J.-C. |last3=Miller |first3=I. M. |last4=Labandeira |first4=C. C. |title=A new Late Cretaceous leaf mine ''Leucopteropsa spiralae'' gen. et sp. nov. (Lepidoptera: Lyonetiidae) represents the first confirmed fossil evidence of the Cemiostominae |year=2021 |journal=Journal of Systematic Palaeontology |volume=19 |issue=2 |pages=131–144 |doi=10.1080/14772019.2021.1881177 |s2cid=233194385 }}</ref>

===Trichopterans===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Archaeopolycentra achupomotet]]''<ref name=FEE444>{{Cite journal|last1=Melnitsky |first1=S. I. |last2=Ivanov |first2=V. D. |title=Two new species of the genus ''Archaeopolycentra'' (Trichoptera: Polycentropodidae) from Cretaceous Taimyr amber |year=2021 |journal=Far Eastern Entomologist |volume=444 |pages=1–7 |doi=10.25221/fee.444.1| issn=1026-051X  |s2cid=244880131 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Melnitsky & Ivanov
|
Late Cretaceous ([[Santonian]])
|
Kheta Formation<br>(Taymyr amber)
|
{{Flag|Russia}}<br>({{Flag|Krasnoyarsk Krai}})
|
A member of the family [[Polycentropodidae]].
|
|-
|
''[[Archaeopolycentra turvalsia]]''<ref name=FEE444 />
|
Sp. nov
|
Valid
|
Melnitsky & Ivanov
|
Late Cretaceous (Santonian)
|
Kheta Formation<br>(Taymyr amber)
|
{{Flag|Russia}}<br>({{Flag|Krasnoyarsk Krai}})
|
A member of the family Polycentropodidae.
|
|-
|
''[[Cernotina fossilinova]]''<ref>{{Cite journal|last1=Wichard |first1=W. |last2=Neumann |first2=C. |title=The polycentropodid genus Cernotina (Insecta, Trichoptera) in Miocene Dominican amber |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=129–133 |doi=10.5194/fr-24-129-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Wichard & Neumann
|
Probably early Miocene
|
[[Dominican amber]]
|
{{Flag|Dominican Republic}}
|
A [[caddisfly]], a species of ''[[Cernotina]]''.
|
|-
|
''[[Electrotrichia|Electrotrichia rasnitsyni]]''<ref>{{Cite journal|last1=Melnitsky |first1=S. I. |last2=Ivanov |first2=V. D. |last3=Perkovsky |first3=E. E. |title=A new species of the fossil genus ''Electrotrichia'' (Insecta: Trichoptera: Hydroptilidae) from Rovno amber (Zhytomyr region, Olevsk locality) |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=421–424 |doi=10.11646/palaeoentomology.4.5.4 |s2cid=240542805 }}</ref>
|
Sp. nov
|
Valid
|
Melnitsky, Ivanov & Perkovsky
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family [[Hydroptilidae]].
|
|-
|
''[[Folindusia aenigmatosa]]''<ref name=PJ555>{{Cite journal|last1=Sukacheva |first1=I. D. |last2=Vasilenko |first2=D. V. |title=Caddisflies (Insecta, Trichoptera) from the upper Mesozoic Chernovskie Kopi locality (Transbaikalia): part 2. Larval cases |year=2021 |journal=Paleontological Journal |volume=55 |issue=5 |pages=545–551 |doi=10.1134/S0031030121050154 |s2cid=238640223 |url=https://elibrary.ru/item.asp?id=46428492 }}</ref>
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Folindusia expansa]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Folindusia inconspicua]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Folindusia inventa]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Folindusia robusta]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Folindusia tortilis]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Holocentropus tutkaktut]]''<ref>{{Cite journal|last1=Melnitsky |first1=S. I. |last2=Ivanov |first2=V. D. |last3=Perkovsky |first3=E. E. |title=A new species of the genus ''Holocentropus'' (Trichoptera: Polycentropodidae) from Rovno amber |year=2021 |journal=Zoosystematica Rossica |volume=30 |issue=2 |pages=298–302 |doi=10.31610/zsr/2021.30.2.298 |s2cid=245185514 }}</ref>
|
Sp. nov
|
Valid
|
Melnitsky, Ivanov & Perkovsky
|
Eocene ([[Priabonian]])
|
Rovno amber
|
{{Flag|Ukraine}}
|
A species of ''[[Holocentropus]]''.
|
|-
|
''[[Palaeopsilotreta|Palaeopsilotreta kachini]]''<ref name=Palaeoentomology44Odontoceridae>{{Cite journal|last1=Wichard |first1=W. |last2=Müller |first2=P. |last3=Xu |first3=C.-P. |title=The odontocerid genera ''Psilotreta'' and ''Palaeopsilotreta'' (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=331–338 |doi=10.11646/palaeoentomology.4.4.8 |s2cid=239650542 }}</ref>
|
Sp. nov
|
Valid
|
Wichard, Müller & Xu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Odontoceridae]].
|
|-
|
''Palaeopsilotreta succini''<ref name=Palaeoentomology44Odontoceridae />
|
Sp. nov
|
Valid
|
Wichard, Müller & Xu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Odontoceridae.
|
|-
|
''[[Plectrocnemia kirmikhia]]''<ref>{{Cite journal|last1=Melnitsky |first1=S. I. |last2=Ivanov |first2=V. D. |last3=Perkovsky |first3=E. E. |year=2021 |title=A new species of ''Plectrocnemia'' (Trichoptera: Polycentropodidae) from Rovno amber |journal=Zootaxa |volume=5006 |issue=1 |pages=106–109 |doi=10.11646/zootaxa.5006.1.14 |pmid=34810582 |s2cid=239644442 }}</ref>
|
Sp. nov
|
Valid
|
Melnitsky, Ivanov & Perkovsky
|
Eocene ([[Priabonian]])
|
Rovno amber
|
{{Flag|Ukraine}}
|
A species of ''[[Plectrocnemia]]''.
|
|-
|
''[[Psilotreta fossilis]]''<ref name=Palaeoentomology44Odontoceridae />
|
Sp. nov
|
Valid
|
Wichard, Müller & Xu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''[[Psilotreta]]''.
|
|-
|
''[[Purbimodus|Purbimodus khramovi]]''<ref>{{Cite journal |last1=Sukatsheva |first1=I. D. |last2=Aristov |first2=D. S. |title=New data on Cretaceous caddisflies family Vitimotauliidae (Insecta, Trichoptera) from Buryatia (Khasurty locality) |year=2021 |journal=Paleontological Journal |volume=55 |issue=4 |pages=391–395 |doi=10.1134/S0031030121330017 |s2cid=237376766 |url=https://elibrary.ru/item.asp?id=46227215 }}</ref>
|
Sp. nov
|
Valid
|
Sukatsheva & Aristov
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A member of the family [[Vitimotauliidae]].
|
|-
|
''[[Terrindusia nova]]''<ref name=PJ555 />
|
Sp. nov
|
Valid
|
Sukacheva & Vasilenko
|
Early Cretaceous
|
|
{{Flag|Russia}}
|
A larval case of a caddisfly.
|
|-
|
''[[Wormaldia denticulata]]''<ref name=WangetalCRWormaldia>{{Cite journal|last1=Wang |first1=J. |last2=Zhang |first2=W. |last3=Shih |first3=C. |last4=Ren |first4=D. |title=Two new mid-Cretaceous caddisflies (Trichoptera, Philopotamidae) from amber of Northern Myanmar |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104891 |doi=10.1016/j.cretres.2021.104891 |bibcode=2021CrRes.12604891W }}</ref>
|
Sp. nov
|
In press
|
Wang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A caddisfly, a species of ''[[Wormaldia]]''.
|
|-
|
''[[Wormaldia diplobifurca]]''<ref name=WangetalCRWormaldia />
|
Sp. nov
|
In press
|
Wang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A caddisfly, a species of ''Wormaldia''.
|
|-
|}

==Clade Antliophora==
===Dipterans===
====New Dipteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Acartophthalmites|Acartophthalmites crassipes]]''<ref name=Clusiomitidae>{{Cite journal|last1=Roháček |first1=J. |last2=Hoffeins |first2=C. |title=Clusiomitidae, a new family of Eocene fossil Acalyptratae, with revision of ''Acartophthalmites'' Hennig and ''Clusiomites'' Gen. Nov. (Diptera) † |year=2021 |journal=Insects |volume=12 |issue=12 |pages=Article 1123 |doi=10.3390/insects12121123 |pmid=34940210 |pmc=8705662 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Roháček & Hoffeins
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of [[Opomyzoidea]] belonging to the new family [[Clusiomitidae]].
|
|-
|
''Acartophthalmites luridus''<ref name=Clusiomitidae />
|
Sp. nov
|
Valid
|
Roháček & Hoffeins
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of Opomyzoidea belonging to the new family Clusiomitidae.
|
|-
|
''Acartophthalmites rugosus''<ref name=Clusiomitidae />
|
Sp. nov
|
Valid
|
Roháček & Hoffeins
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of Opomyzoidea belonging to the new family Clusiomitidae.
|
|-
|
''[[Aleksiana]]''<ref>{{Cite journal|last1=Krzemiński |first1=W. |last2=Kopeć |first2=K. |last3=Soszyńska-Maj |first3=A. |last4=Skibińska |first4=K. |title=''Aleksiana rasnitsyni'' gen. et sp. nov. (Diptera, Limoniidae) to honour Alexandr Rasnitsyn |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=436–440 |doi=10.11646/palaeoentomology.4.5.8 |s2cid=240522901 }}</ref>
|
Gen. et sp. et comb. nov
|
Valid
|
Krzemiński ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the [[crane fly]] family [[Limoniidae]]. The type species is ''A. rasnitsyni''; genus also includes ''"[[Antocha]]" lapra'' Podenas & Poinar (2009).
|
|-
|
''[[Allarete patriciae]]''<ref name=Palaeoentomology46Nel>{{Cite journal|last=Nel |first=A. |title=New Cecidomyiidae from the Lowermost Eocene French amber (Diptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=620–628 |doi=10.11646/palaeoentomology.4.6.11 |s2cid=245179601 }}</ref>
|
Sp. nov
|
Valid
|
Nel
|
Early Eocene
|
Oise amber
|
{{Flag|France}}
|
A member of the family [[Cecidomyiidae]].
|
|-
|
''[[Allodia paleoafricana]]''<ref>{{Cite journal|last1=Bouju |first1=V. |last2=Rosse-Guillevic |first2=S. |last3=Griffon |first3=M. |last4=Bojarski |first4=B. |last5=Szwedo |first5=J. |last6=Perrichot |first6=V. |title=The genus ''Allodia'' (Diptera: Mycetophilidae) in Miocene Ethiopian amber |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=339–346 |doi=10.5194/fr-24-339-2021 |doi-access=free |url=https://fr.copernicus.org/articles/24/339/2021/ }}</ref>
|
Sp. nov
|
Valid
|
Bouju ''et al.''
|
Early [[Miocene]]
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A species of ''[[Allodia]]''.
|
|-
|
''[[Apocephalus chiapanecus]]''<ref name=PalaeoworldPhoridae>{{Cite journal|last1=Solórzano-Kraemer |first1=M. M. |last2=Bourdeau |first2=C. |last3=Rosse-Guillevic |first3=S. |last4=Hammel |first4=J. U. |last5=Xu |first5=C. |last6=Brown |first6=B. V. |title=Parasitic hump-backed flies (Diptera: Phoridae) from Middle Miocene ambers |year=2021 |journal=Palaeoworld |volume=in press |doi=10.1016/j.palwor.2021.11.002 |s2cid=244126413 |url=https://hal-insu.archives-ouvertes.fr/insu-03431983/file/Solorzano-2021.pdf }}</ref>
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
Mexican amber
|
{{Flag|Mexico}}
|
A species of ''[[Apocephalus]]''.
|
|-
|
''[[Apocephalus dominicanus]]''<ref name=PalaeoworldPhoridae />
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A species of ''Apocephalus''.
|
|-
|
''[[Apocephalus miocenus]]''<ref name=PalaeoworldPhoridae />
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
|
|
A species of ''Apocephalus''.
|
|-
|
''[[Araripogon|Araripogon pulchrus]]''<ref>{{Cite journal|last1=Lamas |first1=C. J. E. |last2=Sampronha |first2=S. |last3=Ribeiro |first3=G. C. |title=A new robber fly from the Lower Cretaceous (Aptian) Crato Formation of NE Brazil (Insecta: Diptera: Asilidae) |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105114 |doi=10.1016/j.cretres.2021.105114 }}</ref>
|
Sp. nov
|
In press
|
Lamas, Sampronha & Ribeiro
|
Early Cretaceous (Aptian)
|
Crato Formation
|
{{Flag|Brazil}}
|
A member of the family [[Asilidae]].
|
|-
|
''[[Bashkonia]]''<ref>{{Cite journal|last=Lukashevich |first=E. D. |year=2021 |title=The oldest Diptera (Insecta) from the Upper Buntsandstein (early Middle Triassic) of Europe |journal=Zootaxa |volume=5067 |issue=1 |pages=135–143 |doi=10.11646/zootaxa.5067.1.10 |pmid=34810754 |s2cid=243901965 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Lukashevich
|
[[Middle Triassic]]
|
[[Röt Formation]]
|
{{Flag|Germany}}
|
A member of the family [[Nadipteridae]]. Genus includes new species ''B. franconica''.
|
|-
|
''[[Bibio castaneipennis]]''<ref>{{Cite journal|last1=Skartveit |first1=J. |last2=Wedmann |first2=S. |title=A revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany |year=2021 |journal=Zootaxa |volume=4909 |issue=1 |pages=zootaxa.4909.1.1 |doi=10.11646/zootaxa.4909.1.1 |pmid=33756592 |s2cid=232338865 }}</ref>
|
Sp. nov
|
Valid
|
Skartveit & Wedmann
|
[[Oligocene]]
|
|
{{Flag|Germany}}
|
A species of ''[[Bibio (fly)|Bibio]]''.
|
|-
|
''[[Bibio succineus]]''<ref>{{Cite journal|last=Skartveit |first=J. |title=A new fossil species of the genus ''Bibio'', with an update on bibionid flies from Baltic and Rovno amber (Diptera, Bibionidae) |year=2021 |journal=Deutsche Entomologische Zeitschrift |volume=68 |issue=1 |pages=81–99 |doi=10.3897/dez.68.60611 |doi-access=free |hdl=11250/2731857 |hdl-access=free }}</ref>
|
Sp. nov
|
Valid
|
Skartveit
|
Eocene
|
[[Baltic amber]]
|
Europe (Baltic Sea region)
|
A species of ''Bibio''.
|
|-
|
''[[Burmahesperinus]]''<ref>{{Cite journal|last1=Ševčík |first1=J. |last2=Skartveit |first2=J. |last3=Krzemiński |first3=W. |last4=Skibińska |first4=K. |title=A Peculiar New Genus of Bibionomorpha (Diptera) with Brachycera-Like Modification of Antennae from Mid-Cretaceous Amber of Myanmar |year=2021 |journal=Insects |volume=12 |issue=4 |pages=Article 364 |doi=10.3390/insects12040364 |pmid=33923888 |pmc=8073775 |doi-access=free }}</ref>
|
Gen. et 3 sp. nov
|
Valid
|
Ševčík ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Bibionidae]]. The type species is ''B. antennatus''; genus also includes ''B. conicus'' and ''B. pedicellatus''.
|
|-
|
''[[Burmasymmerus]]''<ref name=Burmasymmerus>{{Cite journal|last1=Ševčík |first1=J. |last2=Hippa |first2=H. |last3=Burdíková |first3=N. |title=Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera |year=2021 |journal=Insects |volume=13 |issue=1 |pages=Article 19 |doi=10.3390/insects13010019 |pmid=35055862 |pmc=8779109 |doi-access=free }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Ševčík, Hippa & Burdíková
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Ditomyiidae]]. The type species is ''B. korneliae''; genus also includes ''B. wieslawi''.
|
|-
|
''[[Burmatricha]]''<ref name=Burmasymmerus />
|
Gen. et sp. nov
|
Valid
|
Ševčík, Hippa & Burdíková
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Sciaroidea]] of uncertain phylogenetic placement. The type species is ''B. mesozoica''.
|
|-
|
''[[Cascomixticus]]''<ref>{{Cite journal|last1=Poinar |first1=G. |last2=Vega |first2=F. E. |title=A new genus of Apsilocephalidae (Diptera) in Mid-Cretaceous Burmese amber |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=1 |pages=183–190 |doi=10.37819/biosis.002.01.0091 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar & Vega
|
Cretaceous (Albian/Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Apsilocephalidae]]. The type species is ''C. tubuliferous''.
|
|-
|
''[[Clusiomites]]''<ref name=Clusiomitidae />
|
Fam. et gen. et comb. et sp. nov
|
Valid
|
Roháček & Hoffeins
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is ''"[[Acartophthalmites]]" clusioides'' Roháček (2016); genus also includes new species ''C. ornatus''.
|
|-
|
''[[Crenoptychoptera|Crenoptychoptera gaoi]]''<ref>{{Cite journal|last1=Liu |first1=Y.M. |last2=Huang |first2=D.Y. |title=A new species of ''Crenoptychoptera'' Kalugina, 1985 (Diptera: Ptychopteridae) from the Middle–Late Jurassic of Jiyuan Basin, China |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=27–29 |doi=10.11646/palaeoentomology.4.1.5 |s2cid=233927262 }}</ref>
|
Sp. nov
|
Valid
|
Liu & Huang
|
Middle–Late Jurassic
|
Jiyuan Basin
|
{{Flag|China}}
|
A member of the family [[Ptychopteridae]].
|
|-
|
''[[Cretobibio|Cretobibio rasnitsyni]]''<ref>{{Cite journal|last1=Lukashevich |first1=E. D. |last2=Amorim |first2=D. de S. |last3=Ribeiro |first3=G. C. |title=A Gondwanan record of the extinct genus ''Cretobibio'' (Diptera: Bibionidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=468–474 |doi=10.11646/palaeoentomology.4.5.13 |s2cid=244210389 }}</ref>
|
Sp. nov
|
Valid
|
Lukashevich, Amorim & Ribeiro
|
Early Cretaceous ([[Aptian]])
|
Crato Formation
|
{{Flag|Brazil}}
|
A member of the family [[Bibionidae]].
|
|-
|
''[[Cretolimonia|Cretolimonia dayana]]''<ref name=SRCretolimonia>{{Cite journal|last1=Kopeć |first1=K. |last2=Soszyńska-Maj |first2=A. |last3=Kania-Kłosok |first3=I. |last4=Coram |first4=R. A. |last5=Krzemiński |first5=W. |title=Morphology of the oldest fossil subfamily of Limoniidae (Diptera, Architipulinae) in the light of exceptionally preserved Mesozoic material |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 24137 |doi=10.1038/s41598-021-03350-4 |pmid=34921169 |pmc=8683464 |bibcode=2021NatSR..1124137K }}</ref>
|
Sp. nov
|
|
Kopeć ''in'' Kopeć ''et al.''
|
Jurassic/Cretaceous boundary
|
Glushkovo Formation
|
{{Flag|Russia}}<br>({{Flag|Zabaykalsky Krai}})
|
A member of the subfamily [[Architipulinae]].
|
|-
|
''Cretolimonia lukashevichae''<ref name=SRCretolimonia />
|
Sp. nov
|
|
Kopeć & Krzemiński ''in'' Kopeć ''et al.''
|
Jurassic/Cretaceous boundary
|
Dain Formation
|
{{Flag|Russia}}<br>({{Flag|Zabaykalsky Krai}})
|
A member of the subfamily Architipulinae.
|
|-
|
''Cretolimonia mikolajczyki''<ref name=SRCretolimonia />
|
Sp. nov
|
|
Kopeć, Krzemiński, Soszyńska-Maj ''in'' Kopeć ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the subfamily Architipulinae.
|
|-
|
''Cretolimonia pseudojurassica''<ref name=SRCretolimonia />
|
Sp. nov
|
|
Krzemiński ''in'' Kopeć ''et al.''
|
Jurassic/Cretaceous boundary
|
Dain Formation
|
{{Flag|Russia}}<br>({{Flag|Zabaykalsky Krai}})
|
A member of the subfamily Architipulinae.
|
|-
|
''[[Cretothereva]]''<ref>{{Cite journal|last1=Carmo |first1=D. d.D. |last2=Lamas |first2=C. J. E. |last3=Ribeiro |first3=G. C. |title=The oldest fossil Stiletto fly: a new genus and species from the Lower Cretaceous Crato Formation of Brazil (Diptera: Therevidae) |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105039 |doi=10.1016/j.cretres.2021.105039 |s2cid=244212150 }}</ref>
|
Gen. et sp. nov
|
In press
|
Carmo, Lamas & Ribeiro
|
[[Early Cretaceous]]
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A member of the family [[Therevidae]]. Genus includes new species ''C. antiqua''.
|
|-
|
''[[Cretpenthetria]]''<ref>{{Cite journal|last1=Li |first1=L. |last2=Zhang |first2=Q. |last3=Xiao |first3=C. |year=2021 |title=A new genus and species of Bibionidae (Diptera: Bibionoidea) from mid-Cretaceous Kachin amber |journal=Acta Palaeontologica Sinica |volume=60 |issue=2 |pages=243–252 |doi=10.19800/j.cnki.aps.2021018 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210204 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li, Zhang & Xiao
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Bibionidae]]. Genus includes new species ''C. burmensis''.
|
|-
|
''[[Culex ekaterinae]]''<ref>{{Cite journal|last1=Giłka |first1=W. |last2=Harbach |first2=R. E. |last3=Perkovsky |first3=E. E. |year=2021 |title=Mosquitoes (Diptera: Culicidae) in Eocene amber from the Rovno region, Ukraine |journal=Zootaxa |volume=5016 |issue=2 |pages=257–270 |doi=10.11646/zootaxa.5016.2.6 |pmid=34810451 |s2cid=238838914 }}</ref>
|
Sp. nov
|
Valid
|
Giłka, Harbach & Perkovsky
|
Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A mosquito, a species of ''[[Culex]]''.
|
|-
|
''[[Culicoides paleopestis]]''<ref name=HBAnglesea>{{Cite journal|last1=Peñalver |first1=E. |last2=Arillo Aranda |first2=A. |last3=Szadziewski |first3=R. |last4=Stilwell |first4=J. D. |year=2021 |title=Biting midges (Diptera: Ceratopogonidae) from the middle Eocene Anglesea amber (Australia) originated in a subpolar greenhouse earth |journal=Historical Biology |volume=34 |issue=3 |pages=425–435 |doi=10.1080/08912963.2021.1924700 |s2cid=236275915 }}</ref>
|
Sp. nov
|
In press
|
Peñalver ''et al.''
|
[[Eocene]]
|
Anglesea amber
|
{{Flag|Australia}}
|
A species of ''[[Culicoides]]''.
|
|-
|
''[[Cyttaromyia freiwaldi]]''<ref name=SRCyttaromyia>{{Cite journal|last1=Kania-Kłosok|first1=I. |last2=Nel |first2=A. |last3=Szwedo |first3=J. |last4=Jordan-Stasiło |first4=W. |last5=Krzemiński |first5=W. |title=Phylogeny and biogeography of the enigmatic ghost lineage Cylindrotomidae (Diptera, Nematocera) |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 13916 |doi=10.1038/s41598-021-91719-w |pmid=34230501 |pmc=8260593 |bibcode=2021NatSR..1113916K }}</ref>
|
Sp. nov
|
Valid 
|
Kania-Kłosok ''et al.''
|
Eocene
|
[[Green River Formation]]
|
{{Flag|United States}}
|
A member of the family [[Cylindrotomidae]].
|
|-
|
''[[Cyttaromyia gelhausi]]''<ref name=SRCyttaromyia />
|
Sp. nov
|
Valid 
|
Kania-Kłosok ''et al.''
|
Eocene
|
Green River Formation
|
{{Flag|United States}}
|
A member of the family Cylindrotomidae.
|
|-
|
''[[Decessia]]''<ref>{{Cite journal|last1=Krzemiński |first1=W. |last2=Kania-Kłosok |first2=I. |last3=Krzemińska |first3=E. |last4=Ševčík |first4=J. |last5=Soszyńska-Maj |first5=A. |title=Fossils Shed a New Light on the Diversity and Disparity of the Family Limoniidae (Diptera, Nematocera) |year=2021 |journal=Insects |volume=12 |issue=3 |pages=Article 206 |doi=10.3390/insects12030206 |pmid=33804356 |pmc=8000698 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Krzemiński ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the [[crane fly]] family [[Limoniidae]]. The type species is ''D. podenasi''.
|
|-
|
''[[Elephantomyia christelae]]''<ref name=SRElephantomyia>{{Cite journal|last1=Kania-Kłosok |first1=I. |last2=Krzemiński |first2=W. |title=Recent discoveries of new ''Elephantomyia'' (Diptera, Limoniidae) fossils in Baltic amber |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 23647 |year=2021 |doi=10.1038/s41598-021-03022-3 |pmid=34880309 |pmc=8655017 |bibcode=2021NatSR..1123647K |doi-access=free }}</ref>
|
Sp. nov
|
|
Kania-Kłosok & Krzemiński
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A species of ''[[Elephantomyia]]''.
|
|-
|
''[[Elephantomyia prima]]''<ref name=SRElephantomyia />
|
Sp. nov
|
|
Kania-Kłosok & Krzemiński
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A species of ''Elephantomyia''.
|
|-
|
''[[Forcipomyia rasnitsyni]]''<ref>{{Cite journal|last1=Szadziewski |first1=R. |last2=Sontag |first2=E. |last3=Pankowski |first3=M. V. |title=A new biting midge of the genus ''Forcipomyia'' Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=441–444 |doi=10.11646/palaeoentomology.4.5.9 |issn=2624-2826 |s2cid=244172069 }}</ref>
|
Sp. nov
|
Valid
|
Szadziewski, Sontag & Pankowski
|
Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A species of ''[[Forcipomyia]]''.
|
|-
|
''[[Helius hispanicus]]''<ref name=SRHelius>{{Cite journal|last1=Kania-Kłosok |first1=I. |last2=Krzemiński |first2=W. |last3=Arillo |first3=A. |title=Two new long-rostrum cranefly species from the Cretaceous Iberian amber (Diptera, Limoniidae, ''Helius'') |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 12851 |doi=10.1038/s41598-021-91803-1 |doi-access=free |pmid=34145304 |pmc=8213732 |bibcode=2021NatSR..1112851K }}</ref>
|
Sp. nov
|
|
Kania-Kłosok, Krzemiński & Arillo
|
[[Early Cretaceous]] ([[Albian]])
|
Utrillas Group<br>(Álava amber)
|
{{Flag|Spain}}
|
A species of ''[[Helius (fly)|Helius]]''.
|
|-
|
''[[Helius turolensis]]''<ref name=SRHelius />
|
Sp. nov
|
|
Kania-Kłosok, Krzemiński & Arillo
|
Early Cretaceous (Albian)
|
Utrillas Group<br>(San Just amber)
|
{{Flag|Spain}}
|
A species of ''Helius''.
|
|-
|
''[[Hirmoneura messelense]]''<ref>{{Cite journal|last1=Wedmann |first1=S. |last2=Hörnschemeyer |first2=T. |last3=Engel |first3=M. S. |last4=Zetter |first4=R. |last5=Grímsson |first5=F. |title=The last meal of an Eocene pollen-feeding fly |year=2021 |journal=Current Biology |volume=31 |issue=9 |pages=2020–2026.e4 |doi=10.1016/j.cub.2021.02.025 |pmid=33705719 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Wedmann ''et al.''
|
[[Eocene]] ([[Lutetian]])
|
[[Messel Formation]]
|
{{Flag|Germany}}
|
A member of the family [[Nemestrinidae]].
|
|-
|
''[[Lebambromyia sacculifera]]''<ref>{{Cite journal|last1=Badano |first1=D. |last2=Zhang |first2=Q. |last3=Fratini |first3=M. |last4=Maugeri |first4=L. |last5=Bukreeva |first5=I. |last6=Longo |first6=E. |last7=Wilde |first7=F. |last8=Yeates |first8=D. K. |last9=Cerretti |first9=P. |title=Discovery of ''Lebambromyia'' in Myanmar Cretaceous Amber: Phylogenetic and Biogeographic Implications (Insecta, Diptera, Phoroidea) |year=2021 |journal=Insects |volume=12 |issue=4 |pages=Article 354 |doi=10.3390/insects12040354 |pmid=33923404 |pmc=8074229 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Badano, Zhang, Yeates & Cerretti ''in'' Badano ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Phoroidea]].
|
|-
|
''[[Leptotarsus andradei]]''<ref name=CRLeptotarsus>{{Cite journal|last1=Ribeiro |first1=G. C. |last2=Santos |first2=R. R. |last3=Santos |first3=D. |title=Four new species of the genus ''Leptotarsus'' Guérin-Méneville, 1831 (Insecta: Diptera: Tipulidae) from the Lower Cretaceous Crato Formation of Brazil |year=2021 |journal=Cretaceous Research |volume=123 |pages=Article 104776 |doi=10.1016/j.cretres.2021.104776 |s2cid=233902054 }}</ref>
|
Sp. nov
|
Valid
|
Ribeiro, Santos & Santos
|
[[Early Cretaceous]]
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A species of ''[[Leptotarsus]]''.
|
|-
|
''[[Leptotarsus gelhausi]]''<ref name=CRLeptotarsus />
|
Sp. nov
|
Valid
|
Ribeiro, Santos & Santos
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A species of ''Leptotarsus''.
|
|-
|
''[[Leptotarsus ivoneae]]''<ref name=CRLeptotarsus />
|
Sp. nov
|
Valid
|
Ribeiro, Santos & Santos
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A species of ''Leptotarsus''.
|
|-
|
''[[Leptotarsus lemeae]]''<ref name=CRLeptotarsus />
|
Sp. nov
|
Valid
|
Ribeiro, Santos & Santos
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A species of ''Leptotarsus''.
|
|-
|
''[[Mesoacentron]]''<ref name=Mesoacentron>{{Cite journal|last1=Giłka |first1=W. |last2=Zakrzewska |first2=M. |last3=Lukashevich |first3=E. D. |last4=Vorontsov |first4=D. D. |last5=Soszyńska-Maj |first5=A. |last6=Skibińska |first6=K. |last7=Cranston |first7=P. S. |title=Wanted, tracked down and identified: Mesozoic non-biting midges of the subfamily Chironominae (Chironomidae, Diptera) |year=2021 |journal=Zoological Journal of the Linnean Society |volume= 194|issue=3 |pages=874–892 |doi=10.1093/zoolinnean/zlab020 |doi-access=free |hdl=1885/294434 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Giłka, Zakrzewska, Lukashevich & Cranston ''in'' Giłka ''et al.''
|
Late Cretaceous ([[Santonian]])
|
[[Kheta Formation]]<br>(Taimyr amber)
|
{{Flag|Russia}}
|
A member of the family [[Chironomidae]] belonging to the subfamily [[Chironominae]] and the tribe [[Pseudochironomini]]. The type species is ''M. kaluginae''.
|
|-
|
''[[Meunierohelea anglesensis]]''<ref name=HBAnglesea />
|
Sp. nov
|
In press
|
Peñalver ''et al.''
|
Eocene
|
Anglesea amber
|
{{Flag|Australia}}
|
A member of the family [[Ceratopogonidae]].
|
|-
|
''[[Minyohelea nexuosa]]''<ref>{{Cite journal|last1=Pielowska-Ceranowska |first1=A. |last2=Azar |first2=D. |last3=Szwedo |first3=J. |title=New fossil biting midge from the Lower Cretaceous amber of Lebanon (Diptera: Ceratopogonidae) |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105004 |doi=10.1016/j.cretres.2021.105004| issn=0195-6671  |s2cid=239639779 }}</ref>
|
Sp. nov
|
In press
|
Pielowska-Ceranowska, Azar & Szwedo
|
Early Cretaceous ([[Barremian]])
|
[[Lebanese amber]]
|
{{Flag|Lebanon}}
|
A member of the family Ceratopogonidae.
|
|-
|
''[[Myriophora asiatica]]''<ref name=PalaeoworldPhoridae />
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
Zhangpu amber
|
{{Flag|China}}
|
A species of ''[[Myriophora]]''.
|
|-
|
''[[Neoarisemus groehni]]''<ref>{{Cite journal|last=Wagner |first=R. |title=''Neoarisemus groehni'' sp. nov., a notable moth fly (Diptera, Psychodidae, Psychodinae) from Ukrainian amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=19–22 |doi=10.11646/palaeoentomology.4.1.3 |s2cid=233941588 }}</ref>
|
Sp. nov
|
Valid
|
Wagner
|
Eocene
|
|
{{Flag|Ukraine}}
|
A member of the family [[Psychodidae]].
|
|-
|
''[[Orimarguloides]]''<ref>{{Cite journal|last1=Men |first1=Q. |last2=Podenas |first2=S. |title=A new genus of Limoniidae (Diptera: Tipuloidea) from the mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104915 |doi=10.1016/j.cretres.2021.104915 |bibcode=2021CrRes.12604915M }}</ref>
|
Gen. et sp. nov
|
Valid
|
Men & Podenas
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the [[crane fly]] family [[Limoniidae]]. Genus includes new species ''O. simplex''.
|
|-
|
''[[Palaeocentron]]''<ref name=Mesoacentron />
|
Gen. et sp. nov
|
Valid
|
Giłka, Zakrzewska, Lukashevich & Cranston ''in'' Giłka ''et al.''
|
Cretaceous (probably Albian–Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is ''P. krzeminskii''.
|
|-
|
''[[Palaeoglaesum|Palaeoglaesum pilosus]]''<ref name=Insects12Palaeoglaesum>{{Cite journal|last1=Skibińska |first1=K. |last2=Albrycht |first2=M. |last3=Zhang |first3=Q. |last4=Giłka |first4=W. |last5=Zakrzewska |first5=M. |last6=Krzemiński |first6=W. |title=Diversity of the Fossil Genus ''Palaeoglaesum'' Wagner (Diptera, Psychodidae) in the Upper Cretaceous Amber of Myanmar |year=2021 |journal=Insects |volume=12 |issue=3 |pages=Article 247 |doi=10.3390/insects12030247 |pmid=33809471 |pmc=8000030 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Skibińska, Krzemiński & Zhang ''in'' Skibińska ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Psychodidae]] belonging to the subfamily [[Bruchomyiinae]].
|
|-
|
''Palaeoglaesum stebneri''<ref name=Insects12Palaeoglaesum />
|
Sp. nov
|
Valid
|
Skibińska & Krzemiński ''in'' Skibińska ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
|
|-
|
''Palaeoglaesum teres''<ref name=Insects12Palaeoglaesum />
|
Sp. nov
|
Valid
|
Skibińska & Albrycht ''in'' Skibińska ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
|
|-
|
''[[Paleoplatyura agnieszkae]]''<ref name=Insects131Paleoplatyura>{{Cite journal|last1=Ševčík |first1=J. |last2=Krzemiński |first2=W. |last3=Skibińska |first3=K. |title=Extant Genus in the Mesozoic: ''Paleoplatyura'' Meunier (Diptera: Keroplatidae) Found in the Cretaceous Amber of Myanmar |year=2021 |journal=Insects |volume=13 |issue=1 |pages=Article 24 |doi=10.3390/insects13010024 |pmid=35055867 |pmc=8778176 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Ševčík, Krzemiński & Skibińska
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''[[Paleoplatyura]]''.
|
|-
|
''[[Paleoplatyura magnifica]]''<ref name=Insects131Paleoplatyura />
|
Sp. nov
|
Valid
|
Ševčík, Krzemiński & Skibińska
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''Paleoplatyura''.
|
|-
|
''[[Paleoplatyura miae]]''<ref name=Insects131Paleoplatyura />
|
Sp. nov
|
Valid
|
Ševčík, Krzemiński & Skibińska
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''Paleoplatyura''.
|
|-
|
''[[Paleotelmatoscopus]]''<ref>{{Cite journal|last1=Curler |first1=G. R. |last2=Skibińska |first2=K. |title=''Paleotelmatoscopus'', a proposed new genus for some fossil moth flies (Diptera, Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species |year=2021 |journal=Zootaxa |volume=4927 |issue=4 |pages=505–524 |doi=10.11646/zootaxa.4927.4.2|issn=1175-5326  |pmid=33756688 |s2cid=232339113 }}</ref>
|
Gen. et comb. et sp. nov
|
Valid
|
Curler & Skibińska
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Psychodidae]]. The type species is ''"[[Pericoma]]" formosa'' Meunier (1905); genus also includes new species ''P. madrizi''.
|
|-
|
''[[Polymera alexanderi]]''<ref>{{Cite journal|last1=Krzemiński |first1=W. |last2=Kopeć |first2=K. |last3=Pełczyńska |first3=A. |last4=Soszyńska-Maj |first4=A. |title=''Polymera alexanderi'' n. sp., a new species of Limoniidae (Diptera) from Dominican amber |year=2021 |journal=Acta Zoologica Cracoviensia |volume=64 |issue=2 |pages=169–174 |doi=10.3409/azc.64.03 |s2cid=245628471 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Krzemiński ''et al.''
|
Miocene
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A species of ''[[Polymera]]''.
|
|-
|
''[[Protopenthetria]]''<ref>{{Cite journal|last1=Li |first1=L. |last2=Skibińska |first2=K. |last3=Krzemiński |first3=W. |last4=Wang |first4=B. |last5=Xiao |first5=C. |last6=Zhang |first6=Q. |title=A new March fly ''Protopenthetria skartveiti'' gen. nov. et sp. nov. (Diptera, Bibionidae, Plecinae) from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104924 |doi=10.1016/j.cretres.2021.104924|bibcode=2021CrRes.12704924L |issn=0195-6671 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Bibionidae]]. Genus includes new species ''P. skartveiti''.
|
|-
|
''[[Rhabdomastix asiatica]]''<ref name=EZJRhabdomastix>{{Cite journal|last1=Kania-Kłosok |first1=I. |last2=Jordan-Stasiło |first2=W. |last3=Kopeć |first3=K. |last4=Janiszewska |first4=K. |last5=Krzemiński |first5=W. |title=The first stage of the evolution of ''Rhabdomastix'' (Diptera, Limoniidae) and the taxonomic implications of genus |year=2021 |journal=The European Zoological Journal |volume=88 |issue=1 |pages=1152–1169 |doi=10.1080/24750263.2021.2003880 |s2cid=244765716 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Kania-Kłosok ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''[[Rhabdomastix]]''.
|
|-
|
''[[Rhabdomastix cretacica]]''<ref name=EZJRhabdomastix />
|
Sp. nov
|
Valid
|
Kania-Kłosok ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''Rhabdomastix''.
|
|-
|
''[[Rhabdomastix krzeminskae]]''<ref name=EZJRhabdomastix />
|
Sp. nov
|
Valid
|
Kania-Kłosok ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''Rhabdomastix''.
|
|-
|
''[[Rhabdomastix myanmae]]''<ref name=EZJRhabdomastix />
|
Sp. nov
|
Valid
|
Kania-Kłosok ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A species of ''Rhabdomastix''.
|
|-
|
''[[Rhipidoxylomyia rasnitsyni]]''<ref name=Palaeoentomology46Nel />
|
Sp. nov
|
Valid
|
Nel
|
Early Eocene
|
Oise amber
|
{{Flag|France}}
|
A member of the family Cecidomyiidae.
|
|-
|
''[[Stempellinella gilkai]]''<ref>{{Cite journal|last1=Zakrzewska |first1=M. |last2=Jankowska |first2=A. |title=A new fossil ''Stempellinella'' from Eocene Baltic amber, with systematic notes and amendment of the generic diagnosis (Diptera: Chironomidae) |year=2021 |journal=Zootaxa |volume=4908 |issue=4 |pages=505–514 |doi=10.11646/zootaxa.4908.4.4 |pmid=33756601 |s2cid=232340439 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Zakrzewska & Jankowska
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A species of ''[[Stempellinella]]''.
|
|-
|
''[[Trichoneura xavieri]]''<ref>{{Cite journal|last1=Kania-Kłosok |first1=I. |last2=Krzemiński |first2=W. |last3=Kopeć |first3=K. |last4=Arillo |first4=A. |title=The Oldest Evolutionary Lineage of ''Trichoneura'' Loew, 1850 (Diptera, Limoniidae) and the First Evidence of This Genus in Cretaceous Spanish Amber |year=2021 |journal=Insects |volume=12 |issue=5 |pages=Article 411 |doi=10.3390/insects12050411 |pmid=34063589 |pmc=8147636 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Kania-Kłosok ''et al.''
|
[[Early Cretaceous]] ([[Albian]])
|
Utrillas Group
|
{{Flag|Spain}}
|
A species of ''[[Trichoneura (fly)|Trichoneura]]''.
|
|-
|
''[[Zhangsolva|Zhangsolva burmensis]]''<ref>{{cite book |last1=Zhang |first1=Q. |last2=Zhang |first2=J. |year=2021 |chapter=New record of ''Zhangsolva'' Nagatomi & Yang, 1998 (Diptera, Zhangsolvidae) from mid-Cretaceous Burmese amber and its implications |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |volume=521 |pages=139–148 |publisher=The Geological Society of London |doi=10.1144/SP521-2021-150 |s2cid=245059011 }}</ref>
|
Sp. nov
|
In press
|
Zhang & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Zhangsolvidae]].
|
|-
|}

====Dipteran research====
* Baranov ''et al.'' (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.<ref>{{Cite journal|last1=Baranov |first1=V. A. |last2=Engel |first2=M. S. |last3=Hammel |first3=J. |last4=Hörnig |first4=M. K. |last5=van de Kamp |first5=T. |last6=Zuber |first6=M. |last7=Haug |first7=J. T. |title=Synchrotron-radiation computed tomography uncovers ecosystem functions of fly larvae in an Eocene forest |year=2021 |journal=Palaeontologia Electronica |volume=24 |issue=1 |pages=Article number 24.1.a07 |doi=10.26879/1129 |doi-access=free }}</ref>

===Mecopterans===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Conicholcorpa|Conicholcorpa longa]]''<ref>{{Cite journal|last1=Zhang |first1=Y. |last2=Shih |first2=P. J. M. |last3=Wang |first3=J. |last4=McNamara |first4=M. E. |last5=Shih |first5=C. |last6=Ren |first6=D. |last7=Gao |first7=T. |year=2021 |title=Jurassic scorpionflies (Mecoptera) with swollen first metatarsal segments suggesting sexual dimorphism |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 47 |doi=10.1186/s12862-021-01771-3 |pmid=33743581 |pmc=7981964 |doi-access=free }}</ref>
|
Sp. nov
|
|
Zhang, Shih & Ren ''in'' Zhang ''et al.''
|
[[Middle Jurassic]]
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A [[Mecoptera|scorpionfly]] belonging to the family [[Holcorpidae]].
|
|-
|
''[[Glanomerope]]''<ref>{{Cite journal|last1=Poschmann |first1=M. J. |last2=Nel |first2=A. |title=The first Permian scorpionfly from Germany (Insecta, Panorpida: Protomeropidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=231–236 |doi=10.11646/palaeoentomology.4.3.10 |s2cid=237881817 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poschmann & Nel
|
[[Permian]]
|
[[Meisenheim Formation]]
|
{{Flag|Germany}}
|
A scorpionfly belonging to the family [[Protomeropidae]]. Genus includes new species ''G. virgoferroa''.
|
|-
|
''[[Longiphlebia|Longiphlebia incompleta]]''<ref>{{Cite journal|last1=Lian |first1=X. |last2=Cai |first2=C. |last3=Huang |first3=D. |year=2021 |title=A new orthophlebiid scorpionfly (Insecta, Orthophlebiidae) from the Late Jurassic Linglongta biota of northern China |journal=Historical Biology |volume=33 |issue=12 |pages=3585–3589 |doi=10.1080/08912963.2021.1878512 |s2cid=233934478 }}</ref>
|
Sp. nov
|
In press
|
Lian, Cai & Huang
|
[[Late Jurassic]]
|
[[Tiaojishan Formation]]
|
{{Flag|China}}
|
A [[Mecoptera|scorpionfly]] belonging to the family [[Orthophlebiidae]].
|
|-
|
''[[Mesopsyche|Mesopsyche jinsuoguanensis]]''<ref name=Zootaxa49953>{{Cite journal|last1=Lian |first1=X. |last2=Cai |first2=C. |last3=Huang |first3=D. |year=2021 |title=New species of ''Mesopsyche'' Tillyard, 1917 (Mecoptera: Mesopsychidae) from the Triassic of northwestern China |journal=Zootaxa |volume=4995 |issue=3 |pages=565–572 |doi=10.11646/zootaxa.4995.3.10 |pmid=34810550 |s2cid=237782965 }}</ref>
|
Sp. nov
|
Valid
|
Lian, Cai & Huang
|
[[Middle Triassic]]
|
Tongchuan Formation
|
{{Flag|China}}
|
A member of the family [[Mesopsychidae]]. Originally described as a species of ''Mesopsyche''; transferred by Lian ''et al.'' (2024) to the genus ''[[Turbidapsyche]]''.<ref name=Turbidapsyche>{{Cite journal|last1=Lian |first1=X.-N. |last2=Zhang |first2=Y. |last3=Cai |first3=C.-Y. |last4=Feng |first4=Z. |last5=Huang |first5=D.-Y. |title=A new genus of Mesopsychidae from the late Middle Triassic Tongchuan entomofauna and its taxonomic and palaeogeographic implications |year=2024 |journal=Mesozoic |volume=1 |issue=2 |pages=144–158 |doi=10.11646/mesozoic.1.2.7 |doi-access=free }}</ref>
|
|-
|
''Mesopsyche liaoi''<ref name=Zootaxa49953 />
|
Sp. nov
|
Valid
|
Lian, Cai & Huang
|
[[Late Triassic]]
|
Huangshanjie Formation
|
{{Flag|China}}
|
A member of the family Mesopsychidae. Originally described as a species of ''Mesopsyche''; transferred by Lian ''et al.'' (2024) to the genus ''Turbidapsyche''.<ref name=Turbidapsyche />
|
|-
|}

==Clade Archaeorthoptera==
===†Caloneurodea===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Aviobiella]]''<ref>{{Cite journal|last1=Nel |first1=A. |last2=Roques |first2=P. |title=The second Caloneurodea from the Moscovian of Avion, France (Insecta, Archaeorthoptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=320–322 |doi=10.11646/palaeoentomology.4.4.5|issn=2624-2826  |s2cid=239666503 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel & Roques
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}
|
A member of [[Archaeorthoptera]] belonging to the order [[Caloneurodea]]. Genus includes new species ''A. garroustei''.
|
|-
|
''[[Oudardgramma]]''<ref>{{cite journal |last1=Nel |first1=A. |last2=Roques |first2=P. |year=2021 |title=A curious new Caloneurodea from the Bashkirian of the Pas-de-Calais, France (Insecta Archaeorthoptera) |journal=Biosis: Biological Systems |volume=2 |issue=4 |pages=358–360 |doi=10.37819/biosis.001.04.0159 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
|
Nel & Roques
|
Carboniferous (Bashkirian)
|
|
{{Flag|France}}
|
The type species is ''O. bruayensis''.
|
|-
|
''[[Paraspiculum]]''<ref name=Contracladus />
|
Gen. et sp. nov
|
In press
|
Dvořák ''et al.''
|
Carboniferous (Moscovian)
|
|
{{Flag|Germany}}
|
A member of Archaeorthoptera belonging to the group [[Caloneurodea]]. Genus includes new species ''P. brauckmannii''.
|
|-
|}

===†Cnemidolestodea===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Avionxixia]]''<ref name=Archaeogerarus />
|
Gen. et sp. nov
|
Valid
|
Nel & Roques
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}
|
A member of Archaeorthoptera belonging to the family [[Cnemidolestidae]]. The type species is ''A. gui''.
|
|-
|
''[[Ctenoptilus|Ctenoptilus frequens]]''<ref>{{Cite journal|last1=Chen |first1=L. |last2=Gu |first2=J.-J. |last3=Yang |first3=Q. |last4=Ren |first4=D. |last5=Blanke |first5=A. |last6=Béthoux |first6=O. |year=2021 |title=Ovipositor and mouthparts in a fossil insect support a novel ecological role for early orthopterans in 300 million years old forests |journal=eLife |volume=10 |pages=e71006 |doi=10.7554/eLife.71006 |pmid=34844668 |pmc=8631945 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Chen ''et al.''
|
Carboniferous ([[Bashkirian]] to [[Moscovian (Carboniferous)|Moscovian]])
|
Yanghugou Formation
|
{{Flag|China}}
|
A member of Archaeorthoptera belonging to the family [[Ctenoptilidae]].
|
|-
|
''[[Palatinarkema]]''<ref>{{Cite journal|last1=Nel |first1=A. |last2=Poschmann |first2=M. J. |title=A new representative of the "orthopteroid" family Cnemidolestidae from the lower Permian of Germany |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=3 |pages=641–646 |doi=10.4202/app.00879.2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel & Poschmann
|
Early Permian (probably [[Asselian]]–?[[Sakmarian]])
|
Meisenheim  Formation
|
{{Flag|Germany}}
|
A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is ''P. prokopi''.
|
|-
|
''[[Piesbergopterum]]''<ref name=Contracladus />
|
Gen. et 2 sp. nov
|
In press
|
Dvořák ''et al.''
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}<br>
{{Flag|Germany}}
|
A member of Archaeorthoptera belonging to the group [[Cnemidolestodea]]. Genus includes new species ''P. punctatum'' Dvořák ''et al.'' and ''P. avionensis'' Nel & Roques.<ref>{{Cite journal|last1=Nel |first1=A. |last2=Roques |first2=P. |title=The second species of the cnemidolestid genus ''Piesbergopterum'' from the Upper Carboniferous of Avion, Northern France (Archaeorthoptera: Cnemidolestidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=323–325 |doi=10.11646/palaeoentomology.4.4.6 |s2cid=239687328 }}</ref>
|
|-
|}

===Orthopterans===
====New Orthopteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Amberotridactylus]]''<ref name=Amberotridactylus>{{Cite journal|last1=Du |first1=X. |last2=Xu |first2=C. |last3=Zhang |first3=L. |title=Two new pygmy mole crickets from mid-Cretaceous Kachin amber of northern Myanmar (Orthoptera: Tridactyloidea: Tridactylidae) |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105002 |doi=10.1016/j.cretres.2021.105002 | issn=0195-6671  |s2cid=238648688 }}</ref>
|
Gen. et sp. nov
|
In press
|
Du, Xu & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Tridactylidae]]. Genus includes new species ''A. cheni''.
|
|-
|
''[[Archaboilus|Archaboilus polyneurus]]''<ref>{{Cite journal|last1=Gu |first1=J.-J. |last2=Yang |first2=X. |last3=Huang |first3=R. |last4=Yue |first4=Y. |last5=Ren |first5=D. |title=New species and material of Hagloidea (Insecta, Ensifera) from the Yanliao biota of China |year=2021 |journal=ZooKeys |issue=1033 |pages=183–190 |doi=10.3897/zookeys.1033.63571 |doi-access=free |pmid=33958925 |pmc=8084857 }}</ref>
|
Sp. nov
|
Valid
|
Gu, Yue & Ren ''in'' Gu ''et al.''
|
[[Middle Jurassic]] ([[Bathonian]]–[[Callovian]])
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
An [[ensifera]]n belonging to the group [[Hagloidea]] and the family [[Haglidae]].
|
|-
|
''[[Caelielca]]''<ref name=Caelielca>{{Cite journal|last=Uchida |first=K. |title=Two new genera of the Elcanidae (order: Orthoptera) from middle Cretaceous Burmese amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105092 |doi=10.1016/j.cretres.2021.105092 |s2cid=244377365 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
In press
|
Uchida
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Elcanidae]]. Genus includes new species ''C. spinocrus''.
|
|-
|
''[[Cascogryllus|Cascogryllus setosus]]''<ref name=Amberotridactylus />
|
Sp. nov
|
In press
|
Du, Xu & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Tridactylidae.
|
|-
|
''[[Cratoelcana|Cratoelcana rasnitsyni]]''<ref name=HBgrasshoppers>{{Cite journal|last1=Nel |first1=A. |last2=Jouault |first2=C. |title=New grasshoppers (Orthoptera: Elcanidae, Locustopsidae) from the Lower Cretaceous Crato Formation suggest a biome homogeneity in Central Gondwana |journal=Historical Biology |year=2022 |volume=34 |issue=10 |pages=2070–2078 |doi=10.1080/08912963.2021.2000602 |s2cid=244167833 |url=https://hal.sorbonne-universite.fr/hal-03439696/file/Nel%20et%20Jouault%20-%202021%20-%20New%20grasshoppers%20%28Orthoptera%20Elcanidae%2C%20Locustops.pdf }}</ref>
|
Sp. nov
|
In press
|
Nel & Jouault
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A member of the family Elcanidae.
|
|-
|
''[[Hukawnelca]]''<ref name=Caelielca />
|
Gen. et sp. nov
|
In press
|
Uchida
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Elcanidae. Genus includes new species ''H. gracile''.
|
|-
|
''[[Panorpidium|Panorpidium spica]]''<ref>{{Cite journal|last1=Kim |first1=D.-Y. |last2=Lee |first2=M. |last3=Nam |first3=G.-S. |last4=Park |first4=T.-Y. S. |title=The first orthopteran fossils from the Lower Cretaceous (Albian) Jinju Formation of Korea: ethological implications for elcanids |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104843 |doi=10.1016/j.cretres.2021.104843 |bibcode=2021CrRes.12504843K }}</ref>
|
Sp. nov
|
In press
|
Kim ''et al.''
|
[[Early Cretaceous]] ([[Albian]])
|
[[Jinju Formation]]
|
{{Flag|South Korea}}
|
An [[orthoptera]]n belonging to the family Elcanidae.
|
|-
|
''[[Permohagla]]''<ref>{{Cite journal|last=Gorochov |first=A. V. |title=The oldest representative of the superfamily Hagloidea (Orthoptera) from the Upper Permian of Russia, with remarks on katydid-like forms from the Paleozoic |year=2021 |journal=Paleontological Journal |volume=55 |issue=5 |pages=552–558 |doi=10.1134/S0031030121050063 |s2cid=238640705 |url=https://elibrary.ru/item.asp?id=46428495 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Gorochov
|
Late Permian
|
|
{{Flag|Russia}}<br/>({{Flag|Kemerovo Oblast}})
|
A member of [[Hagloidea]]. Genus includes new species ''P. tomica''.
|
|-
|
''[[Phyllotridactylus]]''<ref>{{Cite journal|last1=Xu |first1=X. |last2=Wang |first2=B. |last3=Fan |first3=L. |last4=Jarzembowski |first4=E. A. |last5=Fang |first5=Y. |last6=Wang |first6=H. |last7=Li |first7=T. |last8=Zhuo |first8=D. |last9=Ding |first9=M. |last10=Engel |first10=M. S. |title=Widespread mimicry and camouflage among mid-Cretaceous insects |year=2021 |journal=Gondwana Research |volume=101 |pages=94–102 |doi=10.1016/j.gr.2021.07.025|issn=1342-937X  }}</ref>
|
Gen. et sp. nov
|
In press
|
Xu ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Tridactylidae. Genus includes new species ''P. wangi''.
|
|-
|
''[[Pseudoacrida|Pseudoacrida sennlaubi]]''<ref name=HBgrasshoppers />
|
Sp. nov
|
In press
|
Nel & Jouault
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A member of the family [[Locustopsidae]].
|
|-
|}

====Orthopteran research====
* Revision of the fossil record of the family [[Trigonidiidae]] is published by Desutter-Grandcolas ''et al.'' (2021).<ref>{{Cite journal|last1=Desutter-Grandcolas |first1=L. |last2=Hugel |first2=S. |last3=Nel |first3=A. |last4=Warren |first4=B. H. |last5=Souza-Dias |first5=P. |last6=Chintauan-Marquier |first6=I. C. |title=Updated diagnoses for the cricket family Trigonidiidae (Insecta: Orthoptera: Grylloidea) and its subfamilies (Trigonidiinae, Nemobiinae), with a review of the fossil record |year=2021 |journal=Zoologischer Anzeiger |volume=294 |pages=80–91 |doi=10.1016/j.jcz.2021.06.004 |url=https://hal.archives-ouvertes.fr/hal-03366682/file/Desutter-Grandcolas%20et%20al.%20-%202021%20-%20Updated%20diagnoses%20for%20the%20cricket%20family%20Trigonidi.pdf }}</ref>

===†Titanoptera===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Theiatitan]]''<ref>{{Cite journal|last1=Schubnel |first1=T. |last2=Legendre |first2=F. |last3=Roques |first3=P. |last4=Garrouste |first4=R. |last5=Cornette |first5=R. |last6=Perreau |first6=M. |last7=Perreau |first7=N. |last8=Desutter-Grandcolas |first8=L. |last9=Nel |first9=A. |title=Sound vs. light: wing-based communication in Carboniferous insects |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 794 |doi=10.1038/s42003-021-02281-0 |doi-access=free |pmid=34239029 |pmc=8266802 }}</ref>
|
Fam. et gen. et sp. nov
|
Valid
|
Schubnel, Roques & Nel ''in'' Schubnel ''et al.''
|
Carboniferous ([[Moscovian (Carboniferous)|Moscovian]])
|
|
{{Flag|France}}
|
A member of [[Titanoptera]] belonging to the new family [[Theiatitanidae]]. The type species is ''T. azari''.
|[[File:Oldest-Carboniferous-Titanoptera-Theiatitan-azari-gen-sp-nov-holotype.png|thumb|center|upright|''Theiatitan azari'']]
|-
|}

===Other archaeorthopterans===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Archaeogerarus]]''<ref name=Archaeogerarus>{{Cite journal|last1=Nel |first1=A. |last2=Roques |first2=P. |title=A new family and two new genera from Avion, Northern France, confirm the high Moscovian (late Carboniferous) diversity of the insect superorder Archaeorthoptera |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=4 |pages=879–884 |doi=10.4202/app.00940.2021 |s2cid=244879144 |doi-access=free }}</ref>
|
Fam. et gen. et sp. nov
|
Valid
|
Nel & Roques
|
[[Carboniferous]] ([[Moscovian (Carboniferous)|Moscovian]])
|
|
{{Flag|France}}
|
A member of [[Archaeorthoptera]] belonging to the new family [[Archaeogeraridae]]. The type species is ''A. schubneli''.
|
|-
|
''[[Avionugonioneura]]''<ref>{{Cite journal|last1=Nel |first1=A. |last2=Roques |first2=P. |year=2021 |title=A new strange Archaeorthoptera from the Moscovian of Avion (France) (Insecta, Polyneoptera) |journal=Historical Biology |volume= 34|issue=9 |pages=1769–1771 |doi=10.1080/08912963.2021.1978082 |s2cid=239222016 |url=https://hal.sorbonne-universite.fr/hal-03356138/file/Nel%20et%20Roques%20-%202021%20-%20A%20new%20strange%20Archaeorthoptera%20from%20the%20Moscovian%20.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel & Roques
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}
|
A member of Archaeorthoptera possibly belonging to the family [[Nugonioneuridae]]. Genus includes new species ''A. jouaulti''.
|
|-
|
''[[Contracladus]]''<ref name=Contracladus>{{Cite journal|last1=Dvořák |first1=T. |last2=Pecharová |first2=M. |last3=Leipner |first3=A. |last4=Nel |first4=A. |last5=Prokop |first5=J. |year=2021 |title=New archaeorthopteran insects from the Pennsylvanian of Piesberg reveal unexpected mosaic of morphological traits and colouration pattern of the tegmina |journal=Historical Biology |volume=33 |issue=12 |pages=3375–3383 |doi=10.1080/08912963.2020.1867127 |s2cid=234189184 }}</ref>
|
Gen. et sp. nov
|
In press
|
Dvořák ''et al.''
|
[[Carboniferous]] ([[Moscovian (Carboniferous)|Moscovian]])
|
|
{{Flag|Germany}}
|
A member of [[Archaeorthoptera]] of uncertain phylogenetic placement. Genus includes new species ''C. impar''.
|
|-
|
''[[Duquesnia]]''<ref>{{Cite journal|last1=Nel |first1=A. |last2=Roques |first2=P. |last3=Duquesne |first3=H. |title=A further strange Archaeorthoptera from the Moscovian of Nord-Pas-de-Calais (France) (Insecta, Polyneoptera) |year=2021 |journal=Zootaxa |volume=5047 |issue=2 |pages=165–170 |doi=10.11646/zootaxa.5047.2.5 |pmid=34810773 |s2cid=244251921 |url=https://hal.sorbonne-universite.fr/hal-03385759/file/Nel%20et%20al.%20-%202021%20-%20A%20further%20strange%20Archaeorthoptera%20from%20the%20Moscov.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel & Roques ''in'' Nel, Roques & Duquesne
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}
|
A member of Archaeorthoptera. Genus includes new species ''D. gallica''.
|
|-
|
''[[Penanoptera]]''<ref>{{Cite journal|last1=Nel |first1=A. |last2=Poschmann |first2=M. J. |title=A new archaeorthopteran insect from the Early Permian of the Saar-Nahe Basin, SW Germany |year=2021 |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=301 |issue=1 |pages=79–84 |doi=10.1127/njgpa/2021/0998 |s2cid=237729949 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel & Poschmann
|
Early Permian
|
Saar–Nahe Basin
|
{{Flag|Germany}}
|
A member of [[Archaeorthoptera]]. Genus includes new species ''P. manseri''.
|
|-
|}

==Clade Coleopterida==
===Coleoptera===
====Adephaga====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Antephilorhizus]]''<ref>{{Cite journal|last1=Kirichenko-Babko |first1=M. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |year=2021 |title=A new genus and species of Lebiini (Coleoptera: Carabidae) from late Eocene Rovno amber |journal=Historical Biology |volume= 34|issue=3 |pages=436–442 |doi=10.1080/08912963.2021.1924701 |s2cid=241825804 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kirichenko-Babko, Perkovsky & Vasilenko
|
Eocene
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
A [[ground beetle]] belonging to the tribe [[Lebiini]]. Genus includes new species ''A. pripiatiensis''.
|
|-
|
''[[Balticeler]]''<ref>{{Cite journal|last1=Schmidt |first1=J. |last2=Scholz |first2=S. |last3=Maddison |first3=D. R. |title=''Balticeler kerneggeri'' gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae) |year=2021 |journal=Deutsche Entomologische Zeitschrift |volume=68 |issue=1 |pages=207–224 |doi=10.3897/dez.68.66181 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Schmidt & Maddison ''in'' Schmidt, Scholz & Maddison
|
Eocene
|
[[Baltic amber]]
|
Europe (Baltic Sea region)
|
A [[ground beetle]] belonging to the subfamily [[Trechinae]]. The type species is ''B. kerneggeri''.
|
|-
|
''[[Calosoma kimi]]''<ref>{{Cite journal|last1=Lee |first1=S. B. |last2=Nam |first2=G.-S. |year=2021 |title=A new species of fossil ''Calosoma'' (Coleoptera: Carabidae) from the Geumgwangdong Formation (Early Miocene), South Korea |journal=Zootaxa |volume=5072 |issue=1 |pages=1–11 |doi=10.11646/zootaxa.5072.1.1 |pmid=35390890 |s2cid=248024592 }}</ref>
|
Sp. nov
|
Valid
|
Lee & Nam
|
Early Miocene
|
[[Geumgwangdong Formation]]
|
{{Flag|South Korea}}
|
A species of ''[[Calosoma]]''.
|
|-
|}
====Archostemata====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Jurodes shef]]''<ref>{{Cite journal|last1=Yan |first1=E. V. |last2=Strelnikova |first2=O. D. |last3=Ponomarenko |first3=A. G. |title=Jurodidae (Coleoptera: Archostemata) from Transbaikalia: new findings and redescription of type material |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=604–613 |doi=10.11646/palaeoentomology.4.6.9 |s2cid=245203743 }}</ref>
|
Sp. nov
|
Valid
|
Yan, Strelnikova & Cai
|
[[Early Cretaceous]] ([[Berriasian]]-[[Barremian]])
|
Gusinoe Ozero Group
|
{{Flag|Russia}}<br>({{Flag|Buryatia}})
|
A [[Jurodidae|Jurodid beetle]].
|
|-
|
''[[Kirejtomma]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Cai |first2=C.-Y. |year=2021 |title=Revisiting the morphology of the Cretaceous ommatid beetle ''Clessidromma palmeri'' (Coleoptera: Archostemata: Ommatidae) |journal=Papéis Avulsos de Zoologia |volume=61 |pages=e20216195 |doi=10.11606/1807-0205/2021.61.95 |s2cid=240196850 |doi-access=free }}</ref>
|
Gen. et comb. nov
|
Valid
|
Li & Cai
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Ommatidae|Ommatid beetle]]<br/> The type species is  ''"[[Clessidromma]]" zengi'' ([[2020 in paleoentomology|2020]]).
|
[[File:Kirejtomma zengi holotype.png|thumb|center|upright|''[[Kirejtomma zengi]]'']]
|-
|
''[[Limnomma]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Tihelka |first2=E. |last3=Pang |first3=H. |last4=Huang |first4=D.-Y. |last5=Cai |first5=C.-Y. |title=''Limnomma'', a new genus of Ommatidae from the Middle Jurassic Daohugou beds (Coleoptera, Archostemata) |journal=Deutsche Entomologische Zeitschrift |year=2021 |volume=68 |issue=2 |pages=299–308 |doi=10.3897/dez.68.71880 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai ''in'' Li ''et al.''
|
[[Middle Jurassic]]
|
[[Haifanggou Formation]]
|
{{Flag|China}}
|
An [[Ommatidae|Ommatid beetle]].<br/> The type species is ''L. daohugouense''.
|
[[File:Limnomma daohugouense holotype.jpg|thumb|center|upright|''[[Limnomma daohugouense]]'']]
|-
|
''[[Omma axsmithi]]''<ref>{{cite book |last1=Jarzembowski |first1=E. A. |last2=Zheng |first2=D. |last3=Zhao |first3=X. |year=2021 |chapter=Is the beetle ''Omma'' (Insecta: Coleoptera) a living fossil? |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |volume=521 |pages=161–169 |publisher=The Geological Society of London |doi=10.1144/SP521-2021-56 |s2cid=244091355 }}</ref>
|
Sp. nov
|
In press
|
Jarzembowski, Zheng & Zhao
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An ''[[Omma]]'' species [[Ommatidae|Ommatid beetle]].
|
|-
|
''[[Omma forte]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Huang |first2=D.-Y. |last3=Cai |first3=C.-Y. |title=New species of ''Omma'' Newman from mid-Cretaceous Burmese amber (Coleoptera, Archostemata, Ommatidae) |year=2021 |journal=Deutsche Entomologische Zeitschrift |volume=68 |issue=2 |pages=341–348 |doi=10.3897/dez.68.74174 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Li & Cai<br/> ''in'' Li, Huang & Cai
|
Cretaceous <br/>(Albian to Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
An ''[[Omma]]'' species [[Ommatidae|Ommatid beetle]].
|
|-
|
''[[Paraodontomma|Paraodontomma leptocristatum]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Yamamoto |first2=S. |last3=Huang |first3=D.-Y. |last4=Cai |first4=C.-Y. |year=2021 |title=New species of ''Paraodontomma'' from mid-Cretaceous Burmese amber with muscle tissue preservation (Coleoptera: Archostemata: Ommatidae) |journal=Papéis Avulsos de Zoologia |volume=61 |pages=e20216153 |doi=10.11606/1807-0205/2021.61.53 |s2cid=236224305 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Li, Yamamoto & Cai ''in'' Li ''et al.''
|
Cretaceous (Albian to Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Ommatidae|Ommatid beetle]].
|
[[File:Paraodontomma leptocristatum.jpg|thumb|center|upright|''[[Paraodontomma leptocristatum]]'']]
|-
|
''[[Permocatinus]]''<ref name=Permocatinus>{{Cite journal |last=Ponomarenko |first=A. G. |title=Permian triaplids (Coleoptera, Triaplidae), yet another group of Mesozoic beetles from the terminal Permian |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=650–661 |doi=10.1134/S0031030121050130 |s2cid=245010239 |url=https://www.elibrary.ru/item.asp?id=46621796 |access-date=2021-10-15 |archive-date=2021-10-28 |archive-url=https://web.archive.org/web/20211028171149/https://www.elibrary.ru/item.asp%3Fid%3D46621796 |url-status=dead }}</ref>
|
Gen. et sp. nov
|
Valid
|
Ponomarenko
|
Late Permian
|
|
{{Flag|Russia}}
|
A [[Catiniidae (beetle)|Coleocatiniid beetle]].<br/> The type species is ''P. tomiensis''.
|
|-
|
''[[Tomiaplus]]''<ref name=Permocatinus />
|
Gen. et comb. et sp. nov
|
Valid
|
Ponomarenko
|
Late Permian
|
|
{{Flag|Russia}}
|
A [[Triaplidae|Triaplid beetle]].<br/> The type species is ''"[[Triaplus]]" sibiricus'' ([[2013 in paleontology|2013]]), <br/>as well as new species ''T. minimus''.
|
|-
|
''[[Tunguskagyrus|Tunguskagyrus уani]]''<ref name=Permocatinus />
|
Sp. nov
|
Valid
|
Ponomarenko
|
Late Permian
|
|
{{Flag|Russia}}
|
A [[Triaplidae|Triaplid beetle]].
|
|-
|
''[[Zygadenia|Zygadenia alexrasnitsyni]]''<ref>{{Cite journal|last1=Strelnikova |first1=O. D. |last2=Yan |first2=E. V. |title=Redescriptions of beetles of the ''Notocupes'' generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=499–514 |doi=10.11646/palaeoentomology.4.5.15 |s2cid=240527664 }}</ref>
|
Sp. nov
|
Valid
|
Strelnikova & Yan
|
Early Cretaceous
|
[[Zaza Formation]]
|
{{Flag|Russia}}
|
An [[Ommatidae|Ommatid beetle]].
|
|-
|}

====Myxophaga====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Triamyxa]]''<ref>{{Cite journal|last1=Qvarnström |first1=M. |last2=Fikáček |first2=M. |last3=Wernström |first3=J. V. |last4=Huld |first4=S. |last5=Beutel |first5=R. G. |last6=Arriaga-Varela |first6=E. |last7=Ahlberg |first7=P. E. |last8=Niedźwiedzki |first8=G. |date=June 30, 2021|title=Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin|journal=Current Biology |volume=31 |issue=15 |pages=3374–3381.e5 |doi=10.1016/j.cub.2021.05.015 |pmid=34197727|s2cid=235691750 |doi-access=free }}</ref>
|
Fam., Gen. et sp. nov
|
Valid
|
Qvarnström ''et al.''
|
[[Late Triassic]] ([[Carnian]])
|
Keuper succession (Drawno Beds)
|
{{Flag|Poland}}
|
A [[Triamyxidae|Triamyxid beetle]] in the suborder [[Myxophaga]].<br/> The type species is ''T. coprolithica''.
|
[[File:Triamyxa coprolithica type specimens.jpg|thumb|center|upright|''[[Triamyxa coprolithica]]'']]
|-
|}

====Polyphaga====

=====Bostrichiformia=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Archaenosodendron]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Tihelka |first2=E. |last3=Dahan |first3=L. |last4=Huang |first4=D.-Y. |last5=Cai |first5=C.-Y. |title=On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea) |year=2021 |journal=Zootaxa |volume=5082 |issue=3 |pages=223–244 |doi=10.11646/zootaxa.5082.3.2 |pmid=35390968 |s2cid=245319655 |doi-access=free }}</ref>
|
Gen. comb. et 3 sp. nov
|
Valid
|
Li & Cai ''in'' Li ''et al.''
|
Cretaceous (Albian to Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Nosodendridae|nosodendrid beetle]].<br/>The type species is ''"[[Nosodendron]]" cretaceum'';<br/> genus also includes new species ''A. angulare'', ''A. explanatum'' and ''A. remotidens''.
|
|-
|
''[[Cretomegatoma]]''<ref name=Cretomegatoma>{{Cite journal|last=Háva |first=J. |title=Two new genera and species of Dermestidae (Coleoptera) from Cretaceous Burmese amber |year=2021 |journal=Euroasian Entomological Journal |volume=20 |issue=6 |pages=343–345 |doi=10.15298/euroasentj.20.6.08 |s2cid=247374410 |url=https://www.researchgate.net/publication/357395546 |doi-access=free }}</ref>
|
Gen. et comb. nov
|
Valid
|
Háva
|
Late Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Dermestidae|carpet beetle]].<br/> The type species is ''"[[Megatoma]]" atypica'' ([[2017 in paleoentomology|2017]]).
|
|-
|
''[[Mesonosa]]''<ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Li |first2=Y.-D. |last3=Huang |first3=D.-Y. |last4=Cai |first4=C.-Y. |title=Are wounded-tree beetles living fossils? A new nosodendrid genus from Burmese amber with bilobed tarsi (Coleoptera: Nosodendridae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=50–56 |doi=10.11646/palaeoentomology.4.1.10 |s2cid=233894693 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tihelka ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Nosodendridae|wounded-tree beetle]].<br/> The type species is ''M. scandens''.
|
|-
|
''[[Petalium bruteno]]''<ref name=BJC211Petalium>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |title=Two fossil species of Petaliini White (Coleoptera: Ptinidae: Dorcatominae) from Eocene Baltic amber |journal=Baltic Journal of Coleopterology |volume=21 |issue=1 |pages=11–18 |url=http://www.bjc.sggw.pl/viewart.php?bookid=42&artid=733 }}</ref>
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A ''[[Petalium]]'' species [[Ptinidae|death watch beetle]].
|
|-
|
''[[Petalium widewuto]]''<ref name=BJC211Petalium />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A ''[[Petalium]]'' species [[Ptinidae|death watch beetle]].
|
|-
|
''[[Trogoderma ainu]]''<ref>{{Cite journal|last1=Perkovsky |first1=E.E. |last2=Háva |first2=J. |last3=Zaitsev |first3=A.A. |title=The first finding of a skin beetle (Coleoptera, Dermestidae) from Sakhalinian amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=184–192 |doi=10.1134/S0031030121020118 |s2cid=233746555 |url=https://elibrary.ru/item.asp?id=44595947 }}</ref>
|
Sp. nov
|
Valid
|
Perkovsky, Háva & Zaitsev
|
Middle Eocene
|
[[Sakhalin amber]]
|
{{Flag|Russia}}
|
A ''[[Trogoderma]]'' species [[Dermestidae|carpet beetle]].
|
|-
|
''[[Tuberphradonoma]]''<ref name=Cretomegatoma />
|
Gen. et sp. nov
|
Valid
|
Háva
|
Late Cretaceous<br/>Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Dermestidae|carpet beetle]].<br/> The type species is ''T. burmitica''.
|
|-
|
''[[Xyletinus lobanovi]]''<ref>{{Cite journal|last1=Bukejs |first1=A. |last2=Alekseev |first2=V. I. |last3=Háva |first3=J. |title=A new species of ''Xyletinus'' Latreille, 1809 (Coleoptera: Ptinidae: Xyletininae) from Eocene Baltic amber, with a key to known fossil species |year=2021 |journal=Caucasian Entomological Bulletin |volume=17 |issue=1 |pages=179–184 |doi=10.23885/181433262021171-179184 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Bukejs, Alekseev & Háva
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A ''[[Xyletinus]]''species [[Ptinidae|death watch beetle]].
|
|-
|}

=====Cucujiformia=====
======New Cucujiformian taxa======
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Aberrokorynetes|Aberrokorynetes oceanojubilaei]]''<ref>{{Cite journal|last1=Kolibáč |first1=J. |last2=Alekseev |first2=V. I. |last3=Kairišs |first3=K. |last4=Bukejs |first4=A. |title=Systematic placement and new data on the checkered beetles ''Aberrokorynetes'' Winkler and ''Visokorynetes'' Winkler (Coleoptera: Cleridae) from Eocene Baltic amber obtained from X-ray tomography |journal=Historical Biology |year=2022 |volume=34 |issue=10 |pages=1972–1980 |doi=10.1080/08912963.2021.1994561 |s2cid=240056175 }}</ref>
|
Sp. nov
|
In press
|
Kolibáč ''et al.''
|
[[Eocene]]
|
[[Baltic amber]]
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A [[Korynetinae]] [[Cleridae|checkered beetle]].
|
|-
|
''[[Aliattalus]]''<ref>{{Cite journal|last=Tshernyshev |first=S. E. |title=A new genus and species of soft-winged flower beetles (Coleoptera, Malachiidae) in Baltic amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=1 |pages=90–95 |doi=10.1134/S0031030121010056 | issn=0031-0301 |s2cid=232070647 |url=https://www.elibrary.ru/item.asp?id=44492499 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tshernyshev
|
Late Eocene
|
Baltic amber
|
{{flag|Europe}}
|
A [[Malachiinae]] [[Melyridae|malachite beetle]].<br/> The type species is ''A. intercalaris''.
|
|-
|
''[[Archaeosciaphilus|Archaeosciaphilus gyrommatus]]''<ref>{{Cite journal|last1=Legalov |first1=A. A. |last2=Poinar |first2=G. |title=A new species of the genus ''Archaeosciaphilus'' Legalov, 2012 (Coleoptera: Сurculionidae) with a list of Entiminae weevils described from Baltic amber |journal=Historical Biology |year=2022 |volume=34 |issue=11 |pages=2218–2223 |doi=10.1080/08912963.2021.2007530 |s2cid=252905207 }}</ref>
|
Sp. nov
|
In press
|
Legalov & Poinar
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A [[Polydrusini]] [[Entiminae]] [[Curculionidae|broad nosed weevil]].
|
|-
|
''[[Arkagalarhinus]]''<ref>{{Cite journal|last=Legalov |first=A. A. |title=First record of the family Chilecarinae (Coleoptera; Ithyceridae) from the Upper Cretaceous Arkagala Formation, Magadan Region, Russian Far East |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 105001 |doi=10.1016/j.cretres.2021.105001 |bibcode=2021CrRes.12805001L }}</ref>
|
Gen. et sp. nov
|
In press
|
Legalov
|
Late Cretaceous<br/>([[Santonian]]-[[Campanian]])
|
Arkagala Formation
|
{{Flag|Russia}}
|
A [[Chilecarinae]] [[Ithyceridae|ithycerid weevil]].<br/> The type species is ''A. zherikhini''.
|
|-
|
''[[Asiomira dubrovinae]]''<ref>{{Cite journal|last1=Nabozhenko |first1=M. V. |last2=Bukejs |first2=A. |title=The first fossilized comb-clawed beetle of the genus ''Asiomira'' Dubrovina, 1973 (Coleoptera: Tenebrionidae: Alleculinae) from Baltic Amber and notes on the distribution of extant species of the genus |year=2021 |journal=Zootaxa |volume=5082 |issue=2 |pages=177–184 |doi=10.11646/zootaxa.5082.2.7 |pmid=35390973 |s2cid=245227981 }}</ref>
|
Sp. nov
|
Valid
|
Nabozhenko & Bukejs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
An [[Alleculinae]] [[darkling beetle]].
|
|-
|
''[[Aulacoscelis santanensis]]''<ref>{{Cite journal|last=Legalov |first=A. A. |year=2021 |title=First record of orsodacnid leaf beetle (Coleoptera: Orsodacnidae) from the Lower Cretaceous of Brazil |journal=Historical Biology |volume=34 |issue=3 |pages=453–457 |doi=10.1080/08912963.2021.1929204 |s2cid=236395989 }}</ref>
|
Sp. nov
|
In press
|
Legalov
|
[[Early Cretaceous]]<br/>([[Aptian]]-Albian)
|
[[Santana Group]]
|
{{Flag|Brazil}}
|
An [[Orsodacnidae|orsodacnid leaf beetle]].
|
|-
|
''[[Baltocar|Baltocar sontagae]]''<ref name=FRBukejsLegalov>{{Cite journal|last1=Bukejs |first1=A. |last2=Legalov |first2=A. A. |title=Two new species of the family Rhynchitidae (Coleoptera: Curculionoidea) from Eocene Baltic amber, with key to species and assumed trophic relationships |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=117–127 |doi=10.5194/fr-24-117-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Bukejs & Legalov
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Sayrevilleinae]] [[Rhynchitidae|tooth-nosed snout weevil]].
|
|-
|
''[[Baltosidis]]''<ref>{{Cite journal|last=Szawaryn |first=K. |title=The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles |year=2021 |journal=Zoological Journal of the Linnean Society |volume=193 |issue=4 |pages=1294–1309 |doi=10.1093/zoolinnean/zlaa180 |doi-access=free }}</ref>
|
Gen. et 3 sp. nov
|
Valid
|
Szawaryn
|
Eocene
|
Baltic amber
|
{{Flag|Europe}}
|
A [[Microweiseini]] [[Microweiseinae]] [[Coccinellidae|lady beetle]].<br/> The type species is ''B. damzeni'';<br/> genus also includes ''B. damgaardi'' and ''B. szadziewskii''.
|
|-
|
''[[Bronchotibia]]''<ref>{{Cite journal|last1=Poinar |first1=G. |last2=Legalov |first2=A. A. |year=2021 |title=First record of the tribe Lymantini (Coleoptera: Curculionidae) from Dominican amber |journal=Historical Biology |volume=34 |pages=67–71 |doi=10.1080/08912963.2021.1893717 |s2cid=242522689 }}</ref>
|
Gen. et sp. nov
|
In press
|
Poinar & Legalov
|
Miocene<br/>(Burdigalian)
|
[[Dominican amber]]
|
{{Flag|Dominican Republic}}
|
A [[Lymantini]] [[Molytinae]] [[Curculionidae|weevil]].<br/> The type species is ''B. adunatus''.
|
|-
|
''[[Cathartosilvanus perkovskyi]]''<ref>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |title=''Cathartosilvanus perkovskyi'' sp. nov. - first record of Silvanidae (Coleoptera: Cucujoidea) from Eocene Rovno amber |journal=Baltic Journal of Coleopterology |volume=21 |issue=1 |pages=19–24 |url=http://www.bjc.sggw.pl/viewart.php?bookid=42&artid=734 }}</ref>
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A [[Silvanidae|silvanid beetle]].
|
|-
|
''[[Ceutorhynchus zerovae]]''<ref>{{Cite journal|last1=Legalov |first1=A. A. |last2=Nazarenko |first2=V. Yu. |last3=Vasilenko |first3=D. V. |last4=Perkovsky |first4=E. E. |title=''Ceutorhynchus'' Germar (Coleoptera, Curculionidae) as proxy for Eocene core Brassicaceae: first record of the genus from Rovno amber |year=2021 |journal=Journal of Paleontology |volume= 96|issue=2 |pages=379–386 |doi=10.1017/jpa.2021.82 |s2cid=240546489 }}</ref>
|
Sp. nov
|
Valid
|
Legalov ''et al.''
|
Late Eocene
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
A [[Curculionidae|weevil]].
|
|-
|
''[[Coslonatus]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Huang |first2=D.-Y. |last3=Cai |first3=C.-Y. |title=An aberrant colydiine-like tenebrionoid beetle from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea: Zopheridae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=614–619 |doi=10.11646/palaeoentomology.4.6.10 |s2cid=245201548 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai 
|
[[Cenomanian]]
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Colydiinae]]? [[Zopheridae]] [[Tenebrionoidea|tenebrionoid beetle]].<br/> The type species is ''C. rasnitsyni''.
|
|-
|
''[[Cretaidgia]]''<ref>{{Cite journal|last1=Zhao |first1=Y. |last2=Liu |first2=Z. |last3=Yu |first3=Y. |last4=Shih |first4=C. |last5=Ślipiński |first5=A. |last6=Ren |first6=D. |title=First species of Prionoceridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Cleroidea) |year=2021 |journal=Cretaceous Research |volume=133 |pages=Article 105118 |doi=10.1016/j.cretres.2021.105118 |s2cid=245058205 }}</ref>
|
Gen. et sp. nov
|
In press
|
Zhao, Liu & Yu
|
Late Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Prionoceridae|prionocerid beetle]].<br/> The type species is ''C. burmensis''.
|
|-
|
''[[Diapus ethiopicus]]''<ref name=PalaeoworldDiapus>{{Cite journal|last1=Solórzano-Kraemer |first1=M. M. |last2=Hammel |first2=J. U. |last3=Kunz |first3=R. |last4=Xu |first4=C.-P. |last5=Cognato |first5=A. I. |title=Miocene pinhole borer ambrosia beetles: new species of ''Diapus'' (Coleoptera: Curculionidae: Platypodinae) |journal=Palaeoworld |date=2023 |issue=4 |pages=658–668 |volume=32 |doi=10.1016/j.palwor.2021.10.001 |s2cid=240104595 }}</ref>
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
[[Ethiopian amber]]
|
{{Flag|Ethiopia}}
|
A [[Platypodinae]] [[Diapus|pinhole borer ambrosia]] [[Curculionidae|weevil]].
|
|-
|
''[[Diapus resinae]]''<ref name=PalaeoworldDiapus />
|
Sp. nov
|
In press
|
Solórzano-Kraemer ''et al.''
|
Miocene
|
[[Zhangpu amber]]
|
{{Flag|China}}
|
A [[Platypodinae]] [[Diapus|pinhole borer ambrosia]] [[Curculionidae|weevil]].
|
|-
|
''[[Dorytomus mikhailovi]]''<ref>{{Cite journal|last1=Legalov |first1=A. A. |last2=Nazarenko |first2=V. Y. |last3=Perkovsky |first3=E. E. |year=2021 |title=A new species of the genus ''Dorytomus'' Germar, 1817 (Coleoptera, Curculionidae) from Rovno amber |journal=Zootaxa |volume=5006 |issue=1 |pages=95–100 |doi=10.11646/zootaxa.5006.1.12|issn=1175-5326 |pmid=34810584 |s2cid=242796435 }}</ref>
|
Sp. nov
|
Valid
|
Legalov, Nazarenko & Perkovsky
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A species of ''[[Dorytomus]]'' [[Curculioninae]] [[weevil]].
|
|-
|
''[[Electroxylita]]''<ref name=Electroxylita>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |year=2021 |title=Fossil Melandryidae (Coleoptera: Tenebrionoidea) from Eocene Baltic amber of the Sambian peninsula: new genus, six new species, new records and key to described taxa |journal=Zootaxa |volume=4965 |issue=1 |pages=142–166 |doi=10.11646/zootaxa.4965.1.7 |pmid=33903499 |s2cid=233408753 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Melandryidae|false darkling beetle]].<br/> The type species is ''E. chronographica''.
|
|-
|
''[[Eurocrossotus]]''<ref>{{Cite journal|last=Vitali |first=F. |title=''Eurocrossotus alekseevi'' gen. n., sp. n. (Coleoptera: Cerambycidae) from Baltic amber, the first fossil member of the tribe Crossotini Thomson, 1864 |journal=Baltic Journal of Coleopterology |volume=21 |issue=2 |pages=221–228 |url=http://www.bjc.sggw.pl/viewart.php?bookid=43&artid=764 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Vitali
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[longhorn beetle]] belonging to the tribe [[Crossotini]]. The type species is ''E. alekseevi''.
|
|-
|
''[[Glaesotropis rohdendorfi]]''<ref>{{Cite journal|last1=Legalov |first1=A. A. |last2=Nazarenko |first2=V. Y. |last3=Perkovsky |first3=E. E. |title=A new species of the genus ''Glaesotropis'' Gratshev and Zherikhin, 1995 (Coleoptera, Anthribidae) from Rovno amber |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=1–7 |doi=10.5194/fr-24-1-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Legalov, Nazarenko & Perkovsky
|
Eocene<br/>([[Priabonian]])
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
A [[zygaenodini]]n [[Anthribinae|anthribine]] [[Anthribidae|anthribid fungus weevil]].
|
|-
|
''[[Glesoconomorphus|Glesoconomorphus ekaterinae]]''<ref>{{Cite journal|last1=Telnov |first1=D. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |last4=Yamamoto |first4=S. |title=The first fossil Coleoptera record from the Volyn Region, Ukraine, with description of a new ''Glesoconomorphus'' (Coleoptera, Mycteridae) in syninclusion with Winterschmidtiidae (Acari) and a key to species |year=2021 |journal=ZooKeys |issue=1068 |pages=189–201 |doi=10.3897/zookeys.1068.75391 |pmid=34819765 |pmc=8592962 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Telnov ''et al.''
|
Late Eocene<br/>Priabonian
|
Rovno amber
|
{{Flag|Ukraine}}
|
A [[Mycteridae|mycterid palm beetle]].
|
|-
|
''[[Gracilenticrus]]''<ref>{{Cite journal|last1=Yu |first1=Y.-L. |last2=Li |first2=Y.-D. |last3=Kolibáč |first3=J. |last4=Ślipiński |first4=A. |last5=Ren |first5=D. |last6=Pang |first6=H. |last7=Li |first7=Z.-Q. |title= A New Genus and Species of Lophocateridae from Mid-Cretaceous Amber of Myanmar (Coleoptera) |year=2021 |journal=Insects |volume=12 |issue=12 |pages=Article 1052 |doi=10.3390/insects12121052 |pmid=34940140 |pmc=8709423 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Yu, Kolibáč & Ślipiński<br/> ''in'' Yu ''et al.''
|
Late Cretaceous<br/> (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Lophocateridae|lophocaterid beetle]].<br/> The type species is ''G. burmiticus''.
|
|-
|
''[[Antirhelus|Heterhelus buzina]]''<ref>{{Cite journal|last1=Kupryjanowicz |first1=J. |last2=Lyubarsky |first2=G. Yu. |last3=Perkovsky |first3=E. E. |title=''Heterhelus buzina'' sp.n. (Coleoptera: Kateretidae) from Rovno amber: the first proxy for ''Sambucus'' in the Eocene of Eastern Europe |year=2021 |journal=Invertebrate Zoology |volume=18 |issue=1 |pages=16–24 |doi=10.15298/invertzool.18.1.02 |s2cid=234840368 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Kupryjanowicz, Lyubarsky & Perkovsky
|
Eocene
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
A [[Kateretidae|Kateretid short-winged flower beetle]]. Originally described as a species of ''[[Heterhelus]]'', but Kirejtshuk, Jenkins Shaw & Smirnov (2023) transferred it to the genus ''Antirhelus''.<ref>{{Cite journal|last1=Kirejtshuk |first1=A. G. |last2=Jenkins Shaw |first2=J. |last3=Smirnov |first3=I. S. |year=2023 |title=A New Subgenus of the Genus ''Phenolia'' (Coleoptera, Nitidulidae) from Myanmar Cretaceous Amber with Taxonomic, Phylogenetic and Bionomic Notes on the 'Nitidulid' Group of Families |journal=Insects |volume=14 |issue=7 |at=647 |doi=10.3390/insects14070647 |pmid=37504653 |pmc=10380190 |doi-access=free }}</ref>
|
|-
|
''[[Isomira lobanovi]]''<ref>{{Cite journal|last1=Nabozhenko |first1=M. V. |last2=Bukejs |first2=A. |title=A new species and a key to ''Isomira'' Mulsant, 1856 (Coleoptera: Tenebrionidae: Alleculinae) from Eocene Baltic amber |year=2021 |journal=Caucasian Entomological Bulletin |volume=17 |issue=1 |pages=51–56 |doi=10.23885/181433262021171-5156 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Nabozhenko & Bukejs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
An ''[[Isomira]]'' species [[Alleculinae|claw comb]] [[darkling beetle]].
|
|-
|
''[[Kaszubia]]''<ref>{{Cite journal |last1=Legalov |first1=A. A. |last2=Kupryjanowicz |first2=J. |last3=Perkovsky |first3=E. E. |title=A new genus of the tribe Cossonini (Coleoptera: Curculionidae) in Baltic amber (Poland) |year=2021 |journal=Paleontological Journal |volume=55 |issue=4 |pages=405–409 |doi=10.1134/S0031030121040109 |s2cid=237378812 |url=https://elibrary.ru/item.asp?id=46227217 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Legalov, Kupryjanowicz & Perkovsky
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Poland}}
|
A [[Cossonini|cossonine]] [[Curculionidae|weevil]].<br/> The type species is ''K. wanati''.
|
|-
|
''[[Kulindrobor]]''<ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Yamamoto |first2=S. |last3=Huang |first3=D. |last4=Cai |first4=C. |title=Enigmatic tenebrionoid beetles from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea) |year=2021 |journal=Cretaceous Research |volume=123 |pages=Article 104779 |doi=10.1016/j.cretres.2021.104779 |bibcode=2021CrRes.12304779T |s2cid=233908153 }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Tihelka ''et al.''
|
Late Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Tenebrionoidea|tenebrionoid beetle]] of uncertain phylogenetic placement.<br/> Genus includes new species ''K. enigmaticus'' and ''K. magnus''.
|
|-
|
''[[Lobanoviella]]''<ref>{{Cite journal|last1=Kirejtshuk |first1=A. G. |last2=Reid |first2=C. A. M. |title=''Lobanoviella andreyi'', new genus and species of Lobanoviellini trib. nov. in the subfamily Palophaginae (Coleoptera, Megalopodidae) from the Eocene Baltic amber, with discussion of similarity with Phloeostichidae |journal=Historical Biology |year=2022 |volume=34 |issue=9 |pages=1894–1900 |doi=10.1080/08912963.2021.1987428 |s2cid=243581689 |url=https://www.tandfonline.com/doi/abs/10.1080/08912963.2021.1987428 }}</ref>
|
Gen. et sp. nov
|
In press
|
Kirejtshuk & Reid
|
Eocene
|
Baltic amber
|
{{flag|Europe}}
|
A [[Lobanoviellini|lobanovielline]] [[Palophaginae|palophagine]] [[Megalopodidae|megalopodid leaf beetle]].<br/> The type species is ''L. andreyi''.
|
|-
|
''[[Madelinia|Madelinia capillata]]''<ref name=Electroxylita />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Melandryidae|false darkling beetle]].
|
|-
|
''[[Melanosiagon]]''<ref name=Melanosiagon>{{Cite journal|last1=Batelka |first1=J. |last2=Prokop |first2=J. |title=The earliest beetle with mouthparts specialized for feeding on nectar is a parasitoid of mid-Cretaceous Hymenoptera |year=2021 |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 207 |doi=10.1186/s12862-021-01930-6 |pmid=34809578 |pmc=8607574 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Batelka & Prokop 
|
Late Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Ripiphoridae|wedge-shaped beetle]].<br/> The type species is ''M. serraticornis''.
|
|-
|
''[[Metaclisa ottoi]]''<ref>{{Cite journal|last1=Nabozhenko |first1=M. V. |last2=McKellar |first2=R. C. |last3=Bukejs |first3=A. |year=2021 |title=The first described darkling beetle of the tribe Metaclisini (Coleoptera: Tenebrionidae) from Eocene Baltic amber |journal=Zootaxa |volume=4999 |issue=3 |pages=279–284 |doi=10.11646/zootaxa.4999.3.7 |pmid=34810485 |s2cid=237770026 }}</ref>
|
Sp. nov
|
Valid
|
Nabozhenko, McKellar & Bukejs
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A''[[Metaclisa]]'' species [[Metaclisini|metaclisine]] [[darkling beetle]].
|
|-
|
''[[Microscapha kugelanni]]''<ref name=Electroxylita />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Melandryidae|false darkling beetle]].
|
|-
|
''[[Microtrogossita]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Tihelka |first2=E. |last3=Leschen |first3=R. A. B. |last4=Yu |first4=Y. |last5=Ślipiński |first5=A. |last6=Pang |first6=H. |last7=Huang |first7=D. |last8=Kolibáč |first8=J. |last9=Cai |first9=C. |title=An exquisitely preserved tiny bark-gnawing beetle (Coleoptera: Trogossitidae) from mid-Cretaceous Burmese amber and the phylogeny of Trogossitidae |year=2021 |journal=Journal of Zoological Systematics and Evolutionary Research |volume= 59|issue=8 |pages=1939–1950 |doi=10.1111/jzs.12515 |s2cid=238712220 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai<br/> ''in'' Li ''et al.''
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Trogossitidae|Trogossitid bark-gnawing beetle]].<br/> The type species is ''M. qizhihaoi''.
|
|-
|
''[[Monolepta rappsilberi]]''<ref>{{Cite journal|last1=Bukejs |first1=A. |last2=Bezděk |first2=J. |last3=Kairišs |first3=K. |last4=Alekseev |first4=V. I. |year=2021 |title=The first extinct species of ''Monolepta'' Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) from Bitterfeld amber, described using X-ray micro-computed tomography |journal=Zootaxa |volume=4995 |issue=2 |pages=345–356 |doi=10.11646/zootaxa.4995.2.7 |pmid=34810567 |s2cid=237740153 }}</ref>
|
Sp. nov
|
Valid
|
Bukejs ''et al.''
|
Late Eocene<br/>Priabonian
|
[[Bitterfeld amber]]
|
{{Flag|Germany}}
|
A ''[[Monolepta]]'' species [[Chrysomelidae|skeletonizing leaf beetle]].
|
|-
|
''[[Neolatitergum]]''<ref>{{Cite journal|last=Zhang |first=Y. |year=2021 |title=New replacement name for the genus ''Latitergum'' Yu, Ślipiński, Leschen, Ren & Pang, 2014 (Insecta: Coleoptera: Trogossitidae) |journal=Zootaxa |volume=4980 |issue=2 |pages=395–396 |doi=10.11646/zootaxa.4980.2.11 |pmid=34186972 |s2cid=235687049 }}</ref>
|
Nom. nov
|
Valid
|
Zhang
|
[[Jurassic]]
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A [[Trogossitidae|Trogossitid bark-gnawing beetle]].<br/> A replacement name for ''"[[Latitergum]]"'' Yu ''et al.'' ([[2014 in paleontology|2014]]).
|
|-
|
''[[Oedemera girulskii]]''<ref>{{Cite journal|last1=Szawaryn |first1=K. |last2=Sontag |first2=E. |last3=Kubisz |first3=D. |title=The first described fossil Oedemeridae (Insecta: Coleoptera) from Baltic amber |year=2021 |journal=Journal of Paleontology |volume= 96|issue=2 |pages=375–378 |doi=10.1017/jpa.2021.102 |s2cid=242027889 }}</ref>
|
Sp. nov
|
Valid
|
Szawaryn, Sontag & Kubisz
|
Late Eocene<br/> Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
An ''[[Oedemera]]'' species [[Oedemeridae|false blister beetle]].
|
|-
|
''[[Pacyclaxyra]]''<ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Huang |first2=D. |last3=Cai |first3=C. |title=A new genus of giant sooty mould beetles from mid-Cretaceous Burmese amber (Coleoptera: Cyclaxyridae) |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104876 |doi=10.1016/j.cretres.2021.104876 |bibcode=2021CrRes.12504876T }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tihelka, Huang & Cai
|
Late Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Cyclaxyridae|sooty mould beetle]].<br/> The type species is ''P. azari''.
|
|-
|
''[[Palaeatalasis]]''<ref>{{Cite journal|last=Legalov |first=A. A. |title=First record of the subfamily Sagrinae (Coleoptera: Chrysomelidae) from the Eocene of North America |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=135–139 |doi=10.5194/fr-24-135-2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Legalov
|
Eocene<br/> ([[Ypresian]])
|
[[Green River Formation]]
|
{{Flag|United States}}<br>({{Flag|Utah}})
|
A [[Megamerini]] [[Sagrinae]] [[leaf beetle]].<br/> The type species is ''P. monrosi''.
|
|-
|
''[[Palaeoanoplus]]''<ref>{{Cite journal|last=Legalov |first=A.A. |title=New tribe Palaeoanoplini trib. nov. (Coleoptera: Curculionidae) in Baltic amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=179–183 |doi=10.1134/S0031030121020076 |issn=0031-0301  |s2cid=233746782 |url=https://elibrary.ru/item.asp?id=44595946 }}</ref>
|
Trib., Gen., et sp. nov
|
Valid
|
Legalov
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Palaeoanoplini]] [[Curculioninae]] [[Curculionidae|weevil]].<br/> The type species is ''P. horridus''.
|
|-
|
''[[Paleopegorhinus]]''<ref>{{Cite journal|last1=Poinar |first1=G. O. |last2=Brown |first2=A. E. |last3=Legalov |first3=A. A. |title=First record of the subfamily Chilecarinae (Coleoptera; Ithyceridae) from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104793 |doi=10.1016/j.cretres.2021.104793 |bibcode=2021CrRes.12404793P |s2cid=233806293 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar, Brown & Legalov
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Chilecarinae]] [[Ithyceridae|new york weevil]].<br/> The type species is ''P. micrommatus''.
|
|-
|
''[[Palaeopeplus]]''<ref>{{Cite journal|last1=Powell |first1=G. S. |last2=Cline |first2=A. R. |title=The First Cillaeinae (Coleoptera: Nitidulidae) Described from Amber |year=2021 |journal=Annales Zoologici |volume=71 |issue=1 |pages=21–26 |doi=10.3161/00034541ANZ2021.71.1.003 |s2cid=232422254 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Powell & Cline
|
Miocene<br/>([[Burdigalina]])
|
[[Dominican amber]]
|
{{Flag|Dominican Republic}}
|
A [[Cillaeinae]] [[sap beetle]].<br/> The type species is ''P. cascus''.
|
|-
|
''[[Pelretes]]''<ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Li |first2=L. |last3=Fu |first3=Y. |last4=Su |first4=Y. |last5=Huang |first5=D. |last6=Cai |first6=C. |title=Angiosperm pollinivory in a Cretaceous beetle |year=2021 |journal=Nature Plants |volume=7 |issue=4 |pages=445–451 |doi=10.1038/s41477-021-00893-2 |pmid=33846595 |s2cid=233223519 }}</ref><ref>{{Cite journal|last1=Bao |first1=T. |last2=Wedmann |first2=S. |last3=Grímsson |first3=F. |last4=Beutel |first4=R. G. |last5=Seyfullah |first5=L. |last6=Bao |first6=L. |last7=Jarzembowski |first7=E. A. |title=Was the kateretid beetle ''Pelretes'' really a Cretaceous angiosperm pollinator? |year=2021 |journal=Nature Plants |volume=8 |issue=1 |pages=38–40 |doi=10.1038/s41477-021-01044-3 |pmid=34949805 |s2cid=245471963 }}</ref><ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Li |first2=L. |last3=Fu |first3=Y. |last4=Su |first4=Y. |last5=Huang |first5=D. |last6=Cai |first6=C. |title=Reply to: ''Pelretes vivificus'' was a pollinator of Cretaceous angiosperms |year=2021 |journal=Nature Plants |volume=8 |issue=1 |pages=41–44 |doi=10.1038/s41477-021-01045-2 |pmid=34949806 |s2cid=245445815 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tihelka ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
Originally described as a [[Kateretidae|short-winged flower beetle]];<br/> subsequently argued to be a [[sap beetle]] belonging to the subfamily Apophisandrinae<ref>{{Cite journal|last1=Peris |first1=D. |last2=Jelínek |first2=J. |last3=Sabatelli |first3=S. |last4=Liu |first4=M.-K. |last5=Peña-Kairath |first5=C. |last6=Zhao |first6=Q. |last7=Cai |first7=C.-Y. |last8=Kairišs |first8=K. |last9=Mähler |first9=B. |last10=Rühr |first10=P. T. |last11=Hammel |first11=J. U. |last12=Audisio |first12=P. |title=Archaic sap beetles (Coleoptera: Nitidulidae) as Cretaceous pollinators |year=2024 |journal=Palaeoentomology |volume=7 |issue=5 |pages=594–610 |doi=10.11646/palaeoentomology.7.5.4 }}</ref> or a member of the separate family Apophisandridae.<ref>{{Cite journal|last1=Kirejtshuk |first1=A. G. |last2=Jenkins Shaw |first2=J. |last3=Smirnov |first3=I. S. |year=2023 |title=A New Subgenus of the Genus ''Phenolia'' (Coleoptera, Nitidulidae) from Myanmar Cretaceous Amber with Taxonomic, Phylogenetic and Bionomic Notes on the 'Nitidulid' Group of Families |journal=Insects |volume=14 |issue=7 |at=647 |doi=10.3390/insects14070647 |pmid=37504653 |pmc=10380190 |doi-access=free }}</ref><br/> The type species is ''P. vivificus ''.
|
|-
|
''[[Phloiotrya inmarinata]]''<ref name=Electroxylita />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A ''[[Phloiotrya]]'' species [[Melandryidae|false darkling beetle]].
|
|-
|
''[[Protodasytes]]''<ref>{{Cite journal|last1=Tihelka |first1=E. |last2=Huang |first2=D. |last3=Perrichot |first3=V. |last4=Cai |first4=C. |year=2021 |title=A previously missing link in the evolution of dasytine soft-winged flower beetles from Cretaceous Charentese amber (Coleoptera, Melyridae) |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=1753–1764 |doi=10.1002/spp2.1360 |s2cid=234862770 |url=https://hal-insu.archives-ouvertes.fr/insu-03196023/file/Tihelka_etal_accepted.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tihelka ''et al.''
|
[[Late Cretaceous]]<br/>([[Cenomanian]])
|
[[Charentese amber]]
|
{{Flag|France}}
|
A [[Melyridae|soft-wing flower beetle]].<br/> The type species is ''P. cretaceus''.
| 
|-
|
''[[Protomauroania]]''<ref>{{Cite journal|last1=Tshernyshev |first1=S.E. |last2=Shcherbakov |first2=M.V. |title=A new genus and species of malachite beetles (Coleoptera, Dasytidae) in Baltic amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=174–178 |doi=10.1134/S0031030121020039 |s2cid=233746774 |url=https://elibrary.ru/item.asp?id=44595945 }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Tshernyshev ''in'' Tshernyshev & Shcherbakov
|
Late Eocene
|
Baltic amber<br>
Rovno amber
|
Baltic Sea region<br>
{{Flag|Ukraine}}
|
A [[Dasytinae]] [[Melyridae|malachite beetle]].<br/> The type species ''P. chaetophorata''<br/> Genus also includes ''P. mikhailovi''.<ref>{{Cite journal|last1=Tshernyshev |first1=S. E. |last2=Perkovsky |first2=E. E. |year=2021 |title=''Protomauroania mikhailovi''—a new species of malachite beetles (Coleoptera, Dasytidae) in Rovno amber |journal=Zootaxa |volume=5006 |issue=1 |pages=189–194 |doi=10.11646/zootaxa.5006.1.20 |pmid=34810576 |s2cid=243574205 }}</ref>
|
|-
|
''[[Protostomopsis]]''<ref>{{Cite journal|last1=Bukejs |first1=A. |last2=Ślipiński |first2=A. |last3=Mitchell |first3=J. L. |last4=McKellar |first4=R. C. |last5=Barbi |first5=M. |last6=Larsson |first6=H. C. E. |last7=Alekseev |first7=V. I. |year=2021 |title=First fossil representative of Cerylonidae (Coleoptera: Coccinelloidea) described using X-ray micro-computed tomography, from Eocene Baltic amber |journal=Zootaxa |volume=5032 |issue=2 |pages=225–236 |doi=10.11646/zootaxa.5032.2.4 |pmid=34811132 |s2cid=239630223 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Bukejs, Ślipiński & Alekseev<br/>''in'' Bukejs ''et al.''
|
Eocene
|
Baltic amber
|
{{Flag|Poland}}<br />
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A [[Cerylonidae|cerylonid beetle]]. The type species is ''P. pandema''.
|
|-
|
''[[Pseudomesauletes lobanovi]]''<ref name=FRBukejsLegalov />
|
Sp. nov
|
Valid
|
Bukejs & Legalov
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Rhynchitinae]] [[Rhynchitidae|tooth-nosed snout weevil]].
|
|-
|
''[[Pulchritudo]]''<ref>{{Cite journal|last1=Krell |first1=F.-T. |last2=Vitali |first2=F. |year=2021 |title=Attenborough's beauty: exceptional pattern preservation in a frog-legged leaf beetle from the Eocene Green River Formation, Colorado (Coleoptera, Chrysomelidae, Sagrinae) |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=2101–2112 |doi=10.1002/spp2.1398 |s2cid=238854571 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Krell & Vitali
|
[[Eocene]]<br/>(Ypresian)
|
[[Green River Formation]]
|
{{Flag|United States}}<br>({{Flag|Colorado}})
|
A [[Sagrinae]] [[leaf beetle]].<br/> The type species is ''P. attenboroughi''.
|
|-
|
''[[Rhamphophorus]]''<ref>{{Cite journal|last1=Poinar |first1=G. |last2=Brown |first2=A. E. |title=A new tribe, genus and species of weevil, ''Rhamphophorus legalovii'' gen. et sp. nov., (Coleoptera, Nemonychidae, Rhamphophorini tribe nov.) in Burmese amber |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104948 |doi=10.1016/j.cretres.2021.104948 }}</ref>
|
Trib., gen. et sp. nov
|
Valid
|
Poinar & Brown
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Rhamphophorini]] [[Cimberidinae]] [[Nemonychidae|pine flower weevil]].<br>The type species is ''R. legalovii''.
|
|-
|
''[[Salignacicola]]''<ref name=Melanosiagon />
|
Gen. et comb. nov
|
Valid
|
Batelka & Prokop 
|
Late Cretaceous<br/> (Cenomanian)
|
|
{{Flag|France}}
|
A [[Ripiphoridae|wedge shaped beetle]].<br> The type species is ''"[[Macrosiagon]]" ebboi'' ([[2004 in paleontology|2004]]).
|
|-
|
''[[Symphora glaesonauta]]''<ref name=Electroxylita />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Melandryidae|false darkling beetle]].
|
|-
|
''[[Symphora pollocki]]''<ref name=Electroxylita />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Middle Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Melandryidae|false darkling beetle]].
|
|-
|
''[[Teredolaemus primigenius]]''<ref>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |last3=Pankowski |first3=M. G. |last4=Ślipiński |first4=A. |year=2021 |title=The first representative of the family Teredidae (Coleoptera: Coccinelloidea) in the fossil record |journal=Historical Biology |volume= 34|issue=11 |pages=1–6 |doi=10.1080/08912963.2021.2009474 |s2cid=244917429 }}</ref>
|
Sp. nov
|
Valid
|
Alekseev ''et al.''
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A [[Teredidae|teredid beetle]].
|
|-
|
''[[Toxorhynchus michalskii]]''<ref name=Zherikhiniellus />
|
Sp. nov
|
Valid
|
Legalov
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Apioninae]] [[Brentidae|straight-snouted weevil]].
|
|-
|
''[[Trematosphindus]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Tihelka |first2=E. |last3=Liu |first3=Z.-H. |last4=Huang |first4=D.-Y. |last5=Cai |first5=C.-Y. |title=New mid-Cretaceous cryptic slime mold beetles and the early evolution of Sphindidae (Coleoptera: Cucujoidea) |year=2021 |journal=Arthropod Systematics & Phylogeny |volume=79 |pages=587–597 |doi=10.3897/asp.79.e72724 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai<br/> ''in'' Li ''et al.''
|
Cretaceous<br/> (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Sphindidae|slime mold beetle]].<br/> The type species is ''T. newtoni''
|
|-
|
''[[Zaiwa pankowskiorum|Zaiwa]]''<ref>{{Cite journal|last1=Lyubarsky |first1=G. Yu. |last2=Tihelka |first2=E. |last3=Cai |first3=C.-Y. |last4=Perkovsky |first4=E. E. |title=''Zaiwa'': A new genus of the family Lophocateridae from mid-Cretaceous Burmese amber (Coleoptera: Cleroidea) |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=218–222 |doi=10.11646/palaeoentomology.4.3.7 |issn=2624-2826  |s2cid=237862213 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Lyubarsky ''et al.''
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Lophocateridae|lophocaterid beetle]].<br/> The type species is ''Z. pankowskiorum''.
|
|-
|
''[[Zherikhiniellus]]''<ref name=Zherikhiniellus>{{Cite journal|last=Legalov |first=A. A. |title=New weevils of the family Brentidae (Coleoptera) in Baltic amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=1 |pages=96–100 |doi=10.1134/S003103012101007X |s2cid=232071010 |url=https://www.elibrary.ru/item.asp?id=44492500 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Legalov
|
late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Nanophyinae]] [[Brentidae|straight-snouted weevil]].<br/> The type species is ''Z. rarus''.
|
|-
|}

=====Cucujiformia research=====
* A piece of [[Burmese amber]] preserving the complete series of free-living stages (except of the last larval [[instar]]) of a Cretaceous  [[parasitoid]] beetle ''[[Paleoripiphorus]]'' is described by Batelka, [[Michael S. Engel|Engel]] & Prokop (2021).<ref>{{Cite journal|last1=Batelka |first1=J. |last2=Engel |first2=M. S. |last3=Prokop |first3=J. |title=The complete life cycle of a Cretaceous beetle parasitoid |year=2021 |journal=Current Biology |volume=31 |issue=3 |pages=R118–R119 |doi=10.1016/j.cub.2020.12.007 |pmid=33561406 |s2cid=231848712 |doi-access=free }}</ref>

=====Elateriformia=====
======New Elateriformian taxa======
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Amberophytum|Amberophytum maculatum]]''<ref>{{Cite journal|last1=Poinar |first1=G. O. |last2=Vega |first2=F. E. |last3=Legalov |first3=A. A. |title=A New Species of the Genus ''Amberophytum'' Yu, Slipinski et Pang, 2019 (Coleoptera: Cerophytidae) from mid-Cretaceous Burmese Amber |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=3 |pages=337–341 |doi=10.37819/biosis.002.03.0125 |s2cid=238697106 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
[[George Poinar Jr.|Poinar]], Vega & Legalov
|
[[Late Cretaceous]]<br/>([[Cenomanian]])
|
[[Burmese amber]]
|
{{Flag|Myanmar}}
|
A [[Cerophytidae|cerophytid rare click beetle]].
|
|-
|
''[[Anoeuma]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Kundrata |first2=R. |last3=Packova |first3=G. |last4=Huang |first4=D.-Y. |last5=Cai |first5=C.-Y. |title=An unusual elateroid lineage from mid-Cretaceous Burmese amber (Coleoptera: Elateroidea) |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 21985 |doi=10.1038/s41598-021-01398-w |pmid=34753998 |pmc=8578672 |bibcode=2021NatSR..1121985L }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li, Kundrata & Cai ''in'' Li ''et al.''
|
Cretaceous ([[Albian]] to Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Elateroidea|elateroid click beetle]] of uncertain phylogenetic placement.<br/> The type species is ''A. lawrencei''.
|
|-
|
''[[Arturmiles]]''<ref>{{Cite journal|last=Fanti |first=F. |title=A new genus of Cantharidae from Eocene Baltic amber found in Poland, with phoretic mites |journal=Baltic Journal of Coleopterology |volume=21 |issue=2 |pages=105–110 |url=http://www.bjc.sggw.pl/viewart.php?bookid=43&artid=748 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Fanti
|
Eocene
|
Prussian Formation<br>(Baltic amber)
|
{{Flag|Poland}}
|
A [[soldier beetle]] belonging to the tribe [[Cantharini]]. The type species is ''A. pankowskiarum''.
|
|-
|
''[[Balistica vetai]]''<ref name=Euryptychoides>{{Cite journal|last=Muona |first=J. |year=2021 |title=False-click beetles from Baltic amber (Coleoptera, Eucnemidae) |journal=Entomologische Blätter und Coleoptera |volume=117 |pages=1–20 |url=https://www.researchgate.net/publication/357381414 }}</ref>
|
Sp. nov
|
Valid
|
Muona
|
Eocene
|
Baltic amber
|
{{flag|Europe}}
|
A [[Dirhagini]] [[Melasinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Baltodascillus]]''<ref>{{Cite journal|last1=Kundrata |first1=R. |last2=Gimmel |first2=M. L. |last3=Packova |first3=G. |last4=Bukejs |first4=A. |last5=Blank |first5=S. M. |title=A new enigmatic lineage of Dascillidae (Coleoptera: Elateriformia) from Eocene Baltic amber described using X-ray microtomography, with notes on Karumiinae morphology and classification |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=141–149 |doi=10.5194/fr-24-141-2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kundrata ''et al.''
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Dascillidae|dascillid beetle]].<br/> The type species is ''B. serraticornis''.
|
|-
|
''[[Burmagrilus]]''<ref>{{Cite journal|last1=Jiang |first1=R. |last2=Song |first2=H. |last3=Zhang |first3=H. |last4=Wang |first4=S. |title=''Burmagrilus cretacus'' gen. et sp. nov., the first Buprestidae from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104866 |doi=10.1016/j.cretres.2021.104866 |bibcode=2021CrRes.12504866J }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jiang, Song & Wang ''in'' Jiang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Buprestidae|metallic wood-boring beetle]].<br/> The type species is ''B. cretacus''.
|
|-
|
''[[Burmomiles|Burmomiles bilineatimaculatus]]''<ref name=CRBurmomiles>{{Cite journal|last1=Yang |first1=Y. |last2=Zhao |first2=W. |last3=Geiser |first3=M. |last4=Zhang |first4=W. |last5=Bai |first5=M. |last6=Liu |first6=H. |title=Review of the genus ''Burmomiles'' Fanti, Damgaard & Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae) |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104961 |doi=10.1016/j.cretres.2021.104961 |bibcode=2021CrRes.12804961Y }}</ref>
|
Sp. nov
|
In press
|
Yang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[soldier beetle]]. Fanti & Müller (2022) considered it to be a species belonging to the genus ''[[Elektrokleinia]]''.<ref name=BJC222>{{Cite journal |last1=Fanti |first1=F. |last2=Müller |first2=P. |title=Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information |year=2022 |journal=Baltic Journal of Coleopterology |volume=22 |issue=2 |pages=331–380 |url=http://www.bjc.sggw.pl/viewart.php?bookid=45&artid=786 }}</ref>
|
|-
|
''[[Burmomiles|Burmomiles laticollis]]''<ref name=CRBurmomiles />
|
Sp. nov
|
In press
|
Yang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A soldier beetle.
|
|-
|
''[[Burmomiles|Burmomiles oblongoculus]]''<ref name=CRBurmomiles />
|
Sp. nov
|
In press
|
Yang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus ''Elektrokleinia''.<ref name=BJC222 />
|
|-
|
''[[Captopus]]''<ref name=Captopus>{{Cite journal|last1=Li |first1=Y. |last2=Huang |first2=D. |last3=Cai |first3=C. |title=New genera and species of the family Throscidae (Coleoptera: Elateroidea) in mid-Cretaceous Burmese amber |year=2021 |journal=Insects |volume=12 |issue=1 |pages=Article 63 |doi=10.3390/insects12010063 |pmid=33445628 |pmc=7826609 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai ''in'' Li, Huang & Cai
|
[[Cretaceous]]<br/> ([[Cenomanian]])
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Throscidae|throscid beetle]].<br/> The type species is ''C. depressiceps''.
|
|-
|
''[[Chelonarium andabata]]''<ref name=FR241Chelonarium>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Mitchell |first2=J. |last3=McKellar |first3=R. C. |last4=Barbi |first4=M. |last5=Larsson |first5=H. C. E. |last6=Bukejs |first6=A. |title=The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea) |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=19–32 |doi=10.5194/fr-24-19-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Alekseev & Bukejs ''in'' Alekseev ''et al.''
|
Eocene
|
[[Baltic amber]]
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Chelonariidae|turtle beetle]].
|
|-
|
''[[Chelonarium dingansich]]''<ref name=FR241Chelonarium />
|
Sp. nov
|
Valid
|
Alekseev & Bukejs ''in'' Alekseev ''et al.''
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Chelonariidae|turtle beetle]].
|
|-
|
''[[Chelonarium dominicanum]]''<ref>{{Cite journal|last1=Poinar |first1=G. |last2=Háva |first2=J. |title=The first representative of the beetle family Chelonariidae in Dominican amber |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=1–4 |doi=10.18476/pale.v14.a1 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Poinar & Háva
|
[[Burdigalian]]
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A [[Chelonariidae|turtle beetle]].
|
|-
|
''[[Coomanius balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Eocene
|
Baltic amber
|
{{Flag|Europe}}
|
A [[Nematodini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Cretocantharis]]''<ref name=Cretocantharis>{{Cite journal|last1=Hsiao |first1=Y. |last2=Li |first2=Y. |last3=Ren |first3=D. |last4=Pang |first4=H. |title=Morphological phylogenetics provide new insights into the classification and evolution of fossil soldier beetles from Mid-Cretaceous Burmese amber (Coleoptera: Cantharidae) |year=2021 |journal=Zoological Journal of the Linnean Society |volume=193 |issue=4 |pages=1271–1293 |doi=10.1093/zoolinnean/zlaa184 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Hsiao ''et al.''
|
Late Cretaceous<br/> (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[soldier beetle]].<br/> The type species is ''C. veda''.
|
|-
|
''[[Cretopachyderes]]''<ref>{{Cite journal|last1=Kundrata |first1=R. |last2=Prosvirov |first2=A. S. |last3=Long |first3=R. |last4=Packova |first4=G. |title=''Cretopachyderes'' gen. nov., a new remarkable click beetle (Coleoptera: Elateridae: Agrypninae) from the mid-Cretaceous Burmese amber |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=347–355 |doi=10.5194/fr-24-347-2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kundrata ''et al.''
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Agrypninae]] [[click beetle]].<br/>  The type species is ''C. burmitinus''.
|
|-
|
''[[Cretophengodes]]''<ref>{{cite journal |last1=Li |first1=Y. |last2=Kundrata |first2=R. |last3=Tihelka |first3=E. |last4=Liu |first4=Z. |last5=Huang |first5=D. |last6=Cai |first6=C. |year=2021 |title=Cretophengodidae, a new Cretaceous beetle family, sheds light on the evolution of bioluminescence |journal=Proceedings of the Royal Society B: Biological Sciences |volume=288 |issue=1943 |pages=Article ID 20202730 |doi=10.1098/rspb.2020.2730 |pmid=33468008 |pmc=7893276 |doi-access=free }}</ref>
|
Fam. nov., Gen. et sp. nov
|
Valid
|
Li, Kundrata, Tihelka & Cai ''in'' Li ''et al.''
|
Cretaceous (Albian or Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Cretophengodidae|cretophengodid]] [[Elateroidea|elateroid beetle]].<br/> The type species is ''C. azari''.
|
|-
|
''[[Dyscharachthis frickae]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[Dyscharachthini]] [[Eucneminae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Electrothroscus]]''<ref name=Captopus />
|
Gen. et sp. nov
|
Valid
|
Li & Cai 
|
Cretaceous<br/> Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Throscidae|throscid false click beetle]].<br/> The type species is ''E. yanpingae''.
|
|-
|
''[[Entomophthalmus antennalis]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[Dirhagini]] [[Melasinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Erdaia bispinulosa]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Orodotini]]? [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Erdaia longicornis]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Orodotini]]? [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Erdaia longitarsis]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Orodotini]]? [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Euryptychoides]]''<ref name=Euryptychoides />
|
Gen. et sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].<br/> The type species is ''E. magnus''.
|
|-
|
''[[Euryptychus decoratus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Euryptychus excavatus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Euryptychus helmwigae]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Euryptychus sieglindae]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Euryptychus waltrautae]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An [[Euryptychini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Hukawngichthyurus|Hukawngichthyurus maha]]''<ref name=Cretocantharis />
|
Sp. nov
|
Valid
|
Hsiao ''et al.''
|
Late Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[soldier beetle]].
|
|-
|
''[[Isorhipis balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
An ''[[Isorhipis]]'' species [[Melasinae|melasine]] [[Eucnemidae|false click beetle]]
|
|-
|
''[[Lycocerus crassepunctatus]]''<ref name=REJ302>{{Cite journal|last=Kazantsev |first=S. V. |title=New species of Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) |year=2021 |journal=Russian Entomological Journal |volume=30 |issue=2 |pages=153–158 |doi=10.15298/rusentj.30.2.08|issn=0132-8069  |s2cid=242185955 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Kazantsev
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[soldier beetle]].
|
|-
|
''[[Malthodes aliger]]''<ref name=REJ302 />
|
Sp. nov
|
Valid
|
Kazantsev
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[soldier beetle]]
|
|-
|
''[[Malthodes headsi]]''<ref name=BJC211>{{Cite journal|last=Fanti |first=F. |title=Two new ''Malthodes'' and a new ''Podistra'' from Eocene Baltic amber (Coleoptera, Cantharidae) |journal=Baltic Journal of Coleopterology |volume=21 |issue=1 |pages=1–9 |url=http://www.bjc.sggw.pl/viewart.php?bookid=42&artid=732 }}</ref>
|
Sp. nov
|
Valid
|
Fanti
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Poland}}
|
A [[soldier beetle]].
|
|-
|
''[[Malthodes jaredi]]''<ref name=BJC211 />
|
Sp. nov
|
Valid
|
Fanti
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Poland}}
|
A soldier beetle, a species of ''Malthodes''.
|
|-
|
''[[Malthodes vladiceniensis]]''<ref>{{Cite journal|last1=Pintilioaie |first1=A.-M. |last2=Fanti |first2=F. |last3=Ionesi |first3=V. |title=First finding of a fossilized Cantharidae (Insecta: Coleoptera) from the Middle Miocene deposits in Romania |year=2021 |journal=Travaux du Muséum National d'Histoire Naturelle "Grigore Antipa" |volume=64 |issue=1 |pages=73–80 |doi=10.3897/travaux.64.e64706 |s2cid=237805010 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Pintilioaie, Fanti & Ionesi
|
Middle Miocene
|
[[Bârnova-Muntele Formation]]
|
{{Flag|Romania}}
|
A [[soldier beetle]].
|
|-
|
''[[Microrhagus vetai|Microrhagus (Emyirhagus) vetai]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A ''[[Microrhagus]]''species [[Melasinae|melasine]] [[Eucnemidae|false click beetle]]
|
|-
|
''[[Murcybolus]]''<ref>{{Cite journal|last1=Li |first1=Y. |last2=Tihelka |first2=E. |last3=Huang |first3=D. |last4=Cai |first4=C. |title=''Murcybolus'' gen. nov., a new net-winged beetle genus from mid-Cretaceous Burmese amber (Coleoptera: Lycidae: Burmolycini) |year=2021 |journal=Zootaxa |volume=4966 |issue=1 |pages=77–83 |doi=10.11646/zootaxa.4966.1.8 |pmid=34186632 |s2cid=235577673 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li & Cai<br/> ''in'' Li ''et al.''
|
Cretaceous<br/> [[Cenomanian]]
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Burmolycini]] [[Dexorinae|dexorine]] [[Lycidae|net-winged beetle]].<br/> The type species is ''M. longiantennus''.
|
|-
|
''[[Nematodes schwertleitae]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Nematodini]] [[Macraulacinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Neocharoides]]''<ref name=Euryptychoides />
|
Gen. et sp. nov
|
Valid
|
Muona
|
Middle Eocene<br/> Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Nematodini]]? [[Macraulacinae]] [[Eucnemidae|false click beetle]].<br/> The type species is ''N. balticus''.
|
|-
|
''[[Palaeocantharis]]''<ref name=Cretocantharis />
|
Gen. et sp. nov
|
Disputed
|
Hsiao ''et al.''
|
Late Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[soldier beetle]]. The type species is ''P. panna''.<br/> Fanti & Müller (2022) considered the genus ''Palaeocantharis'' to a be a [[Synonym (taxonomy)|junior synonym]] of the genus ''[[Elektrokleinia]]'', though the authors maintained ''P. panna'' as a distinct species within the latter genus.<ref name=BJC222 />
|
|-
|
''[[Podistra guthriei]]''<ref name=BJC211 />
|
Sp. nov
|
Valid
|
Fanti
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Poland}}
|
A [[soldier beetle]].
|
|-
|
''[[Poecilochrus balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Eucnemini]] [[Eucneminae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Potergites brittoni]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Dendrocharini]] [[Eucneminae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Pseudomataeopsephus]]''<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Huang |first2=D. |last3=Cai |first3=C. |title=''Pseudomataeopsephus'', a new genus of water penny beetles from mid-Cretaceous Burmese amber (Coleoptera: Psephenidae) |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105055 |doi=10.1016/j.cretres.2021.105055 |s2cid=244242362 }}</ref>
|
Gen. et sp. nov
|
In press
|
Li, Huang & Cai
|
Cretaceous<br/>(Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[water-penny beetle]].<br/>The type species is ''P. burmensis''.
|
|-
|
''[[Pseudopactopus]]''<ref name=Captopus />
|
Gen. et sp. nov
|
Valid
|
Li & Cai <br/>''in'' Li, Huang & Cai
|
Cretaceous<br/> ( Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Throscidae|throscid false click beetle]].<br/> The type species is ''P. robustus''.
|
|-
|
''[[Ptilodactyla eocenica]]''<ref>{{Cite journal|last1=Kundrata |first1=R. |last2=Packova |first2=G. |last3=Kairišs |first3=K. |last4=Bukejs |first4=A. |last5=Hoffmannova |first5=J. |last6=Blank |first6=S. M. |title=The First ''Ptilodactyla'' Illiger, 1807 (Coleoptera: Dryopoidea: Ptilodactylidae) Described from Eocene Baltic Amber |year=2021 |journal=Biology |volume=10 |issue=9 |pages=Article 877 |doi=10.3390/biology10090877 |pmid=34571754 |pmc=8465962 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Kundrata, Bukejs & Blank<br/> ''in'' Kundrata ''et al.''
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A''[[Ptilodactyla]]'' species [[Ptilodactylidae|toe winged beetle]].
|
|-
|
''[[Rhacopus balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A ''[[Rhacopus]]'' species [[Eucnemidae|eucnemid false click beetle]]
|
|-
|
''[[Rhagomicrus balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A [[Dirhagini]] [[Melasinae]] [[Eucnemidae|false click beetle]].
|
|-
|
''[[Rheanischia]]''<ref>{{Cite journal|last1=Li |first1=H. |last2=Chang |first2=H. |last3=Muona |first3=J. |last4=Zhao |first4=Y. |last5=Ren |first5=D. |title=Subfamily Anischiinae (Coleoptera: Eucnemidae) in Early Cretaceous of Northeast China |year=2021 |journal=Insects |volume=12 |issue=2 |pages=Article 105 |doi=10.3390/insects12020105 |pmid=33530615 |pmc=7910996 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
|
Li, Chang & Muona
|
[[Early Cretaceous]]<br/>([[Hauterivian]] to [[Aptian]])
|
[[Yixian Formation]]
|
{{Flag|China}}
|
An [[Anischiinae]] [[Eucnemidae|false click beetle]].<br/> The type species is ''R. brevicornis''.
|
|-
|
''[[Silis curleri]]''<ref name=Zootaxa49961>{{Cite journal|last1=Fanti |first1=F. |last2=Pankowski |first2=M. G. |year=2021 |title=Two new species of soldier beetles (Coleoptera, Cantharidae), the first from the tribe Silini in Dominican amber |journal=Zootaxa |volume=4996 |issue=1 |pages=163–170 |doi=10.11646/zootaxa.4996.1.7 |pmid=34810540 |s2cid=237823176 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Fanti & Pankowski
|
[[Burdigalian]]
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A species of ''[[Silis (beetle)|Silis]]''.
|
|-
|
''[[Silis hegnai]]''<ref name=Zootaxa49961 />
|
Sp. nov
|
Valid
|
Fanti & Pankowski
|
[[Burdigalian]]
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A species of ''Silis''.
|
|-
|
''[[Xylophilus balticus]]''<ref name=Euryptychoides />
|
Sp. nov
|
Valid
|
Muona
|
Late Eocene<br/>Priabonian
|
Baltic amber
|
{{flag|Europe}}
|
A ''[[Xylophilus (beetle)|Xylophilus]]'' species [[Eucnemidae|eucnemid false click beetle]].
|
|-
|}

======Elateriformia research======
* A study on the anatomy and phylogenetic relationships of ''[[Archaeolus]] funestus'' is published by Li ''et al.'' (2021).<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Kundrata |first2=R. |last3=Huang |first3=D.-Y. |last4=Cai |first4=C.-Y. |title=Notes on the Morphology and Systematic Position of ''Archaeolus'' Lin, 1986, from the Jurassic of South China (Coleoptera: Elateroidea) |year=2021 |journal=Insects |volume=12 |issue=10 |pages=Article 876 |doi=10.3390/insects12100876 |pmid=34680645 |pmc=8537516 |doi-access=free }}</ref>
* A study on the morphological diversity of larvae of extant and fossil members of the family [[Brachypsectridae]] is published by Haug ''et al.'' (2021).<ref>{{Cite journal|last1=Haug |first1=J. T. |last2=Zippel |first2=A. |last3=Haug |first3=G. T. |last4=Hoffeins |first4=C. |last5=Hoffeins |first5=H.-W. |last6=Hammel |first6=J. U. |last7=Baranov |first7=V. |last8=Haug |first8=C. |title=Texas beetle larvae (Brachypsectridae) – the last 100 million years reviewed |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=161–183 |doi=10.18476/pale.v14.a8 |doi-access=free }}</ref>

=====Scarabaeiformia=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Goriresina]]''<ref>{{Cite journal|last1=Matalin |first1=A. V. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |year=2021 |title=First record of tiger beetles (Coleoptera, Cicindelidae) from Rovno amber, with the description of a new genus and species |journal=Zootaxa |volume=5016 |issue=2 |pages=243–256 |doi=10.11646/zootaxa.5016.2.5 |pmid=34810452 |s2cid=238845128 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Matalin, Perkovsky & Vasilenko
|
Late Eocene<br/>Priabonian
|
Rovno amber
|
{{Flag|Ukraine}}
|
A [[tiger beetle]].<br/> The type species is ''G. fungifora''.
|
|-
|
''[[Onthophagus pilauco]]''<ref>{{Cite journal|last1=Tello |first1=F. |last2=Verdú |first2=J. R. |last3=Rossini |first3=M. |last4=Zunino |first4=M. |title=''Onthophagus pilauco'' sp. nov. (Coleoptera, Scarabaeidae): evidence of beetle extinction in the Pleistocene–Holocene transition in Chilean Northern Patagonia |year=2021 |journal=ZooKeys |issue=1043 |pages=133–145 |doi=10.3897/zookeys.1043.61706 |doi-access=free |pmid=34168516 |pmc=8219658 }}</ref>
|
Sp. nov
|
Valid
|
Tello ''et al.''
|
Late [[Pleistocene]]
|
[[Pilauco Bajo]]
|
{{Flag|Chile}}
|
A [[Scarabaeidae|scarab beetle]].
|
|-
|}

=====Staphyliniformia=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name 
!Novelty
!Status
!Authors
!Age
!Type locality
!Country
!Notes
!Images
|-
|
''[[Acritus sutirca]]''<ref>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |title=The first extinct species of ''Acritus'' LeConte, 1853 (Histeridae: Abraeinae) from Eocene Baltic amber: a microscopic beetle inclusion studied with X-ray micro-computed tomography |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=223–231 |doi=10.5194/fr-24-223-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br />({{Flag|Kaliningrad Oblast}})
|
A species of ''[[Acritus]]''.
|
|-
|
''[[Bacanius gorskii]]''<ref>{{Cite journal|last1=Alekseev |first1=V. I. |last2=Bukejs |first2=A. |title=A new Eocene ''Bacanius'' species (Histeridae: Dendrophilinae) from Baltic amber |year=2021 |journal=Fossil Record |volume=24 |issue=1 |pages=93–99 |doi=10.5194/fr-24-93-2021 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Alekseev & Bukejs
|
Eocene
|
[[Baltic amber]]
|
{{Flag|Poland}}
|
A species of ''[[Bacanius]]''.
|
|-
|
''[[Burmagluta]]''<ref>{{Cite journal|last1=Yin |first1=Z.-W. |last2=Cai |first2=C.-Y. |title=A new Brachyglutini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Staphylinidae: Pselaphinae) |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104807 |doi=10.1016/j.cretres.2021.104807 |bibcode=2021CrRes.12404807Y |s2cid=233577289 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Yin & Cai
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Brachyglutini]] [[Pselaphinae]] [[rove beetle]].<br/> The type species is ''B. rougemonti''.
|
|-
|
''[[Carinumerus]]''<ref name=Carinumerus>{{Cite journal|last=Caterino |first=M. S. |title=New Fossil Histerid Species from Cretaceous Burmese Amber (Coleoptera: Histeridae) |year=2021 |journal=The Coleopterists Bulletin |volume=75 |issue=1 |pages=211–221 |doi=10.1649/0010-065X-75.1.211 |s2cid=232368396 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Caterino
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Onthophilinae]] [[Histeridae|clown beetle]].<br/> The type species is ''C. maddisoni''.
|
|-
|
''[[Charhyphus serratus]]''<ref>{{Cite journal|last1=Yamamoto |first1=S. |last2=Shavrin |first2=A. V. |last3=Kairišs |first3=K. |year=2021 |title=A second fossil species of the enigmatic rove beetle genus ''Charhyphus'' in Eocene Baltic amber, with implications on the morphology of the female genitalia (Coleoptera: Staphylinidae: Phloeocharinae) |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |volume=113 |pages=39–50 |doi=10.1017/S1755691021000360 |hdl=2115/86861 |s2cid=244931145 |hdl-access=free }}</ref>
|
Sp. nov
|
In press
|
Yamamoto & Shavrin
|
Eocene
|
Baltic amber
|
{{flag|Europe}}
|
A [[Phloeocharinae]] [[Staphylinidae|rove beetle]].
|
|-
|
''[[Cretosaja]]''<ref>{{Cite journal|last1=Sohn |first1=J.-C. |last2=Nam |first2=G. S. |title=New fossil genus and species of carrion beetle (Coleoptera, Silphidae) from the Lower Cretaceous Jinju Formation, South Korea |year=2021 |journal=Journal of Asia-Pacific Entomology |volume=24 |issue=3 |pages=584–587 |doi=10.1016/j.aspen.2021.05.003 |s2cid=236585083 }}</ref>
|
Gen. et sp. nov
|
In press
|
Sohn & Nam
|
[[Early Cretaceous]]
|
[[Jinju Formation]]
|
{{Flag|South Korea}}
|
A [[Silphidae|carrion beetle]].<br/> The type species is ''C. jinjuensis''.
|
|-
|
''[[Druantia aeterna|Druantia]]''<ref name=Carinumerus />
|
Gen. et sp. nov
|
Valid
|
Caterino
|
Cenomianian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Dendrophilinae]] [[Histeridae|clown beetle]].<br/> The type species is ''D. aeterna''.
|
|-
|
''[[Euconnus nathani]]''<ref>{{Cite journal|last1=Jałoszyński |first1=P. |last2=Perkovsky |first2=E. E. |year=2021 |title=A new bizarre species of ''Euconnus'' (''Cladoconnus'') in Upper Eocene Rovno amber (Coleoptera: Staphylinidae: Scydmaeninae) |journal=Zootaxa |volume=5004 |issue=2 |pages=395–400 |doi=10.11646/zootaxa.5004.2.8 |pmid=34811301 |s2cid=237709269 }}</ref>
|
Sp. nov
|
Valid
|
Jałoszyński & Perkovsky
|
Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A ''[[Euconnus]]'' species [[Scydmaeninae]] [[Staphylinidae|rove beetle]].
|
|-
|
''[[Eusphalerum bukejsi]]''<ref>{{Cite journal|last1=Shavrin |first1=A. V. |last2=Kairišs |first2=K. |year=2021 |title=A new species of ''Eusphalerum'' Kraatz, 1857 from the Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae) |journal=Zootaxa |volume=4966 |issue=4 |pages=469–475 |doi=10.11646/zootaxa.4966.4.5 |pmid=34186602 |s2cid=235565793 }}</ref>
|
Sp. nov
|
Valid
|
Shavrin ''in'' Shavrin & Kairišs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A ''[[Eusphalerum]]'' species [[Omaliinae]] [[Staphylinidae|rove beetle]].
|
|-
|
''[[Megabythinus]]''<ref>{{Cite journal|last1=Yin |first1=Z.-W. |last2=Zhao |first2=Q.-H. |last3=Cai |first3=C.-Y. |title=''Megabythinus'' gen. nov., a new stem-group Pselaphinae from mid-Cretaceous Myanmar amber (Coleoptera, Staphylinidae) |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104879 |doi=10.1016/j.cretres.2021.104879 |bibcode=2021CrRes.12504879Y }}</ref>
|
Gen. et sp. nov
|
Valid
|
Yin, Zhao & Cai
|
Cretaceous<br/>Cenomanian
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Bythinini]] [[Pselaphinae]] [[Staphylinidae|rove beetle]].<br/> The type species is ''M. spinitibialis''.
|
|-
|
''[[Paroxyporus]]''<ref>{{Cite journal|last1=Liu |first1=Y. |last2=Tihelka |first2=E. |last3=Thayer |first3=M. K. |last4=Newton |first4=A. F. |last5=Huang |first5=D. |last6=Tian |first6=L. |last7=Cai |first7=C. |title=A transitional fossil sheds light on the early evolution of the Staphylinine group of rove beetles (Coleoptera: Staphylinidae) |year=2021 |journal=Journal of Systematic Palaeontology |volume=19 |issue=4 |pages=321–332 |doi=10.1080/14772019.2021.1917705 |s2cid=235172619 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cai 
|
[[Early Cretaceous]]
|
[[Yixian Formation]]
|
{{Flag|China}}
|
A [[rove beetle]].<br/> The type species is ''P. fossilis''.
|
|-
|
''[[Phasmister]]''<ref name=Carinumerus />
|
Gen. et sp. nov
|
Valid
|
Caterino
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Onthophilinae]] [[Histeridae|clown beetle]].<br/> The type species is ''P. cristatus''.
|
|-
|
''[[Ponohydrochus]]''<ref>{{Cite journal|last1=Prokin |first1=A. A. |last2=Strelnikova |first2=O. D. |title=The first remarkable fossil Hydrochidae (Coleoptera) from the Lower Cretaceous (Hauterivian) of the Buryatia Republic, Russia |year=2021 |journal=Cretaceous Research |volume=123 |pages=Article 104795 |doi=10.1016/j.cretres.2021.104795 |bibcode=2021CrRes.12304795P |s2cid=233957495 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Prokin & Strelnikova
|
[[Early Cretaceous]]<br/>([[Hauterivian]])
|
|
{{Flag|Russia}}
|
A [[Hydrochidae|water scavenger beetle]].<br/> The type species is ''P. buryaticus''.
|
|-
|
''[[Proptomaphaginus alleni]]''<ref>{{Cite journal|last1=Perreau |first1=M. |last2=Haelewaters |first2=D. |last3=Tafforeau |first3=P. |title=A parasitic coevolution since the Miocene revealed by phase-contrast synchrotron X-ray microtomography and the study of natural history collections |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 2672 |doi=10.1038/s41598-020-79481-x |pmid=33514784 |pmc=7846571 |bibcode=2021NatSR..11.2672P }}</ref>
|
Sp. nov
|
Valid
|
Perreau & Haelewaters<br/>''in'' Perreau, Haelewaters & Tafforeau
|
Miocene<br/> Burdigalian
|
Dominican amber
|
{{Flag|Dominican Republic}}
|
A [[Ptomaphagini]] [[Leiodidae|round fungus beetle]].
|
|-
|
''[[Trichophya minor]]''<ref>{{Cite journal|last1=Yamamoto |first1=S. |last2=Newton |first2=A. F. |title=First fossil trichophyine rove beetle from mid-Cretaceous amber of northern Myanmar (Coleoptera: Staphylinidae: Trichophyinae) |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104951 |doi=10.1016/j.cretres.2021.104951 |bibcode=2021CrRes.12704951Y |url=https://zenodo.org/record/4784783 |doi-access=free |hdl=2115/90159 |hdl-access=free }}</ref>
|
Sp. nov
|
Valid
|
Yamamoto & Newton
|
Cretaceous<br/> (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A ''[[Trichophya]]'' species [[Trichophyinae]] [[Staphylinidae|rove beetle]].
|
|-
|
''[[Yethiha]]''<ref name=Carinumerus />
|
Gen. et sp. nov
|
Valid
|
Caterino
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Dendrophilinae]] [[Histeridae|clown beetle]].<br/> The type species is ''Y. peregrina''.
|
|-
|}

====other Coleopteran research====
* A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the [[Permian–Triassic extinction event]] on beetle diversity, is published by Zhao ''et al.'' (2021).<ref>{{Cite journal|last1=Zhao |first1=X. |last2=Yu |first2=Y. |last3=Clapham |first3=M. E. |last4=Yan |first4=E. |last5=Chen |first5=J. |last6=Jarzembowski |first6=E. A. |last7=Zhao |first7=X. |last8=Wang |first8=B. |year=2021 |title=Early evolution of beetles regulated by the end-Permian deforestation |journal=eLife |volume=10 |pages=e72692 |doi=10.7554/eLife.72692 |pmid=34747694 |pmc=8585485 |doi-access=free }}</ref>
* Revision of ''Notocupes undatabdominus'' and ''N. ? multituberatus'' from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus ''[[Notocupes]]'', transfer ''N. ? multituberatus'' to the genus ''[[Lasiosyne]]'', and consider ''"N." undatabdominus'' to be likely a member of the family [[Artematopodidae]].<ref>{{Cite journal|last1=Li |first1=Y.-D. |last2=Huang |first2=D.-Y. |last3=Cai |first3=C.-Y. |year=2021 |title=Revisiting the systematic positions of two ''"Notocupes"'' species from the Lower Cretaceous of South China (Coleoptera: Polyphaga) |journal=Zootaxa |volume=5005 |issue=1 |pages=86–90 |doi=10.11646/zootaxa.5005.1.7 |pmid=34811277 |s2cid=237669437 |doi-access=free }}</ref>
* The oldest solid wood-borer larvae reported to date (belonging to the families [[Buprestidae]] and possibly [[Cerambycidae]]) are described from the Cretaceous amber from Myanmar by Haug ''et al.'' (2021).<ref>{{Cite journal|last1=Haug |first1=C. |last2=Haug |first2=G. T. |last3=Zippel |first3=A. |last4=van der Wal |first4=S. |last5=Haug |first5=J. T. |title=The earliest record of fossil solid-wood-borer larvae—immature beetles in 99 million-year-old Myanmar amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=390–404 |doi=10.11646/palaeoentomology.4.4.14 |issn=2624-2826  |s2cid=237622518 |doi-access=free }}</ref>
* Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from [[pack rat]] middens from [[California]] ([[United States]]) and [[Baja California]] ([[Mexico]]) dating up to ~ 34,355 years before present by Smith ''et al.'' (2021).<ref>{{Cite journal|last1=Smith |first1=A. D. |last2=Kamiński |first2=M. J. |last3=Kanda |first3=K. |last4=Sweet |first4=A. D. |last5=Betancourt |first5=J. L. |last6=Holmgren |first6=C. A. |last7=Hempel |first7=E. |last8=Alberti |first8=F. |last9=Hofreiter |first9=M. |title=Recovery and analysis of ancient beetle DNA from subfossil packrat middens using high-throughput sequencing |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 12635 |doi=10.1038/s41598-021-91896-8 |pmid=34135378 |pmc=8209150 }}</ref>

==Clade Dictyoptera==
===New Dictyopteran taxa===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Afrophaga]]''<ref name=Afrophaga>{{Cite journal|last1=Vršanský |first1=P. |last2=Vršanská |first2=L. |last3=Vasilenko |first3=D. V. |last4=Puškelová |first4=L. |last5=Biroň |first5=A. |title=An isolated Cretaceous analogue of Madagascar on the Adria–Turkey microcontinent indicated by fossils in Brezina, Algeria |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=19–35 |doi=10.1127/pala/2021/0107 |s2cid=240038689 }}</ref>
|
Gen. et sp. nov
|
In press
|
Vršanský ''et al.''
|
[[Cretaceous]] ([[Hauterivian]]?)
|
|
{{Flag|Algeria}}
|
A cockroach belonging to the family [[Corydiidae]]. Genus includes new species ''A. extincta''.
|
|-
|
''[[Alienopterix|Alienopterix mlynskyi]]''<ref name=Nadveruzenie>{{Cite journal|last1=Vršanský |first1=P. |last2=Sendi |first2=H. |last3=Hinkelman |first3=J. |last4=Hain |first4=M. |title=''Alienopterix'' Mlynský et al., 2018 complex in North Myanmar amber supports Umenocoleoidea/ae status |year=2021 |journal=Biologia |volume=76 |issue=8 |pages=2207–2224 |doi=10.1007/s11756-021-00689-x |s2cid=233784192 }}</ref>
|
Sp. nov
|
In press
|
Sendi ''in'' Vršanský ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An [[Alienopteridae|alienopterid]].
|
|-
|
''Alienopterix smidovae''<ref name=Nadveruzenie />
|
Sp. nov
|
In press
|
Hinkelman ''in'' Vršanský ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An alienopterid.
|
|-
|
''[[Arceotermes]]''<ref name=Arceotermes>{{Cite journal|last1=Jiang |first1=R.X. |last2=Zhang |first2=H.R. |last3=Eldredge |first3=K.T. |last4=Song  |first4=X.B. |last5=Li |first5=Y.D. |last6=Tihelka |first6=E. |last7=Huang |first7=D.Y. |last8=Wang |first8=S. |last9=Engel |first9=M.S. |last10=Cai |first10=C.Y. |title=Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine ''Cretotrichopsenius'' (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=374–389 |doi=10.11646/palaeoentomology.4.4.13 |s2cid=239695173 }}</ref>
|
Fam. et gen. et sp. nov
|
Valid
|
[[Michael S. Engel|Engel]] & Jiang ''in'' Jiang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A termite belonging to the new family [[Arceotermitidae]]. Genus includes new species ''A. hospitis''.
|
|-
|
''[[Caligoptera]]''<ref name=Caligoptera>{{Cite journal|last=Sendi |first=H. |title=Highly specialised basal ectobiid cockroaches (Blattaria: Blattoidea) were rare in Burmese amber |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=109–125 |doi=10.1127/pala/2021/0106 |s2cid=243471520 }}</ref>
|
Gen. et sp. nov
|
In press
|
Sendi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family [[Ectobiidae]]. Genus includes new species ''C. hinkelmani''.
|
|-
|
''[[Chuanblatta]]''<ref>{{Cite journal|last1=Liang |first1=J. |last2=Wang |first2=Y. |last3=Shih |first3=C. |last4=Ren |first4=D. |title=''Chuanblatta'' gen. nov. sexually dimorphic cockroaches of Raphidiomimidae (Blattaria) from the Jiulongshan Formation in China |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=3–17 |doi=10.1127/pala/2021/0113|issn=0375-0442  |s2cid=240180893 }}</ref>
|
Gen. et 2 sp. nov
|
In press
|
Liang ''et al.''
|
[[Middle Jurassic]]
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A cockroach belonging to the family [[Raphidiomimidae]]. Genus includes new species ''C. huntianlingensis'' and ''C. lata''.
|
|-
|
''[[Cuniculoblatta]]''<ref>{{Cite journal|last=Hinkelman |first=J. |title=''Cuniculoblatta brevialata'' gen. et sp. n., the second case of brachyptery from Cretaceous North Myanmar amber |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=97–107 |doi=10.1127/pala/2021/0104 |s2cid=240183883 }}</ref>
|
Gen. et sp. nov
|
In press
|
Hinkelman
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach. Genus includes new species ''C. brevialata ''.
|
|-
|
''[[Ectoovia]]''<ref name=Caligoptera />
|
Gen. et sp. nov
|
In press
|
Sendi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family Ectobiidae. Genus includes new species ''E. protecta''.
|
|-
|
''[[Elisama|? Elisama algeriaensis]]''<ref name=Afrophaga />
|
Sp. nov
|
In press
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of the family [[Blattulidae]].
|
|-
|
''[[Eminespina]]''<ref>{{Cite journal|last1=Chen |first1=X.-Y. |last2=Zhang |first2=H.-C. |last3=Shi |first3=X. |title=A new species and genus of Alienopteridae (Blattodea) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Zootaxa |volume=4941 |issue=4 |pages=580–586 |doi=10.11646/zootaxa.4941.4.7 |pmid=33756925 |s2cid=232339877 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Chen, Zhang  Shi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
An alienopterid. Genus includes new species ''E. burma''.
|
|-
|
''[[Fractalia]]''<ref name=Fractalia>{{Cite journal|last1=Vršanský |first1=P. |last2=Hinkelman |first2=J. |last3=Koubová |first3=I. |last4=Sendi |first4=H. |last5=Kúdelová |first5=T. |last6=Kúdela |first6=M. |last7=Barclay |first7=M. |title=A single common ancestor for praying mantids, termites, cave roaches and umenocoleoids |year=2021 |journal=AMBA Projekty |volume=11 |issue=1 |pages=1–16 |url=http://geo.sav.sk/files/vrsansky/Vrsansky_et_al_2021.pdf }}</ref>
|
Fam. et gen. et 2 sp. nov
|
Valid
|
Hinkelman & Vršanský ''in'' Vršanský ''et al.''
|
[[Late Jurassic]]–Late Cretaceous ([[Kimmeridgian]]–Cenomanian)
|
Burmese amber
|
{{Flag|Germany}}<br>
{{Flag|Kazakhstan}}<br>
{{Flag|Myanmar}}
|
A member of [[Corydioidea]], the type genus of the new family [[Fractaliidae]] Vršanský & Hinkelman. The type species is ''F. articulata'' Hinkelman; genus also includes new species ''F. aristovi'' Vršanský.
|
|-
|
''[[Fragosublatta]]''<ref>{{Cite journal|last1=Chen |first1=G. |last2=Xiao |first2=L. |last3=Liang |first3=J. |last4=Shih |first4=C. |last5=Ren |first5=D. |title=A new cockroach (Blattodea, Corydiidae) with pectinate antennae from mid-Cretaceous Burmese amber |year=2021 |journal=ZooKeys |issue=1060 |pages=155–169 |doi=10.3897/zookeys.1060.67216 |doi-access=free |pmid=34690506 |pmc=8486729 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Chen, Shih & Ren ''in'' Chen ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family [[Corydiidae]]. The type species is ''F. pectinata''.
|
|-
|
''[[Glyptotermes abyssinicus]]''<ref>{{Cite journal|last1=Bouju |first1=V. |last2=Jouault |first2=C. |last3=Perrichot |first3=V. |title=The termite genus ''Glyptotermes'' (Isoptera: Kalotermitidae) in Miocene amber from Ethiopia |year=2021 |journal=Journal of Paleontology |volume= 96|issue=2 |pages=387–393 |doi=10.1017/jpa.2021.106 |s2cid=244426915 |url=https://hal-insu.archives-ouvertes.fr/insu-03435971/file/JPA-RA-2021-0060.R2_Proof_hi.pdf }}</ref>
|
Sp. nov
|
Valid
|
Bouju, Jouault & Perrichot
|
Early Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A termite belonging to the family [[Kalotermitidae]].
|
|-
|
''[[Kinneyblatta]]''<ref>{{Cite journal |last1=Schneider |first1=J. W. |last2=Scholze |first2=F. |last3=Germann |first3=S. |last4=Lucas |first4=S. G. |title=The Late Pennsylvanian nearshore insect fauna of the Kinney Brick Quarry invertebrate and vertebrate fossil Lagerstätte, New Mexico |year=2021 |journal=New Mexico Museum of Natural History and Science Bulletin |volume=84 |pages=255–285 |url=https://www.researchgate.net/publication/350896849 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Schneider ''et al.''
|
[[Carboniferous]] ([[Kasimovian]])
|
[[Atrasado Formation]]
|
{{Flag|United States}}<br>({{Flag|New Mexico}})
|
A member of [[Blattodea]] belonging to the family [[Spiloblattinidae]]. The type species is ''K. huberi''.
|
|-
|
''[[Kriedoblatta]]''<ref name=Afrophaga />
|
Gen. et sp. nov
|
In press
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of the family [[Liberiblattinidae]]. Genus includes new species ''K. gondwanensis''.
|
|-
|
''[[Kurablattina|? Kurablattina samsonovi]]''<ref name=Afrophaga />
|
Sp. nov
|
In press
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of the family Liberiblattinidae.
|
|-
|
''[[Labradormantis]]''<ref>{{Cite journal|last1=Demers-Potvin |first1=A. V. |last2=Larsson |first2=H. C. E. |last3=Cournoyer |first3=M. |last4=Béthoux |first4=O. |title=Wing morphology of a new Cretaceous praying mantis solves the phylogenetic jigsaw of early-diverging extant lineages |year=2021 |journal=Systematic Entomology |volume=46 |issue=1 |pages=205–223 |doi=10.1111/syen.12457 |s2cid=231391397 |url=https://escholarship.mcgill.ca/concern/articles/0r967824q }}</ref>
|
Gen. et sp. nov
|
Valid
|
Demers-Potvin ''et al.''
|
[[Late Cretaceous]] ([[Cenomanian]])
|
[[Redmond Formation]]
|
{{Flag|Canada}}
|
A praying mantis belonging to the family [[Baissomantidae]]. Genus includes new species ''L. guilbaulti''.
|
|-
|
''[[Latiblattella basaltica]]''<ref>{{Cite journal|last1=Vršanský |first1=P. |last2=Poschmann |first2=M. J. |last3=Vidlička |first3=L. |title=Oligocene pseudophyllodromiin cockroach from the Enspel Fossillagerstätte in Germany |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=149–167 |doi=10.1127/pala/2021/0110|issn=0375-0442  |s2cid=240039709 }}</ref>
|
Sp. nov
|
In press
|
Vršanský, Poschmann & Vidlička
|
Late [[Oligocene]]
|
|
{{Flag|Germany}}
|
A species of ''[[Latiblattella]]''.
|
|-
|
''[[Latiblattella karlgruberi]]''<ref name=Caligoptera />
|
Sp. nov
|
In press
|
Sendi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
|
|-
|
''[[Laticephalana]]''<ref>{{Cite journal|last1=Luo |first1=C. |last2=Beutel |first2=R. G. |last3=Xu |first3=C. |last4=Jarzembowski |first4=E. A. |title=†''Laticephalana liuyani'' gen. et sp. nov., a new bizarre roachoid of †Umenocoleidae (Insecta, Dictyoptera) from mid-Cretaceous Kachin amber |year=2021 |journal=Proceedings of the Geologists' Association |volume=132 |issue=4 |pages=469–478 |doi=10.1016/j.pgeola.2021.04.004 |bibcode=2021PrGA..132..469L |s2cid=236329819 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Luo ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Umenocoleidae]]. Genus includes new species ''L. liuyani''.
|
|-
|
''[[Lukotermes]]''<ref>{{Cite journal|last1=Perkovsky |first1=E. E. |last2=Nel |first2=A. |title=A new Rovno amber termite genus (Isoptera, Rhinotermitidae) from Styr river basin |year=2021 |journal=Palaeontologia Electronica |volume=24 |issue=1 |pages=Article number 24.1.a05 |doi=10.26879/1127 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Perkovsky & Nel
|
Eocene ([[Priabonian]])
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
A termite belonging to the family [[Rhinotermitidae]]. Genus includes new species ''L. milescaput''.
|
|-
|
''[[Meiatermes|Meiatermes cretacicus]]''<ref>{{Cite journal |last1=Bezerra |first1=F. I. |last2=DeSouza |first2=O. |last3=Ribeiro |first3=G. C. |last4=Mendes |first4=M. |year=2021 |title=A new primitive termite (Isoptera) from the Crato Formation, Araripe Basin, Early Cretaceous of South America |journal=Journal of South American Earth Sciences |volume=109 |pages=Article 103260 |doi=10.1016/j.jsames.2021.103260 |bibcode=2021JSAES.10903260B |s2cid=233807443 |url=http://www.repositorio.ufc.br/handle/riufc/64786 }}</ref>
|
Sp. nov
|
In press
|
Bezerra ''et al.''
|
[[Early Cretaceous]]
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A termite.
|
|-
|
''[[Meloblatta]]''<ref name=Afrophaga />
|
Gen. et sp. nov
|
In press
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of the family [[Mesoblattinidae]]. Genus includes new species ''M. brezinica''.
|
|-
|
''[[Milesitermes]]''<ref name=Milesitermes>{{Cite journal|last1=Jouault |first1=C. |last2=Legendre |first2=F. |last3=Grandcolas |first3=P. |last4=Nel |first4=A. |title=Revising dating estimates and the antiquity of eusociality in termites using the fossilized birth–death process |year=2021 |journal=Systematic Entomology |volume=46 |issue=3 |pages=592–610 |doi=10.1111/syen.12477 |s2cid=235363027 |url=https://hal-insu.archives-ouvertes.fr/insu-03188488/document }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A termite belonging to the family [[Mastotermitidae]]. Genus includes new species ''M. engeli''.
|
|-
|
''[[Miniblattina]]''<ref name=Miniblattina>{{Cite journal|last=Sendi |first=H. |title=Diverse Liberiblattinidae (Insecta: Blattaria) from Lebanese and North Myanmar amber document allometric modifications near lowest size limit |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=127–148 |doi=10.1127/pala/2021/0108 |s2cid=243475716 }}</ref>
|
Gen. et sp. nov
|
In press
|
Sendi
|
[[Early Cretaceous]] ([[Barremian]])
|
[[Lebanese amber]]
|
{{Flag|Lebanon}}
|
A cockroach belonging to the family [[Liberiblattinidae]]. Genus includes new species ''M. libera''.
|
|-
|
''[[Mongolblatta|Mongolblatta sendii]]''<ref>{{Cite journal|last=Hinkelman |first=J. |title=''Mongolblatta sendii'' sp. n. (Mesoblattinidae) from North Myanmar amber links record to Laurasian sediments |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=81–96 |doi=10.1127/pala/2021/0105 |s2cid=240162578 }}</ref>
|
Sp. nov
|
In press
|
Hinkelman
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Mesoblattinidae.
|
|-
|
''[[Morphna cenozoica]]''<ref name=PAABlaberidae>{{Cite journal|last1=Šmídová |first1=L. |last2=Vidlička |first2=L. |last3=Wedmann |first3=S. |title=Appearance of the family Blaberidae (Insecta: Blattaria) during the Cretaceous and a review of fossils of this family |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=71–79 |doi=10.1127/pala/2021/0109 |s2cid=243467634 }}</ref>
|
Sp. nov
|
In press
|
Šmídová, Vidlička & Wedmann
|
[[Eocene]]
|
[[Messel pit]]
|
{{Flag|Germany}}
|
A cockroach belonging to the family [[Blaberidae]].
|
|-
|
''[[Morphna cretacica]]''<ref name=PAABlaberidae />
|
Sp. nov
|
In press
|
Šmídová, Vidlička & Wedmann
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family Blaberidae.
|
|-
|
''[[Nadveruzenie]]''<ref name=Nadveruzenie />
|
Gen. et sp. nov
|
In press
|
Vršanský, Hinkelman & Sendi ''in'' Vršanský ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An alienopterid. Genus includes new species ''N. postava''.
|
|-
|
''[[Nodosigalea|Nodosigalea simplivena]]''<ref name=Olenablatta>{{Cite journal|last=Šmídová |first=L. |title=New genus and species of the families Olidae and Corydiidae (Corydioidea, Blattodea) from mid-Cretaceous Kachin amber |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=61–70 |doi=10.1127/pala/2021/0117 |s2cid=243471507 }}</ref>
|
Sp. nov
|
In press
|
Šmídová
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family [[Corydiidae]].
|
|-
|
''[[Ocelloblattula|Ocelloblattula margarita]]''<ref name=Fractalia />
|
Sp. nov
|
Valid
|
Koubová & Vršanský ''in'' Vršanský ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Corydioidea.
|
|-
|
''[[Olenablatta]]''<ref name=Olenablatta />
|
Gen. et sp. nov
|
In press
|
Šmídová
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family [[Olidae]]. Genus includes new species ''O. vrsanskyi''.
|
|-
|
''[[Otazka]]''<ref name=Afrophaga />
|
Gen. et sp. nov
|
In press
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of the family Mesoblattinidae. Genus includes new species ''O. systematicka''.
|
|-
|
''[[Pabuonqed|Pabuonqed magna]]''<ref>{{Cite journal|last1=Song |first1=Z. |last2=Xu |first2=C. |last3=Li |first3=J. |last4=Jarzembowski |first4=E. A. |last5=Wang |first5=B. |last6=Xiao |first6=C. |title=A new species of Pabuonqedidae (Blattaria: Mastotermitoidea) from mid-Cretaceous Kachin amber |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=53–59 |doi=10.1127/pala/2021/0111 |s2cid=240023055 }}</ref>
|
Sp. nov
|
In press
|
Song ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Mastotermitoidea]] belonging to the family [[Pabuonqedidae]].
|
|-
|
''[[Paekthoblatta]]''<ref>{{Cite journal|last1=So |first1=K. S. |last2=Won |first2=C. G. |last3=Ri |first3=C. J. |last4=Jon |first4=S. H. |last5=Ju |first5=I. Y. |title=''Paekthoblatta'', a New Predatory Cockroach Genus (Insecta: Blattaria: Raphidiomimidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea |year=2021 |journal=Paleontological Journal |volume=55 |issue=8 |pages=906–909 |doi=10.1134/S0031030121080074 |s2cid=245330342 }}</ref>
|
Gen. et sp. nov
|
Valid
|
So ''et al.''
|
Early Cretaceous
|
[[Sinuiju Formation]]
|
{{Flag|North Korea}}
|
A member of the family [[Raphidiomimidae]]. Genus includes new species ''P. coreanica''.
|
|-
|
[[Periplaneta perialla|''Periplaneta'' (?) ''perialla'']]<ref>{{Cite journal|last1=Anisyutkin |first1=L. N. |last2=Perkovsky |first2=E. E. |title=''Periplaneta'' (?) ''perialla'' sp. nov., a cockroach larva of the subfamily Blattinae (Dictyoptera, Blattidae) from Rovno amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=3 |pages=294–300 |doi=10.1134/S0031030121030047 |s2cid=235966737 |url=https://elibrary.ru/item.asp?id=45690953 }}</ref>
|
Sp. nov
|
Valid
|
Anisyutkin & Perkovsky
|
Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A cockroach, possibly a species of ''[[Periplaneta]]''.
|
|-
|
''[[Petropterix|Petropterix fukuiensis]]''<ref name=OyamaetalPAA>{{Cite journal|last1=Oyama |first1=N. |last2=Yukawa |first2=H. |last3=Imai |first3=T. |title=New cockroach assemblage from the Lower Cretaceous Kitadani Formation, Fukui, Japan |year=2021 |journal=Palaeontographica Abteilung A |volume=321 |issue=1–6 |pages=37–52 |doi=10.1127/pala/2021/0112 |s2cid=239140974 }}</ref>
|
Sp. nov
|
In press
|
Oyama, Yukawa & Imai
|
Early Cretaceous
|
[[Kitadani Formation]]
|
{{Flag|Japan}}
|
A member of the family [[Umenocoleidae]].
|
|-
|
''[[Praeblattella|Praeblattella arcuata]]''<ref name=OyamaetalPAA />
|
Sp. nov
|
In press
|
Oyama, Yukawa & Imai
|
Early Cretaceous
|
Kitadani Formation
|
{{Flag|Japan}}
|
A member of the family [[Mesoblattinidae]].
|
|-
|
''Praeblattella inexpecta''<ref name=OyamaetalPAA />
|
Sp. nov
|
In press
|
Oyama, Yukawa & Imai
|
Early Cretaceous
|
Kitadani Formation
|
{{Flag|Japan}}
|
A member of the family Mesoblattinidae.
|
|-
|
''[[Pseudomantina]]''<ref name=Fractalia />
|
Gen. et 2 sp. nov
|
Valid
|
Sendi & Vršanský ''in'' Vršanský ''et al.''
|
Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)
|
Lebanese amber
|
{{Flag|Kazakhstan}}<br>
{{Flag|Lebanon}}
|
A member of the family Liberiblattinidae. The type species is ''P. occisor'' Sendi; genus also includes new species ''P. nigroalba'' Vršanský.
|
|-
|
''[[Reticulitermes grimaldii]]''<ref name=Milesitermes />
|
Sp. nov
|
Valid
|
Jouault ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A termite, a species of ''[[Reticulitermes]]''.
|
|-
|
''[[Vrsanskysajda|Sajda]]''<ref name=Afrophaga />
|
Gen. et sp. nov
|
Junior homonym
|
Vršanský ''et al.''
|
Cretaceous (Hauterivian?)
|
|
{{Flag|Algeria}}
|
A member of [[Holocompsinae]]. Genus includes new species ''S. equatorialis''. The generic name is preoccupied by ''Sajda'' Dworakowska (1981); Jiang, Xing & Li (2023) coined a replacement name ''Vrsanskysajda''.<ref>{{Cite journal|last1=Jiang |first1=L. |last2=Xing |first2=J. |last3=Li |first3=Y. |year=2023 |title=New replacement name for the genus ''Sajda'' Vršanský, 2021 (Blattaria: Corydiidae: Holocompsinae) |journal=Zootaxa |volume=5270 |issue=2 |pages=343–344 |doi=10.11646/zootaxa.5270.2.10 |pmid=37518160 |s2cid=258274946 }}</ref>
|
|-
|
''[[Spinaeblattina|Spinaeblattina baekthoensis]]''<ref>{{Cite journal|last1=So |first1=K. S. |last2=Won |first2=C. G. |last3=Ri |first3=C. J. |last4=Jon |first4=S. H. |last5=Ju |first5=I. Y. |title=A New Species of ''Spinaeblattina'' Hinkelman, 2019 (Insecta, Blattaria, Mesoblattinidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea |year=2021 |journal=Paleontological Journal |volume=55 |issue=8 |pages=910–912 |doi=10.1134/S0031030121080086 |s2cid=245330596 }}</ref>
|
Sp. nov
|
Valid
|
So ''et al.''
|
Early Cretaceous
|
Sinuiju Formation
|
{{Flag|North Korea}}
|
A member of the family [[Mesoblattinidae]].
|
|-
|
''[[Stavba delicata]]''<ref name=Miniblattina />
|
Sp. nov
|
In press
|
Sendi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family Liberiblattinidae.
|
|-
|
''[[Stavba magnoculara]]''<ref name=Miniblattina />
|
Sp. nov
|
In press
|
Sendi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A cockroach belonging to the family Liberiblattinidae.
|
|-
|
''[[Tanytermitalis]]''<ref name=Arceotermes />
|
Gen. et sp. nov
|
Valid
|
Engel & Cai ''in'' Jiang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A termite belonging to the family [[Tanytermitidae]]. Genus includes new species ''T. philetaerus''.
|
|-
|}

===Dictyopteran research===
* A study on the phylogenetic placement of the family [[Umenocoleidae]], based on data from a new specimen of ''[[Umenocoleus]] sinuatus'' from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo ''et al.'' (2021).<ref>{{Cite journal|last1=Liuo |first1=C.-H. |last2=Beutel |first2=R. G. |last3=Thomson |first3=U. R |last4=Zheng |first4=D.-R. |last5=Li |first5=J.-H. |last6=Zhao |first6=X.-Y. |last7=Zhang |first7=H.-C. |last8=Wang |first8=B. |title=Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis |year=2021 |journal=Palaeoworld |volume=31 |pages=121–130 |doi=10.1016/j.palwor.2021.01.003 |s2cid=234208413 }}</ref>
* Description of new [[Alienopteridae|alienopterid]] [[Nymph (biology)|nymphs]] from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo ''et al.'' (2021).<ref>{{Cite journal|last1=Luo |first1=C. |last2=Beutel |first2=R. G. |last3=Engel |first3=M. S. |last4=Liang |first4=K. |last5=Li |first5=L. |last6=Li |first6=J. |last7=Xu |first7=C. |last8=Vršanský |first8=P. |last9=Jarzembowski |first9=E. A. |last10=Wang |first10=B. |year=2021 |title=Life history and evolution of the enigmatic Cretaceous-Eocene Alienopteridae: A critical review |journal=Earth-Science Reviews |volume=225 |pages=Article 103914 |doi=10.1016/j.earscirev.2021.103914 |s2cid=245601183 }}</ref>
* A study on the sensory organs of ''[[Huablattula]] hui'', and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi ''et al.'' (2021).<ref>{{Cite journal|last1=Taniguchi |first1=R. |last2=Nishino |first2=H. |last3=Watanabe |first3=H. |last4=Yamamoto |first4=S. |last5=Iba |first5=Y. |title=Reconstructing the ecology of a Cretaceous cockroach: destructive and high-resolution imaging of its micro sensory organs |year=2021 |journal=The Science of Nature |volume=108 |issue=5 |pages=Article number 45 |doi=10.1007/s00114-021-01755-9 |issn=0028-1042 |pmid=34581877 |bibcode=2021SciNa.108...45T |s2cid=238202024 }}</ref>

==Clade †Glosselytrodea==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Moltenojurina]]''<ref name=Moltenojurina>{{Cite journal|last1=Béthoux |first1=O. |last2=Anderson |first2=J. M. |title=The Polycytellidae viewed as Gondwanan Glosselytrodea |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=550–558 |doi=10.11646/palaeoentomology.4.6.5 |s2cid=245173292 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Béthoux & Anderson
|
Late Triassic ([[Carnian]])
|
Molteno Formation
|
{{Flag|South Africa}}
|
A [[Polycytellidae|polycytellid]] [[Glosselytrodea]]n.<br/> The type species is ''M. parva''.
|
|-
|
''[[Polycytella|Polycytella rasnitsyni]]''<ref name=Moltenojurina />
|
Sp. nov
|
Valid
|
Béthoux & Anderson
|
Late Triassic (Carnian)
|
Molteno Formation
|
{{Flag|South Africa}}
|
A polycytellid Glosselytrodea.
|
|-
|}

==Hymenopterans==
===New Hymenopteran taxa===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Aethemelikertes]]''<ref name=Aethemelikertes>{{Cite journal|last1=Engel |first1=M. S. |last2=Davis |first2=S. R. |title=New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae) |year=2021 |journal=Journal of Melittology |volume=103 |issue=103 |pages=1–52 |doi=10.17161/jom.i103.15655 |s2cid=237834516 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
[[Michael S. Engel|Engel]] ''in'' Engel & Davis
|
[[Eocene]]
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A bee belonging to the tribe [[Melikertini]]. The type species is ''A. emunctorii''.
|
|-
|
''[[Alivespa|Alivespa gracilenta]]''<ref name=CRVespidae>{{Cite journal|last1=Wu |first1=Q. |last2=Yang |first2=Y. |last3=Shih |first3=C. |last4=Zhao |first4=Y. |last5=Ren |first5=D. |title=Two new species of Vespidae (Hymenoptera) in mid-Cretaceous amber from northern Myanmar |year=2021 |journal=Cretaceous Research |volume=132 |pages=Article 105117 |doi=10.1016/j.cretres.2021.105117 }}</ref>
|
Sp. nov
|
In press
|
Wu ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Vespidae]].
|
|-
|
''[[Allommation]]''<ref name=Allommation>{{Cite journal|last1=Rosa |first1=B. B. |last2=Melo |first2=G. A. R. |title=Apoid wasps (Hymenoptera: Apoidea) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=122 |pages=Article 104770 |doi=10.1016/j.cretres.2021.104770 |bibcode=2021CrRes.12204770R |s2cid=234071940 }}</ref>
|
Fam. nov., Gen. et 3 sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Apoidea]] in the new family [[Allommationidae]]. Genus includes new species ''A. procerum'', ''A. elongatum'' and ''A. grande''.
|
|-
|
''[[Amelikertotes]]''<ref name=Aethemelikertes />
|
Gen. et comb. nov
|
Valid
|
Engel ''in'' Engel & Davis
|
Eocene
|
Baltic amber<br>
Bitterfeld amber
|
{{Flag|Germany}}<br>
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A bee belonging to the tribe Melikertini. The type species is ''"[[Melikertes (genus)|Melikertes]]" clypeatus'' Engel (2001).
|
|-
|
''[[Ammosphecium]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Crabroninae]]. Genus includes new species ''A. diabolicum''.
|
|-
|
''[[Anochetus miosumatrensis]]''<ref>{{Cite journal|last1=Ngô-Muller |first1=V. |last2=Garrouste |first2=R. |last3=Schubnel |first3=T. |last4=Pouillon |first4=J.-M. |last5=Christophersen |first5=V. |last6=Christophersen |first6=N. |last7=Nel |first7=A. |title=The first representative of the trap-jaw ant genus ''Anochetus'' Mayr, 1861 in Neogene amber from Sumatra (Hymenoptera: Formicidae) |year=2021 |journal=Comptes Rendus Palevol |volume=20 |issue=2 |pages=21–27 |doi=10.5852/cr-palevol2021v20a2 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Ngô-Muller, Garrouste & Nel ''in'' Ngô-Muller ''et al.''
|
[[Miocene]]
|
|
{{Flag|Indonesia}}
|
An ant, a species of ''[[Anochetus]]''.
|
|-
|
''[[Archaeovespa|Archaeovespa incompleta]]''<ref name=CRVespidae />
|
Sp. nov
|
In press
|
Wu ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Vespidae.
|
|-
|
''[[Archeonesia]]''<ref>{{Cite journal|last1=Tribull |first1=C. M. |last2=Pankowski |first2=M. V. |last3=Colombo |first3=W. D. |title=A new genus and species of Pristocerinae (Hymenoptera, Bethylidae) from upper Eocene Baltic amber with a review of conspecific association from insect fossils |year=2021 |journal=Journal of Hymenoptera Research |volume=85 |pages=119–133 |doi=10.3897/jhr.85.68658 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tribull, Pankowski & Colombo
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A [[Pristocerinae]] [[Bethylidae|Bethylid]].<br/> The type species is ''A. eocena''.
|
[[File:Archeonesia eocena Tribull, Pankowski & Colombo Fig1 A.jpg|thumb|center|upright|''[[Archeonesia eocena]]'']]
|-
|
''[[Arkadiserphus]]''<ref>{{Cite journal|last=Rasnitsyn |first=A. P. |title=First Jurassic representative of the extinct family Peleserphidae (Hymenoptera, Proctotrupoidea) |year=2021 |journal=Journal of Hymenoptera Research |volume=84 |pages=295–300 |doi=10.3897/jhr.84.65493 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
[[Alexandr Rasnitsyn|Rasnitsyn]]
|
[[Late Jurassic]]
|
[[Karabastau Formation]]
|
{{Flag|Kazakhstan}}
|
A member of [[Proctotrupoidea]] belonging to the family [[Peleserphidae]]. The type species is ''A. leleji''.
|
|-
|
''[[Ascogaster latitibialis|Ascogaster (Syntaphus) latitibialis]]''<ref name=Palaeocolastes>{{Cite journal|last1=Belokobylskij |first1=S. A. |last2=Pankowski |first2=M. G. |last3=Pankowski |first3=M. V. |last4=Zaldívar-Riverón |first4=A. |title=A new genus and three new species of fossil braconid wasps (Hymenoptera, Ichneumonoidea) from Eocene Baltic and Rovno ambers |year=2021 |journal=Journal of Paleontology |volume=95 |issue=6 |pages=1259–1272 |doi=10.1017/jpa.2021.47 |bibcode=2021JPal...95.1259B |s2cid=237921761 }}</ref>
|
Sp. nov
|
Valid
|
Belokobylskij ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Braconidae]] belonging to the subfamily [[Cheloninae]].
|
|-
|
''[[Aulacicosmus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Heterogynaidae]]. Genus includes new species ''A. setosus''.
|
|-
|
''[[Austroplebeia fujianica]]''<ref name=JM105>{{Cite journal|last1=Engel |first1=M. S. |last2=Herhold |first2=H. |last3=Davis |first3=S. |last4=Wang |first4=B. |last5=Thomas |first5=J. |title=Stingless bees in Miocene amber of southeastern China (Hymenoptera: Apidae) |year=2021 |journal=Journal of Melittology |volume=105 |issue=105 |pages=1–83 |doi=10.17161/jom.i105.15734 |s2cid=239647786 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Engel ''in'' Engel ''et al.''
|
[[Miocene]] ([[Langhian]])
|
Zhangpu amber
|
{{Flag|China}}
|
A stingless bee, a species of ''[[Austroplebeia]]''.
|
|-
|
''[[Burmabracon]]''<ref>{{Cite journal|last1=Li |first1=L. |last2=Shih |first2=C. |last3=Yang |first3=J. |last4=Wang |first4=L. |last5=Li |first5=D. |last6=Ren |first6=D. |title=New amber record of Braconidae (Insecta: Hymenoptera) from the mid-Cretaceous of Myanmar |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104794 |doi=10.1016/j.cretres.2021.104794 |bibcode=2021CrRes.12404794L |s2cid=233882169 }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Braconidae]] belonging to the subfamily [[Protorhyssalinae]]. Genus includes new species ''B. gracilens'' and ''B. grossus''.
|
|-
|
''[[Burmapyris]]''<ref name=Burmapyris>{{Cite journal|last1=Jouault |first1=C. |last2=Perrichot |first2=V. |last3=Nel |first3=A. |title=New flat wasps from mid-Cretaceous Burmese amber deposits highlight the bethylid antiquity and paleobiogeography (Hymenoptera: Chrysidoidea) |year=2021 |journal=Cretaceous Research |volume=123 |pages=Article 104772 |doi=10.1016/j.cretres.2021.104772 |bibcode=2021CrRes.12304772J |s2cid=234041758 |url=https://hal-insu.archives-ouvertes.fr/insu-03132639/file/jouault-2021.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault, Perrichot & Nel
|
Cretaceous (Albian-Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Bethylidae]] belonging to the subfamily [[Lancepyrinae]]. Genus includes new species ''B. azevedoi''.
|
|-
|
''[[Burmoxyela]]''<ref name=Burmoxyela>{{Cite journal|last1=Zheng |first1=Y. |last2=Hu |first2=H. |last3=Zhang |first3=H. |last4=Chen |first4=J. |last5=Rasnitsyn |first5=A. P. |last6=Zhuo |first6=D. |title=New genus and species of syspastoxyelid sawflies (Insecta, Hymenoptera) from the mid-Cretaceous Kachin amber with a review of the family Syspastoxyelidae |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104940 |doi=10.1016/j.cretres.2021.104940 |bibcode=2021CrRes.12704940Z }}</ref>
|
Gen. et sp. nov
|
Valid
|
Zheng ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A sawfly belonging to the family [[Syspastoxyelidae]]. Genus includes new species ''B. lii''.
|
|-
|
''[[Burmusculus|Burmusculus angustus]]''<ref>{{Cite journal|last1=Li |first1=L. |last2=Shih |first2=C. |last3=Yang |first3=J. |last4=Gao |first4=Y. |last5=Ren |first5=D. |title=Two new wasps from mid-Cretaceous Myanmar amber (Hymenoptera: Apocrita) |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105115 |doi=10.1016/j.cretres.2021.105115 |s2cid=244885445 }}</ref>
|
Sp. nov
|
In press
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Apocrita]] belonging to the family [[Burmusculidae]].
|
|-
|
''[[Buserphites]]''<ref name=Buserphites>{{Cite journal|last1=Herbert |first1=M. C. M. |last2=McKellar |first2=R. C. |title=New genera ''Buserphites'' and ''Mesoserphites'' (Hymenoptera: Serphitidae) from mid-Cretaceous amber of Myanmar |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105025 |doi=10.1016/j.cretres.2021.105025 |s2cid=240331883 |url=https://hal-insu.archives-ouvertes.fr/insu-03418652/file/Herbert-2021.pdf }}</ref>
|
Gen. et 2 sp. nov
|
In press
|
Herbert & McKellar
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Serphitidae]]. Genus includes new species ''B. applanatus'' and ''B. myanmarensis''.
|
|-
|
''[[Chainochora]]''<ref name=Chainochora>{{Cite journal|last1=Chen |first1=H. |last2=van Achterberg |first2=C. |last3=Hong |first3=P. |title=Two new genera of Protobraconinae (Hymenoptera, Braconidae) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104914 |doi=10.1016/j.cretres.2021.104914 |bibcode=2021CrRes.12604914C }}</ref>
|
Gen. et sp. nov
|
Valid
|
Chen, van Achterberg & Hong
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Braconidae]] belonging to the subfamily [[Protobraconinae]]. The type species is ''C. syntoma''.
|
|-
|
''[[Chlorepyris deploegi]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene ([[Priabonian]])
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family [[Bethylidae]].
|
|-
|
''[[Chlorepyris engeli]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris hopei]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris jouaulti]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris mckellari]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris meunieri]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris neli]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber<br>
Rovno amber
|
{{Flag|Russia}}<br>
{{Flag|Ukraine}}
|
A member of the family Bethylidae.
|
|-
|
''[[Chlorepyris perrichoti]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae.
|
|-
|
''[[Christophus]]''<ref name=Christophus>{{Cite journal|last1=Brazidec |first1=M. |last2=Perrichot |first2=V. |last3=Nel |first3=A. |last4=Jouault |first4=C. |title=Two new genera of belytine wasps (Hymenoptera: Diapriidae) from Cretaceous Burmese amber |year=2021 |journal=Proceedings of the Geologists' Association |volume=132 |issue=5 |pages=597–604 |doi=10.1016/j.pgeola.2021.09.006 |bibcode=2021PrGA..132..597B |s2cid=244613026 |url=https://hal-insu.archives-ouvertes.fr/insu-03380285/file/Brazidec_etal_2021_Belytinae%20from%20Burmese%20amber_revised.pdf }}</ref>
|
Gen. et 2 sp. nov
|
In press
|
Brazidec ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Diapriidae]]. Genus includes new species ''C. myanmarensis'' and ''C. ohmkuhnlei''.
|
|-
|
''[[Cretastatus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Astatinae]]. Genus includes new species ''C. longevus''.
|
|-
|
''[[Cretorussus]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Perrichot |first2=V. |last3=Nel |first3=A. |title=Taxonomic addition to the early diverged parasitic wasps (Hymenoptera: Orussoidea) from the mid-Cretaceous Burmese amber |year=2021 |journal=Proceedings of the Geologists' Association |volume=132 |issue=3 |pages=324–331 |doi=10.1016/j.pgeola.2021.02.002 |bibcode=2021PrGA..132..324J |s2cid=234848669 |url=https://hal-insu.archives-ouvertes.fr/insu-03203872/file/jouault-2021.pdf }}</ref>
|
Gen. et sp. nov
|
In press
|
Jouault, Perrichot & Nel
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Orussoidea]]. Genus includes new species ''C. vilhelmseni''.
|
|-
|
''[[Curiosivespa|Curiosivespa sennlaubi]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Rasnitsyn |first2=A. P. |last3=Pouillon |first3=J.-M. |last4=Nel |first4=A. |year=2021 |title=The first ''Curiosivespa'' Rasnitsyn, 1975 (Hymenoptera: Vespidae) from the Lower Cretaceous Crato formation (Brazil) |journal=Annales de la Société Entomologique de France |series=Nouvelle Série |volume= 57|issue=6 |pages=509–513 |doi=10.1080/00379271.2021.1976672 |s2cid=244229524 }}</ref>
|
Sp. nov
|
Valid
|
Jouault, Rasnitsyn & Nel ''in'' Jouault ''et al.''
|
Early Cretaceous
|
Crato Formation
|
{{Flag|Brazil}}
|
A member of the family [[Vespidae]].
|
|-
|
''[[Diameneura]]''<ref>{{Cite journal|last1=Santer |first1=M. |last2=Álvarez-Parra |first2=S. |last3=Nel |first3=A. |last4=Peñalver |first4=E. |last5=Delclòs |first5=X. |title=New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea) |year=2021 |journal=Cretaceous Research |volume=133 |pages=Article 105128 |doi=10.1016/j.cretres.2021.105128 |s2cid=245556639 |doi-access=free |hdl=2445/182542 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
In press
|
Santer & Álvarez-Parra ''in'' Santer ''et al.''
|
Early Cretaceous (Albian)
|
[[Escucha Formation]]
|
{{Flag|Spain}}
|
A member of the family [[Spathiopterygidae]]. The type species is ''D. marveni''.
|
|-
|
''[[Drymomyrmex rasnitsyni]]''<ref>{{Cite journal|last=Radchenko |first=A. G. |title=New species of the fossil ant genus ''Drymomyrmex'' (Hymenoptera, Formicidae, Formicinae) from the late Eocene Rovno amber (Ukraine) |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=544–549 |doi=10.11646/palaeoentomology.4.6.4 |s2cid=245149637 }}</ref>
|
Sp. nov
|
Valid
|
Radchenko
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
An ant belonging to the subfamily [[Formicinae]].
|
|-
|
''[[Elasmosomites|Elasmosomites arkadyleleji]]''<ref>{{Cite journal|last1=Belokobylskij |first1=S. A. |last2=Dubovikoff |first2=D. A. |last3=Manukyan |first3=A. R. |last4=Zharkov |first4=D. M. |title=Braconid parasitoids of ants (Hymenoptera, Braconidae, Euphorinae, Neoneurini) from Baltic amber with a discussion of records of fossil larvae parasitizing ant workers |year=2021 |journal=Journal of Hymenoptera Research |volume=84 |pages=29–43 |doi=10.3897/jhr.84.67749 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Belokobylskij ''in'' Belokobylskij ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Braconidae]] belonging to the subfamily [[Euphorinae]].
|
|-
|
''[[Electrofoenops|Electrofoenops exaltatus]]''<ref name=CRLietalAulacidae>{{Cite journal|last1=Li |first1=L. |last2=Shih |first2=P. J. M. |last3=Zhang |first3=X. |last4=Peng |first4=C. |last5=Li |first5=D. |last6=Ren |first6=D. |title=Geometric Morphometric Analysis of two genera confirm three new wasps from the mid-Cretaceous of Myanmar (Hymenoptera: Aulacidae) |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105057 |doi=10.1016/j.cretres.2021.105057 |s2cid=242552161 }}</ref>
|
Sp. nov
|
In press
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Aulacidae]].
|
|-
|
''[[Embolemus brachypterus]]''<ref name=Palaeoentomology41Embolemus>{{Cite journal|last1=Olmi |first1=M. |last2=Guglielmino |first2=A. |last3=Müller |first3=P. |last4=Capradossi |first4=L. |last5=Perkovsky |first5=E. E. |title=Descriptions of the first females of ''Embolemus'' (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=44–49 |doi=10.11646/palaeoentomology.4.1.9 |s2cid=233944242 }}</ref>
|
Sp. nov
|
Valid
|
Olmi ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Embolemidae]].
|
|-
|
''[[Embolemus micropterus]]''<ref name=Palaeoentomology41Embolemus />
|
Sp. nov
|
Valid
|
Olmi ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Embolemidae.
|
|-
|
''[[Eopelecinus|Eopelecinus marechali]]''<ref>{{Cite journal|last=Jouault |first=C. |title=Mid-Cretaceous Burmese amber pelecinid wasps (Hymenoptera, Pelecinidae) support the hypothesis of an Asian origin of the family |year=2021 |journal=Annales de Paléontologie |volume=107 |issue=1 |pages=Article 102464 |doi=10.1016/j.annpal.2020.102464 |bibcode=2021AnPal.10702464J |s2cid=234321579 |url=https://hal-insu.archives-ouvertes.fr/insu-03142566/file/Jouault_Eopelecinus_marechalivf.pdf }}</ref>
|
Sp. nov
|
Valid
|
Jouault
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Pelecinidae]].
|
|-
|
''[[Eoteredon]]''<ref>{{Cite journal|last1=Archibald |first1=S. B. |last2=Aase |first2=A. |last3=Nel |first3=A. |year=2021 |title=The second North American fossil horntail wood-wasp (Hymenoptera: Siricidae), from the early Eocene Green River Formation |journal=Zootaxa |volume=4999 |issue=4 |pages=325–334 |doi=10.11646/zootaxa.4999.4.2 |pmid=34810481 |s2cid=237762599 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Archibald, Aase & Nel
|
Early Eocene
|
[[Green River Formation]]
|
{{Flag|United States}}<br>({{Flag|Wyoming}})
|
A [[horntail]]. Genus includes new species ''E. lacoi''.
|
|-
|
''[[Ephedrus rasnitsyni]]''<ref>{{Cite journal|last1=Davidian |first1=E. M. |last2=Kaliuzhna |first2=M. O. |last3=Perkovsky |first3=E. E. |title=First aphidiine wasp from the Sakhalinian amber |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=Supplement to 3 |pages=S59–S65 |doi=10.4202/app.00843.2020 |s2cid=236723591 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Davidian & Kaliuzhna ''in'' Davidian, Kaliuzhna & Perkovsky
|
Middle Eocene
|
Sakhalinian amber
|
{{Flag|Russia}}
|
A member of the family [[Braconidae]] belonging to the subfamily [[Aphidiinae]].
|
|-
|
''[[Exilaulacus|Exilaulacus advenus]]''<ref name=CRLietalAulacidae />
|
Sp. nov
|
In press
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Aulacidae.
|
|-
|
''Exilaulacus eximius''<ref name=CRLietalAulacidae />
|
Sp. nov
|
In press
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Aulacidae.
|
|-
|
''[[Formica ribbeckei]]''<ref>{{Cite journal|last1=Radchenko |first1=A. G. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |title=''Formica'' species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber |year=2021 |journal=Journal of Hymenoptera Research |volume=82 |pages=237–251 |doi=10.3897/jhr.82.64599 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Radchenko & Perkovsky ''in'' Radchenko, Perkovsky & Vasilenko
|
Late Eocene
|
[[Rovno amber]]
|
{{Flag|Ukraine}}
|
An ant, a species of ''[[Formica]]''.
|
|-
|
''[[Glenocephalus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Apoidea in the new family [[Cirrosphecidae]]. Genus includes new species ''G. mandibularis''.
|
|-
|
''[[Gloxinius]]''<ref name=Gloxinius>{{Cite journal|last1=Colombo |first1=W. D. |last2=Perkovsky |first2=E. E. |last3=Waichert |first3=C. |last4=Azevedo |first4=C. O. |title=Synopsis of the fossil flat wasps Epyrinae (Hymenoptera, Bethylidae), with description of three new genera and 10 new species |year=2021 |journal=Journal of Systematic Palaeontology |volume=19 |issue=1 |pages=39–89 |doi=10.1080/14772019.2021.1882593 |s2cid=232313171 }}</ref>
|
Gen. et comb. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae; a new genus for ''"Epyris" bifossatus'' Brues (1939).
|
|-
|
''[[Grandixyela]]''<ref>{{Cite journal|last1=Zheng |first1=Y. |last2=Kopylov |first2=D. S. |last3=Zhang |first3=Q. |last4=Zhang |first4=H.-G. |last5=Qin |first5=M. |last6=Chen |first6=J. |last7=Zhang |first7=H.-C. |title=Discovery of a new symphytan wasp in the mid-Cretaceous Kachin amber (Hymenoptera, Syspastoxyelidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=629–636 |doi=10.11646/palaeoentomology.4.6.12 |s2cid=245173388 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Zheng ''in'' Zheng ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Syspastoxyelidae]]. The type species is ''G. rasnitsyni''.
|
|-
|
''[[Haidomelikertes]]''<ref name=Aethemelikertes />
|
Gen. et sp. nov
|
Valid
|
Engel ''in'' Engel & Davis
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A bee belonging to the tribe Melikertini. The type species is ''H. uraeus''.
|
|-
|
''[[Haptodioctes]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Apoidea in the new family [[Cirrosphecidae]]. Genus includes new species ''H. apiformis''.
|
|-
|
''[[Helosericus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Psenini]]. Genus includes new species ''H. verrucosus''.
|
|-
|
''[[Heteroichneumon]]''<ref name=Heteroichneumon>{{Cite journal|last1=Kopylov |first1=D. S. |last2=Zhang |first2=Q. |last3=Zhang |first3=H.-C. |title=The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=592–603 |doi=10.11646/palaeoentomology.4.6.8 |s2cid=245157688 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kopylov, Zhang & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Ichneumonidae]]. The type species is ''H. rasnitsyni''.
|
|-
|
''[[Heteropimpla|Heteropimpla pulverulenta]]''<ref name=Heteroichneumon />
|
Sp. nov
|
Valid
|
Kopylov, Zhang & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Ichneumonidae.
|
|-
|
''[[Holepyris rasnitsyni]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae.
|
|-
|
''[[Holepyris terayamai]]''<ref name=Gloxinius />
|
Sp. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Bethylidae.
|
|-
|
''[[Holopsenella|Holopsenella antiqua]]''<ref name=CRHolopsenella>{{Cite journal|last1=Lepeco |first1=A. |last2=Melo |first2=G. A. R. |title=The wasp genus †''Holopsenella'' in mid-Cretaceous Burmese amber (Hymenoptera: †Holopsenellidae stat. nov.) |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105089 |doi=10.1016/j.cretres.2021.105089|issn=0195-6671  |s2cid=244055567 }}</ref>
|
Sp. nov
|
In press
|
Lepeco & Melo
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Aculeata]] of uncertain phylogenetic placement.
|
|-
|
''Holopsenella burmitica''<ref name=CRHolopsenella />
|
Sp. nov
|
In press
|
Lepeco & Melo
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Aculeata of uncertain phylogenetic placement.
|
|-
|
''Holopsenella gothica''<ref name=CRHolopsenella />
|
Sp. nov
|
In press
|
Lepeco & Melo
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Aculeata of uncertain phylogenetic placement.
|
|-
|
''[[Hybristodryinus|Hybristodryinus castaneus]]''<ref name=JHR81>{{Cite journal|last1=Olmi |first1=M. |last2=Chen |first2=H.-Y. |last3=Shih |first3=C. |last4=Müller |first4=P. |last5=Capradossi |first5=L. |last6=Ren |first6=D. |last7=Perkovsky |first7=E. E. |last8=Guglielmino |first8=A. |title=New species of ''Hybristodryinus'' Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts |year=2021 |journal=Journal of Hymenoptera Research |volume=81 |pages=43–55 |doi=10.3897/jhr.81.57792 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Olmi, Guglielmino & Chen ''in'' Olmi ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Dryinidae]].
|
|-
|
''Hybristodryinus viriosus''<ref>{{Cite journal|last1=Wang |first1=Z. |last2=Olmi |first2=M. |last3=Shih |first3=C. |last4=Ren |first4=D. |last5=Gao |first5=T. |title=New species of ''Hybristodryinus'' from mid-Cretaceous Myanmar amber (Hymenoptera, Dryinidae) |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104861 |doi=10.1016/j.cretres.2021.104861|bibcode=2021CrRes.12504861W | issn=0195-6671 }}</ref>
|
Sp. nov
|
Valid
|
Wang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Dryinidae.
|
|-
|
''Hybristodryinus  zaifui''<ref name=JHR81 />
|
Sp. nov
|
Valid
|
Chen, Olmi & Perkovsky ''in'' Olmi ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Dryinidae.
|
|-
|
''[[Janzenella theia]]''<ref>{{Cite journal|last1=Bremer |first1=J. |last2=van de Kamp |first2=T. |last3=Talamas |first3=E. J. |title=''Janzenella theia'' Bremer & Talamas (Platygastroidea, Janzenellidae): a new species from Baltic amber |year=2021 |journal=Journal of Hymenoptera Research |volume=87 |pages=223–233 |doi=10.3897/jhr.87.67256 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Bremer & Talamas ''in'' Bremer, van de Kamp & Talamas
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A species of ''[[Janzenella]]''.
|
|-
|
''[[Karataus|Karataus ryonsangensis]]''<ref name=WonSoCRwasps>{{Cite journal|last1=Won |first1=C. G. |last2=So |first2=K. S. |title=Two new wasps (Insecta: Hymenoptera) from the Lower Cretaceous Sinuiju Formation of Ryonsang-dong, North Phyongan Province, Democratic People's Republic of Korea |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105086 |doi=10.1016/j.cretres.2021.105086 |s2cid=243805918 }}</ref>
|
Sp. nov
|
In press
|
Won & So
|
Early Cretaceous ([[Barremian]]–[[Aptian]])
|
[[Sinuiju Formation]]
|
{{Flag|North Korea}}
|
A wasp belonging to the family [[Ephialtitidae]].
|
|-
|
''[[Keratodellitha]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Maréchal |first2=A. |last3=Condamine |first3=F. L. |last4=Wang |first4=B. |last5=Nel |first5=A. |last6=Legendre |first6=F. |last7=Perrichot |first7=V. |title=Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth–death process |year=2022|volume=194|issue=4|pages=1396–1423 |journal=Zoological Journal of the Linnean Society |doi=10.1093/zoolinnean/zlab034| url= https://doi.org/10.1093/zoolinnean/zlab034}}</ref>
|
Gen. et 3 sp. nov
|
Valid
|
Jouault ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Evanioidea]] belonging to the family [[Othniodellithidae]]. Genus includes new species ''K. anubis'', ''K. basilisci'' and ''K. kirin''.
|
|-
|
''[[Kleistochora]]''<ref name=Chainochora />
|
Gen. et sp. nov
|
Valid
|
Chen, van Achterberg & Hong
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is ''K. dolichura''.
|
|-
|
''[[Lelejus]]''<ref>{{Cite journal|last1=Radchenko |first1=A. G. |last2=Proshchalykin |first2=M. Yu. |title=New extinct ant genus (Hymenoptera, Formicidae, Myrmicinae) from late Eocene Rovno amber |year=2021 |journal=Journal of Hymenoptera Research |volume=84 |pages=271–282 |doi=10.3897/jhr.84.65238 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Radchenko & Proshchalykin
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
An ant belonging to the subfamily [[Myrmicinae]]. The type species is ''L. venustus''.
|
|-
|
''[[Lonchodryinus groehni]]''<ref>{{Cite journal|last1=Olmi |first1=M. |last2=Vasilenko |first2=D. V. |last3=Capradossi |first3=L. |last4=Perkovsky |first4=E. E. |last5=Guglielmino |first5=A. |year=2021 |title=A new species of ''Lonchodryinus'' (Hymenoptera, Dryinidae) from Upper Eocene Baltic amber |journal=Zootaxa |volume=5020 |issue=2 |pages=328–336 |doi=10.11646/zootaxa.5020.2.5 |pmid=34810999 |s2cid=238658382 }}</ref>
|
Sp. nov
|
Valid
|
Olmi ''et al.''
|
Late Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Dryinidae]].
|
|-
|
''[[Mael longiceps|Mael]]''<ref name=Gloxinius />
|
Gen. et comb. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Eocene (Priabonian)
|
Baltic amber
|
{{Flag|Russia}}
|
A member of the family Bethylidae belonging to the subfamily [[Scleroderminae]]; a new genus for ''"Calyoza" longiceps'' Brues (1923).
|
|-
|
''[[Megalyrhyssalus]]''<ref name=Megalyrhyssalus>{{Cite journal|last1=Belokobylskij |first1=S. A. |last2=Jouault |first2=C. |title=Two new striking braconid genera (Hymenoptera: Braconidae) from the mid-Cretaceous Burmese amber |year=2021 |journal=Proceedings of the Geologists' Association |volume=132 |issue=4 |pages=426–437 |doi=10.1016/j.pgeola.2021.04.003 |bibcode=2021PrGA..132..426B |s2cid=236271642 |url=https://hal-insu.archives-ouvertes.fr/insu-03206260 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Belokobylskij & Jouault
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Braconidae]]. Genus includes new species ''M. clavicornis''.
|
|-
|
''[[Megarolium]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Bembicinae]]. Genus includes new species ''M. paradoxum''.
|
|-
|
''[[Merascylla]]''<ref name=Gloxinius />
|
Gen. et comb. nov
|
Valid
|
Colombo & Azevedo ''in'' Colombo ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Bethylidae belonging to the subfamily [[Pristocerinae]]; a new genus for ''"Epyris" atavellus'' Cockerell (1920).
|
|-
|
''[[Mesoserphites]]''<ref name=Buserphites />
|
Gen. et 5 sp. nov
|
In press
|
Herbert & McKellar
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Serphitidae. Genus includes new species ''M. annulus'', ''M. giganteus'', ''M. engeli'', ''M. scutatus'' and ''M. viraneacapitis''.
|
|-
|
''[[Meteorus arasnitsyni]]''<ref name=Palaeocolastes />
|
Sp. nov
|
Valid
|
Belokobylskij ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A species of ''[[Meteorus]]''.
|
|-
|
''[[Microtypus eocenus]]''<ref name=Palaeocolastes />
|
Sp. nov
|
Valid
|
Belokobylskij ''et al.''
|
Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Braconidae belonging to the subfamily [[Microtypinae]].
|
|-
|
''[[Mintara parva|Mintara]]''<ref name=Christophus />
|
Gen. et sp. nov
|
In press
|
Brazidec ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Diapriidae. Genus includes new species ''M. parva''.
|
|-
|
''[[Odontosericus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Psenini. Genus includes new species ''O. burmensis''.
|
|-
|
''[[Orthosyntexis]]''<ref>{{Cite journal|last1=Gao |first1=J. |last2=Engel |first2=M. S. |last3=Shih |first3=C. |last4=Ren |first4=D. |last5=Gao |first5=T. |title=A new genus of anaxyelid wood wasps from the mid-Cretaceous and the phylogeny of Anaxyelidae (Hymenoptera) |year=2021 |journal=Journal of Hymenoptera Research |volume=86 |pages=151–169 |doi=10.3897/jhr.86.73161 |doi-access=free }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Gao, Engel, Shih, Gao ''in'' Gao ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Anaxyelidae]]. The type species is ''O. elegans''; genus also includes ''O. thanti''.
|
|-
|
''[[Palaeocolastes]]''<ref name=Palaeocolastes />
|
Gen. et sp. nov
|
Valid
|
Belokobylskij ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family Braconidae belonging to the subfamily [[Exothecinae]]. The type species is ''P. bruesi''.
|
|-
|
''[[Paleoaulacus]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Nel |first2=A. |title=A new genus and species of Aulacidae (Hymenoptera: Evanioidea) from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104799 |doi=10.1016/j.cretres.2021.104799 |bibcode=2021CrRes.12404799J |s2cid=233846750 |url=https://hal-insu.archives-ouvertes.fr/insu-03161875/file/1-s2.0-S019566712100046X-main.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault & Nel
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Aulacidae]]. Genus includes new species ''P. minutus''.
|
|-
|
''[[Paleoxyela]]''<ref name=Paleoxyela>{{Cite journal|last1=Jouault |first1=C. |last2=Aase |first2=A. |last3=Nel |first3=A. |title=Past ecosystems drive the evolution of the early diverged Symphyta (Hymenoptera: Xyelidae) since the earliest Eocene |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=379–393 |doi=10.5194/fr-24-379-2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault, Aase & Nel
|
Eocene ([[Chadronian]])
|
[[Florissant Formation]]
|
{{Flag|United States}}<br>({{Flag|Colorado}})
|
A member of the family [[Xyelidae]]. The type species is ''P. nearctica''.
|
|-
|
''[[Peleserphus qufuensis]]''<ref>{{cite book |last1=Yu |first1=Y. |last2=Hu |first2=G. |last3=Zhu |first3=Y. |last4=Li |first4=Y. |last5=Jiang |first5=W. |last6=Rasnitsyn |first6=A. P. |last7=Zhang |first7=H. |last8=Zhang |first8=Q. |year=2021 |chapter=New material of Peleserphidae (Proctotrupoidea, Hymenoptera) in the mid-Cretaceous Burmese (Kachin) Amber |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |publisher=The Geological Society of London |doi=10.1144/SP521-2021-128 |s2cid=244645180 }}</ref>
|
Sp. nov
|
In press
|
Yu ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Peleserphidae]].
|
|-
|
''[[Pheidole pauchil]]''<ref>{{Cite journal|last1=Varela-Hernández |first1=F. |last2=Riquelme |first2=F. |title=A New Ant Species of the Genus ''Pheidole'' Westwood, 1839 from Miocene Mexican Amber |year=2021 |journal=Southwestern Entomologist |volume=46 |issue=1 |pages=75–82 |doi=10.3958/059.046.0107 |s2cid=232357684 }}</ref>
|
Sp. nov
|
Valid
|
Varela-Hernández & Riquelme
|
Miocene
|
[[Mexican amber]]
|
{{Flag|Mexico}}
|
An ant, a species of ''[[Pheidole]]''.
|
|-
|
''[[Pinguixyela]]''<ref name=Burmoxyela />
|
Gen. et comb. nov
|
Valid
|
Zheng ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A sawfly belonging to the family Syspastoxyelidae; a new genus for ''"[[Syspastoxyela]]" pinguis''.
|
|-
|
''[[Platyxyela|Platyxyela tenuis]]''<ref name=Scleroxyela>{{Cite journal|last1=Zheng |first1=Y. |last2=Hu |first2=H. |last3=Chen |first3=D. |last4=Chen |first4=J. |last5=Zhang |first5=H. |last6=Rasnitsyn |first6=A. P. |title=New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China |year=2021 |journal=European Journal of Taxonomy |issue=733 |pages=146–159 |doi=10.5852/ejt.2021.733.1229 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Zheng ''et al.''
|
[[Middle Jurassic]] ([[Callovian]]-[[Bathonian]] boundary)
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A member of the family [[Xyelidae]].
|
|-
|
''[[Pompilopterus|Pompilopterus ryonsangensis]]''<ref name=WonSoCRwasps />
|
Sp. nov
|
In press
|
Won & So
|
Early Cretaceous (Barremian–Aptian)
|
Sinuiju Formation
|
{{Flag|North Korea}}
|
A wasp belonging to the family [[Angarosphecidae]].
|
|-
|
''[[Pristinopterus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Bembicinae. Genus includes new species ''P. spectabilis''.
|
|-
|
''[[Pristopsen]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Psenini. Genus includes new species ''P. obscurus''.
|
|-
|
''[[Prosyntexis|Prosyntexis sennlaubi]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Nel |first2=A. |title=A new species of ''Prosyntexis'' Sharkey, 1990 (Hymenoptera: Sepulcidae) from the Lower Cretaceous Crato Formation of Brazil confirmed by geometric morphometric analysis |year=2021 |journal=Palaeoentomology |volume=4 |issue=2 |pages=171–177 |doi=10.11646/palaeoentomology.4.2.6 |s2cid=235511059 |url=https://hal-insu.archives-ouvertes.fr/insu-03206270/document }}</ref>
|
Sp. nov
|
Valid
|
Jouault & Nel
|
Early Cretaceous ([[Aptian]])
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A member of the family [[Sepulcidae]].
|
|-
|
''[[Protopyris]]''<ref name=Burmapyris />
|
Gen. et sp. nov
|
Valid
|
Jouault & Nel ''in'' Jouault, Perrichot & Nel
|
[[Early Cretaceous]] ([[Albian]])
|
Hkamti amber
|
{{Flag|Myanmar}}
|
A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species ''P. myanmarensis''.
|
|-
|
''[[Prototeleia]]''<ref>{{Cite journal|last1=Talamas |first1=E. J. |last2=Popovici |first2=O. |last3=Shih |first3=C. |last4=Ren |first4=D. |title=''Prototeleia'' Talamas, Popovici, Shih & Ren: A new genus of Platygastridae from Burmese amber |year=2021 |journal=Journal of Hymenoptera Research |volume=87 |pages=67–80 |doi=10.3897/jhr.87.65472 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Talamas ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Platygastridae]]. The type species is ''P. kleio''.
|
|-
|
''[[Proxyelia]]''<ref name=Paleoxyela />
|
Gen. et sp. nov
|
Valid
|
Jouault, Aase & Nel
|
Eocene ([[Ypresian]])
|
[[Green River Formation]]
|
{{Flag|United States}}<br>({{Flag|Wyoming}})
|
A member of the family Xyelidae. The type species is ''P. pankowskii''.
|
|-
|
''[[Ptilocosmus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Heterogynaidae. Genus includes new species ''P. corniculatus''.
|
|-
|
''[[Ramageoptera]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Brazidec |first2=M. |title=A new protopristocerine wasp (Hymenoptera: Bethylidae) from the mid-Cretaceous Burmese amber |year=2021 |journal=Annales de Paléontologie |volume=107 |issue=4 |pages=Article 102522 |doi=10.1016/j.annpal.2021.102522 |bibcode=2021AnPal.10702522J |s2cid=244791642 |url=https://hal-insu.archives-ouvertes.fr/insu-03457367/file/BU-Ramageoptera.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault & Brazidec
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Bethylidae belonging to the subfamily [[Protopristocerinae]]. Genus includes new species ''R. platycephala''.
|
|-
|
''[[Rasnichneumon]]''<ref name=Heteroichneumon />
|
Gen. et 2 sp. nov
|
Valid
|
Kopylov, Zhang & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Ichneumonidae. The type species is ''R. alexandri''; genus also includes ''R. gracilis''.
|
|-
|
''[[Rasnitsynum]]''<ref>{{Cite journal|last1=Olmi |first1=M. |last2=Guglielmino |first2=A. |last3=Parise |first3=G. |last4=Capradossi |first4=L. |last5=Perkovsky |first5=E. E. |title=Discovery of a new genus and species of Dryinidae (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber: ''Rasnitsynum burmense'' gen. et sp. nov. |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=416–420 |doi=10.11646/palaeoentomology.4.5.3 |s2cid=240513429 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Olmi ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Dryinidae]]. Genus includes new species ''R. burmense''.
|
|-
|
''[[Rogichneumon]]''<ref name=Heteroichneumon />
|
Gen. et sp. nov
|
Valid
|
Kopylov, Zhang & Zhang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Ichneumonidae. The type species is ''R. braconidicus''.
|
|-
|
''[[Rovenosa alexrasnitsyni]]''<ref name=Palaeoentomology46Rovenosa>{{Cite journal|last1=Manukyan |first1=A. R. |last2=Zhindarev |first2=L. A. |title=Fossil Darwin wasps (Hymenoptera: Ichneumonidae) from Baltic amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=637–647 |doi=10.11646/palaeoentomology.4.6.13 |s2cid=245191818 }}</ref>
|
Sp. nov
|
Valid
|
Manukyan ''in'' Manukyan & Zhindarev
|
Late Eocene
|
Prussian Formation<br>(Baltic amber)
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of the family Ichneumonidae.
|
|-
|
''[[Rovenosa khalaimi]]''<ref name=Palaeoentomology46Rovenosa />
|
Sp. nov
|
Valid
|
Manukyan ''in'' Manukyan & Zhindarev
|
Late Eocene
|
Prussian Formation<br>(Baltic amber)
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A member of the family Ichneumonidae.
|
|-
|
''[[Sakhalinencyrtus]]''<ref>{{Cite journal|last1=Simutnik |first1=S. A. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |title=''Sakhalinencyrtus leleji'' Simutnik gen. et sp. nov. of earliest Encyrtidae (Hymenoptera, Chalcidoidea) from Sakhalinian amber |year=2021 |journal=Journal of Hymenoptera Research |volume=84 |pages=361–372 |doi=10.3897/jhr.84.66367 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Simutnik ''in'' Simutnik, Perkovsky & Vasilenko
|
Middle Eocene
|
Sakhalinian amber
|
{{Flag|Russia}}
|
A member of the family [[Encyrtidae]]. The type species is ''S. leleji''.
|
|-
|
''[[Sakhalinoctonus]]''<ref>{{Cite journal|last1=Davidian |first1=E. M. |last2=Manukyan |first2=A. R. |last3=Belokobylskij |first3=S. A. |title=A new genus and species of the aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) from the Eocene Sakhalinian amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=537–543 |doi=10.11646/palaeoentomology.4.6.3 |s2cid=245166207 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Davidian ''in'' Davidian, Manukyan & Belokobylskij
|
Middle Eocene
|
Sakhalinian amber
|
{{Flag|Russia}}
|
A member of the family [[Braconidae]] belonging to the subfamily [[Aphidiinae]]. Genus includes new species ''S. alexrasnitsyni''.
|
|-
|
''[[Scleroxyela]]''<ref name=Scleroxyela />
|
Gen. et sp. nov
|
Valid
|
Zheng ''et al.''
|
Middle Jurassic (Callovian-Bathonian boundary)
|
Jiulongshan Formation
|
{{Flag|China}}
|
A member of the family Xyelidae. The type species is ''S. daohugouensis''.
|
|-
|
''[[Spalangiopelta darlingi]]''<ref name=PeerJSpalangiopelta>{{Cite journal|last1=Moser |first1=M. |last2=Burks |first2=R. A. |last3=Ulmer |first3=J. M. |last4=Heraty |first4=J. M. |last5=van de Kamp |first5=T. |last6=Krogmann |first6=L. |title=Taxonomic description and phylogenetic placement of two new species of ''Spalangiopelta'' (Hymenoptera: Pteromalidae: Ceinae) from Eocene Baltic amber |year=2021 |journal=PeerJ |volume=9 |pages=e10939 |doi=10.7717/peerj.10939 |pmid=34113480 |pmc=8162234 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Moser ''in'' Moser ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Pteromalidae]].
|
|-
|
''[[Spalangiopelta semialba]]''<ref name=PeerJSpalangiopelta />
|
Sp. nov
|
Valid
|
Moser ''in'' Moser ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family Pteromalidae.
|
|-
|
''[[Spheciellus]]''<ref name=Allommation />
|
Fam. nov., Gen. et sp. nov.
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Apoidea in the new family [[Spheciellidae]]. Genus includes new species ''S. aenigmaticus''.
|
|-
|
''[[Stephanorhyssalus]]''<ref name=Megalyrhyssalus />
|
Gen. et sp. nov
|
Valid
|
Belokobylskij & Jouault
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Braconidae. Genus includes new species ''S. longiscapus''.
|
|-
|
''[[Tetragonula florilega]]''<ref name=JM105 />
|
Sp. nov
|
Valid
|
Engel ''in'' Engel ''et al.''
|
Miocene (Langhian)
|
Zhangpu amber
|
{{Flag|China}}
|
A stingless bee, a species of ''[[Tetragonula]]''.
|
|-
|
''[[Thigmocosmus]]''<ref name=Allommation />
|
Gen. et sp. nov
|
Valid
|
Rosa & Melo
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Heterogynaidae. Genus includes new species ''T. longicornis''.
|
|-
|
''[[Tyrannomecia]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Nel |first2=A. |title=The oldest Cenozoic ant fossil: †''Tyrannomecia'' gen. nov. (Formicidae: Myrmeciinae) from the Palaeocene Menat Formation (France) |journal=Historical Biology |year=2022 |volume=34 |issue=11 |pages=2241–2248 |doi=10.1080/08912963.2021.2010192 |s2cid=245194843 |url=https://hal-insu.archives-ouvertes.fr/insu-03477166/file/Menat-myrme.pdf }}</ref>
|
Gen. et sp. nov
|
In press
|
Jouault & Nel
|
[[Paleocene]]
|
[[Menat Formation]]
|
{{Flag|France}}
|
An ant belonging to the [[Crown group#Stem groups|stem group]] of the subfamily [[Myrmeciinae]]. Genus includes new species ''T. inopinata''.
|
|-
|
''[[Vespula? hassiaca]]''<ref>{{Cite journal|last1=Abels |first1=J. |last2=Wedmann |first2=S. |title=A fossil wasp (Hymenoptera: Vespidae: ''Vespula? hassiaca'') from the Eocene of Messel |year=2021 |journal=Palaeontographica Abteilung A |volume=323 |issue=1–3 |pages=105–117 |doi=10.1127/pala/2021/0122 |s2cid=245248645 }}</ref>
|
Sp. nov
|
In press
|
Abels & Wedmann
|
Eocene
|
[[Messel pit]]
|
{{Flag|Germany}}
|
A member of the family [[Vespidae]].
|
|-
|
''[[Xeris dorbnikensis]]''<ref>{{Cite journal|last1=Manukyan |first1=A.R. |last2=Smirnova |first2=A.V. |title=New data on the family Siricidae (Hymenoptera, Symphyta) in Baltic amber |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=157–165 |doi=10.1134/S003103012102009X |issn=0031-0301  |s2cid=233746801 |url=https://elibrary.ru/item.asp?id=44595943 }}</ref>
|
Sp. nov
|
Valid
|
Manukyan ''in'' Manukyan & Smirnova
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[horntail]]. Originally described as a species of ''[[Xeris]]''; Archibald & [[Alexandr Rasnitsyn|Rasnitsyn]] (2022) transferred it to the genus ''[[Urocerus]]''.<ref>{{Cite journal |last1=Archibald |first1=S. B. |last2=Rasnitsyn |first2=A. P. |title=The early Eocene ''Eourocerus anguliterreus'' gen. et sp. nov (Hymenoptera, Siricidae) from Republic, Washington |year=2022 |journal=Zootaxa |volume=5105 |issue=2 |pages=289–295 |doi=10.11646/zootaxa.5105.2.8 |pmid=35391300 |s2cid=247240762 }}</ref>
|
|-
|
''[[Zigrasimecia|Zigrasimecia goldingot]]''<ref>{{Cite journal|last1=Zhuang |first1=Y. |last2=Ran |first2=H. |last3=Li |first3=X. |last4=Feng |first4=C. |last5=Liu |first5=Y. |title=A new species of the iron maiden ant based on an alate female from mid-Cretaceous Burmese amber (Hymenoptera: Formicidae: †''Zigrasimecia'') |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105056 |doi=10.1016/j.cretres.2021.105056 |s2cid=240812443 }}</ref>
|
Sp. nov
|
In press
|
Zhuang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
An ant in the tribe [[Sphecomyrmini]].
|
|-
|}

===Hymenopteran Research===
* Guo ''et al.'' (2021) report the discovery of two [[queen ant]] specimens from Burmese amber, belonging to the species ''[[Haidomyrmex]] cerberus'' and including the first known [[alate]] queen in ''Haidomyrmex''.<ref>{{Cite journal|last1=Guo |first1=Y. |last2=Shih |first2=C. |last3=Zhuo |first3=D. |last4=Ren |first4=D. |last5=Zhao |first5=Y. |last6=Gao |first6=T. |title=The first queen-worker association for Cretaceous Formicidae: the winged caste of ''Haidomyrmex cerberus'' |year=2021 |journal=ZooKeys |issue=1048 |pages=69–78 |doi=10.3897/zookeys.1048.66920 |pmid=34305419 |pmc=8280096 |doi-access=free }}</ref>
* Cooccurrence of ants belonging to extant [[Holarctic]] genus ''[[Lasius]]'' and a tropical [[weaver ant]] in the same piece of amber is reported from the [[Bitterfeld amber]] ([[Germany]]) by Radchenko & Perkovsky (2021).<ref>{{Cite journal|last1=Radchenko |first1=A. G. |last2=Perkovsky |first2=E. E. |title=Wheeler's dilemma revisited: first ''Oecophylla''–''Lasius'' syninclusion and other ants syninclusions in the Bitterfeld amber (late Eocene) |year=2021 |journal=Invertebrate Zoology |volume=18 |issue=1 |pages=47–65 |doi=10.15298/invertzool.18.1.05 |s2cid=234800986 |doi-access=free }}</ref>
* Original claims that both ''[[Melittosphex burmensis]]'' and ''[[Discoscapa apicula]]'' were primitive [[bee]]s have been rejected by Rosa & Melo, who instead consider the former to be an [[Aculeata|aculeate wasp]] of uncertain affinity, and the latter to belong within the [[Crabronidae|crabronid wasp]] subfamily [[Crabroninae]], a lineage not directly related to bees.<ref name=Allommation />

==Clade Neuropterida==
===Neuropterans===
====New Neuropteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Ankyloleon]]''<ref>{{Cite journal|last1=Badano |first1=D. |last2=Fratini |first2=M. |last3=Maugeri |first3=L. |last4=Palermo |first4=F. |last5=Pieroni |first5=N. |last6=Cedola |first6=A. |last7=Haug |first7=J. T. |last8=Weiterschan |first8=T. |last9=Velten |first9=J. |last10=Mei |first10=M. |last11=Di Giulio |first11=A. |last12=Cerretti |first12=P. |title=X-ray microtomography and phylogenomics provide insights into the morphology and evolution of an enigmatic Mesozoic insect larva |year=2021 |journal=Systematic Entomology |volume=46 |issue=3 |pages=672–684 |doi=10.1111/syen.12482 |s2cid=235363035 |doi-access=free |hdl=11573/1552955 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Badano, Haug & Cerretti ''in'' Badano ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
An early representative of the clade [[Myrmeleontiformia]]. The type species is ''A. caudatus''.
|
|-
|
''[[Astioberotha|Astioberotha coutreti]]''<ref>{{Cite journal|last=Jouault |first=C. |title=A new species of thorny lacewing (Neuroptera: Rhachiberothidae: Paraberothinae) from mid-Cretaceous Burmese amber with novel raptorial foreleg structure |year=2021 |journal=Proceedings of the Geologists' Association |volume=133 |pages=32–39 |doi=10.1016/j.pgeola.2021.11.001 |s2cid=244457471 |url=https://hal-insu.archives-ouvertes.fr/insu-03445490 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Jouault
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Rhachiberothidae]] belonging to the subfamily [[Paraberothinae]].
|
|-
|
''[[Burmotachinymphes]]''<ref>{{Cite journal|last1=Cao |first1=L.-R. |last2=Wang |first2=B. |last3=Liu |first3=X.-Y. |title=A new genus and species of Mesochrysopidae (Neuroptera) from the mid-Cretaceous Burmese amber |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=77–84 |doi=10.11646/palaeoentomology.4.1.12 |s2cid=233963511 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cao, Wang & Liu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Mesochrysopidae]]. Genus includes new species ''M. bilobata''.
|
|-
|
''[[Calobabinskaia]]''<ref name=Calobabinskaia>{{Cite journal|last1=Lu |first1=X. |last2=Wang |first2=B. |last3=Liu |first3=X. |title=New Cretaceous antlion-like lacewings promote a phylogenetic reappraisal of the extinct myrmeleontoid family Babinskaiidae |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 16431 |year=2021 |doi=10.1038/s41598-021-95946-z |pmid=34385561 |pmc=8361207 |bibcode=2021NatSR..1116431L |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Lu, Wang & Liu
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Babinskaiidae]]. The type species is ''C. xiai''.
|
|-
|
''[[Fuscopolystoechotes]]''<ref name=Fuscopolystoechotes>{{Cite journal|last1=Xu |first1=Y. |last2=Zheng |first2=B. |last3=Fang |first3=H. |last4=Shih |first4=C. |last5=Ren |first5=D. |last6=Wang |first6=Y. |year=2021 |title=New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae) |journal=Zootaxa |volume=5067 |issue=2 |pages=259–266 |doi=10.11646/zootaxa.5067.2.7 | issn=1175-5326  |pmid=34810745 |s2cid=243981058 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Xu, Zheng, Shih & Wang ''in'' Xu ''et al.''
|
Middle Jurassic
|
Jiulongshan Formation
|
{{Flag|China}}
|
A member of the family [[Ithonidae]]. Genus includes new species ''F. reticulatus''.
|
|-
|
''[[Karenina cuneiformis]]''<ref>{{Cite journal|last=Nakamine |first=H. |title=A new species of Mesochrysopidae (Neuroptera) from the Lower Cretaceous Crato Formation of Brazil |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=115–119 |doi=10.18476/pale.v14.a4 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Nakamine
|
[[Early Cretaceous]] ([[Aptian]])
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A member of the family [[Mesochrysopidae]].
|
|-
|
''[[Lichenipolystoechotes|Lichenipolystoechotes fenestratus]]''<ref name=Fuscopolystoechotes />
|
Sp. nov
|
Valid
|
Xu, Fang, Shih & Wang ''in'' Xu ''et al.''
|
Middle Jurassic
|
Jiulongshan Formation
|
{{Flag|China}}
|
A member of the family Ithonidae.
|
|-
|
''[[Osmylidia donnae]]''<ref name="Protosmylinae2021">{{cite journal |last1=Makarkin |first1=V.N. |last2=Archibald |first2=S.B. |last3=Mathewes |first3=R.W. |year=2021 |title=New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks |journal=Zootaxa |volume=4980 |issue=1 |pages=142–156 |doi=10.11646/zootaxa.4980.1.9 |pmid=34186986|s2cid=235685548 }}</ref>
|
Sp. nov
|
Valid
|
Makarkin, Archibald, & Mathewes
|
[[Ypresian]]
|
[[Coldwater Beds|Quilchena site]]
|
{{Flag|Canada}}<br/>{{Flag|British Columbia}}
|
A [[Protosmylinae|protosmyline]] [[Osmylidae|Osmylid lance lacewing]] 
|
|-
|
''[[Osmylidia glastrai]]''<ref name="Protosmylinae2021"/>
|
Sp. nov
|
Valid
|
Makarkin, Archibald, & Mathewes
|
[[Ypresian]]
|
[[Klondike Mountain Formation]]
|
{{Flag|United States}}<br/>{{Flag|Washington}}
|
A [[Protosmylinae|protosmyline]] [[Osmylidae|Osmylid lance lacewing]] 
|
|-
|
''[[Osmylidia picta]]''<ref name="Protosmylinae2021"/>
|
Comb. nov
|
Valid
|
(Hagen)
|
[[Priabonian]]
|
[[Baltic amber]]
|
{{Flag|Europe}}
|
A [[Protosmylinae|protosmyline]] [[Osmylidae|Osmylid lance lacewing]]<br/> Moved from ''"[[Protosmylus]]" picta'' ([[1913 in paleontology|1913]])
|
|-
|
''[[Osmylidia taliae]]''<ref name="Protosmylinae2021"/>
|
Sp. nov
|
Valid
|
Makarkin, Archibald, & Mathewes
|
[[Ypresian]]
|
[[Green River Formation]]<br/> Parachute Creek Member
|
{{Flag|United States}}<br/>{{Flag|Colorado}}
|
A [[Protosmylinae|protosmyline]] [[Osmylidae|Osmylid lance lacewing]] 
|
|-
|
''[[Osmyloberotha]]''<ref>{{Cite journal|last=Khramov |first=A. V. |year=2021 |title=''Osmyloberotha'', an unusual new genus of beaded lacewings (Neuroptera: Berothidae) from Burmese amber |journal=Zootaxa |volume=5060 |issue=2 |pages=95–99 |doi=10.11646/zootaxa.5060.2.5 |pmid=34811173 |s2cid=240194221 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Khramov
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Berothidae|beaded lacewing]]. Genus includes new species ''O. simpla''.
|
|-
|
''[[Paradoxoberotha]]''<ref>{{Cite journal|last1=Nakamine |first1=H. |last2=Yamamoto |first2=S. |last3=Takahashi |first3=Y. |last4=Li |first4=H. |last5=Liu |first5=X. |title=A remarkable new thorny lacewing from mid-Cretaceous amber from northern Myanmar (Neuroptera: Rhachiberothidae) |year=2021 |journal=PalZ |volume=96 |issue=2 |pages=207–217 |doi=10.1007/s12542-021-00589-0 |s2cid=244780786 }}</ref>
|
Gen. et sp. nov
|
In press
|
Nakamine ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Rhachiberothidae]]. Genus includes new species ''P. chimaera''.
|
|-
|
''[[Paraneliana]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Nel |first2=A. |title=A new myrmeleontoid genus and species from the Lower Cretaceous Crato Formation (Neuroptera, Babinskaiidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=204–208 |doi=10.11646/palaeoentomology.4.3.4 |s2cid=237819174 |url=https://hal-insu.archives-ouvertes.fr/insu-03272782/file/Jouault_Nel_Paraneliana.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jouault & Nel
|
Early Cretaceous
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A member of the family [[Babinskaiidae]]. Genus includes new species ''P. sennlaubi''.
|
|-
|
''[[Rasnitsyneura]]''<ref>{{Cite journal|last1=Makarkin |first1=V. N. |last2=Ansorge |first2=J. |last3=Khramov |first3=A. V. |title=Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae ''s.l.'' (Neuroptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=516–531 |doi=10.11646/palaeoentomology.4.6.1 |s2cid=245181387 }}</ref>
|
Gen. et comb. nov
|
Valid
|
Makarkin, Ansorge & Khramov
|
[[Early Jurassic]] ([[Toarcian]])
|
|
{{Flag|Germany}}
|
A member of the family [[Ithonidae]] ''sensu lato'' belonging to the subfamily [[Epigambriinae]]. The type species is ''"[[Prohemerobius]]" aequabilis'' Bode (1953).
|
|-
|
''[[Sinonothochrysa]]''<ref>{{Cite journal|last1=Huang |first1=D. |last2=Lian |first2=X. |last3=Jouault |first3=C. |last4=Gao |first4=J. |last5=Nel |first5=A. |year=2021 |title=First green lacewing (Insecta: Neuroptera: Chrysopidae) from the Palaeocene Sanshui basin of Guangdong, South China |journal=Historical Biology |volume=34 |issue=4 |pages=704–712 |doi=10.1080/08912963.2021.1943381 |s2cid=237756011 |url=https://hal-insu.archives-ouvertes.fr/insu-03272764/file/Huang_et_al_Chrysopidae.pdf }}</ref>
|
Gen. et sp. nov
|
In press
|
Huang & Nel ''in'' Huang ''et al.''
|
[[Paleocene]]
|
[[Buxin Formation]]
|
{{Flag|China}}
|
A member of the family [[Chrysopidae]]. Genus includes new species ''S. zhangi''
|
|-
|
''[[Stenobabinskaia]]''<ref name=Calobabinskaia />
|
Gen. et sp. nov
|
Valid
|
Lu, Wang & Liu
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Babinskaiidae. The type species is ''S. punctata''.
|
|-
|
''[[Xiaobabinskaia]]''<ref name=Calobabinskaia />
|
Gen. et sp. nov
|
Valid
|
Lu, Wang & Liu
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Babinskaiidae. The type species is ''X. lepidotricha''.
|
|-
|}

====Neuropteran research====
* Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.<ref>{{Cite journal|last1=Haug |first1=G. T. |last2=Haug |first2=C. |last3=Haug |first3=J. T. |title=The morphological diversity of spoon-winged lacewing larvae and the first possible fossils from 99 million-year-old Kachin amber, Myanmar |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=133–152 |doi=10.18476/pale.v14.a6 |doi-access=free }}</ref>
* New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug ''et al.'' (2021), who also review the morphological diversity of fossil lacewing larvae.<ref>{{Cite journal|last1=Haug |first1=J. T. |last2=Baranov |first2=V. |last3=Müller |first3=P. |last4=Haug |first4=C. |title=New extreme morphologies as exemplified by 100 million-year-old lacewing larvae |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 20432 |doi=10.1038/s41598-021-99480-w |pmid=34650142 |pmc=8517013 |doi-access=free }}</ref>

===Raphidiopterans===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Archiinocellia protomaculata]]''<ref name="Archibald2021">{{Cite journal|last1=Archibald |first1=S. B. |last2=Makarkin |first2=V. N. |title=Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation |year=2021 |journal=Zootaxa |volume=4951 |issue=1 |pages=41–79 |doi=10.11646/zootaxa.4951.1.2 |pmid=33903413 |s2cid=233411745 }}</ref>
|
comb nov
|
valid
|
([[Michael S. Engel|Engel]])
|
[[Lutetian]]
| 
[[Green River Formation]]
|
{{Flag|USA}}<br/>{{flag|Colorado}}
|
A [[Raphidiidae|raphidiid]] [[snakefly]]. <br/>Moved from ''[[Agulla protomaculata]]'' ([[2011 in arthropod paleontology|2011]])<ref name="Engel2011a">{{Cite journal|last1=Engel |first1=Michael S. |year=2011 |title=A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae)|journal= Transactions of the Kansas Academy of Science |volume=114 |issue= 1–2 |pages = 77–87 |doi=10.1660/062.114.0107 |s2cid=85011364 }}</ref>
|
[[file:Archiinocellia protomaculata holotype NMNH-USNM PAL 569865 img1.tif|thumb|center|upright|''[[Archiinocellia protomaculata]]'']]  
|-
|
''[[Electrobaissoptera|Electrobaissoptera liui]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Engel |first2=M. S. |last3=Nel |first3=A. |title=A new baissopterid snakefly (Raphidioptera: Baissopteridae) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105028 |doi=10.1016/j.cretres.2021.105028 |s2cid=239253683 }}</ref>
|
Sp. nov
|
In press
|
Jouault, [[Michael S. Engel|Engel]] & Nel
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Baissopteridae|baissopterid]] snakefly.
|
|-
|
''[[Megaraphidia|Megaraphidia antiquissima]]''<ref name="Archibald2021"/>
|
Sp. nov
|
Valid
|
Archibald & Makarkin
|
[[Ypresian]]
|
[[McAbee Fossil Beds|Tranquille Formation]]
|
{{Flag|Canada}}<br/>{{Flag|British Columbia}}
|
A [[Raphidiidae|raphidiid]] [[snakefly]]. 
|
|-
|
''[[Megaraphidia|Megaraphidia hopkinsi]]''<ref name="Archibald2021"/>
|
Sp. nov
|
Valid
|
Archibald & Makarkin
|
[[Ypresian]]
|
[[Allenby Formation]]
|
{{Flag|Canada}}<br/>{{Flag|British Columbia}}
|
A [[Raphidiidae|raphidiid]] [[snakefly]]. 
|
|-
|
''[[Megaraphidia|Megaraphidia klondika]]''<ref name="Archibald2021"/>
|
Sp. nov
|
Valid
|
Archibald & Makarkin
|
[[Ypresian]]
|
[[Klondike Mountain Formation]]
|
{{Flag|United States}}<br/>{{Flag|Washington}}
|
A [[Raphidiidae|raphidiid]] [[snakefly]]. 
|
|-
|
''[[Megaraphidia|Megaraphidia ootsa]]''<ref name="Archibald2021"/>
|
Sp. nov
|
Valid
|
Archibald & Makarkin
|
[[Ypresian]]
|
[[Driftwood Canyon Provincial Park|"Unnamed unit" Ootsa Group]]
|
{{Flag|Canada}}<br/>{{Flag|British Columbia}}
|
A [[Raphidiidae|raphidiid]] [[snakefly]]. 
|
|-
|
''[[Mesoraphidia|Mesoraphidia baekthoensis]]''<ref name=CRMesoraphidia>{{Cite journal|last1=Ri |first1=C.-J. |last2=Won |first2=C.-G. |last3=So |first3=K.-S. |last4=Ho |first4=M.-S. |title=New snakeflies (Insecta: Raphidioptera: Mesoraphidiidae) from the Lower Cretaceous Sinuiju Formation of the Democratic People's Republic of Korea |year=2021 |journal=Cretaceous Research |volume=132 |pages=Article 105105 |doi=10.1016/j.cretres.2021.105105 |s2cid=244700502 }}</ref>
|
Sp. nov
|
In press
|
Ri ''et al.''
|
Early Cretaceous (Barremian–Aptian)
|
Sinuiju Formation
|
{{Flag|North Korea}}
|
A member of the family [[Mesoraphidiidae]].
|
|-
|
''Mesoraphidia ryonsangensis''<ref name=CRMesoraphidia />
|
Sp. nov
|
In press
|
Ri ''et al.''
|
Early Cretaceous (Barremian–Aptian)
|
Sinuiju Formation
|
{{Flag|North Korea}}
|
A member of the family Mesoraphidiidae.
|
|-
|}

==Clade †Palaeodictyopteroidea==
===†Megasecoptera===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Anhuihymen]]''<ref>{{Cite journal|last1=Huang |first1=D. |last2=Fu |first2=Y. |last3=Lian |first3=X. |last4=Gao |first4=J. |last5=Nel |first5=A. |year=2021 |title=The first Chinese Protohymenidae (Palaeoptera: Megasecoptera) |journal=Historical Biology |volume=34 |issue=3 |pages=458–461 |doi=10.1080/08912963.2021.1929950 |s2cid=236405039 |url=https://hal.sorbonne-universite.fr/hal-03238506 }}</ref>
|
Gen. et sp. nov
|
In press
|
Huang & Nel ''in'' Huang ''et al.''
|
Middle [[Permian]]
|
[[Yinping Formation]]
|
{{Flag|China}}
|
A [[Protohymenidae|protohymenid]] [[Megasecoptera]].<br/> The type species is ''A. medianelongata''.
|
|-
|}

===†Palaeodictyoptera===
====New Palaeodictyopteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Dictyoneura goujonorum]]''<ref>{{Cite journal|last1=Logghe |first1=A. |last2=Garrouste |first2=R. |last3=Steyer |first3=J.-S. |last4=Pouillon |first4=J.-M. |last5=Nel |first5=A. |year=2021 |title= The first Palaeodictyoptera (Insecta) from the Carboniferous-Permian basin of Graissessac (France) |journal=Historical Biology |volume=34 |issue=3 |pages=486–492 |doi=10.1080/08912963.2021.1929953 |s2cid=236354115 |url=https://hal.sorbonne-universite.fr/hal-03257982 }}</ref>
|
Sp. nov
|
In press
|
Logghe ''et al.''
|
[[Carboniferous]]-Permian ([[Gzhelian]]-[[Asselian]])
|
|
{{Flag|France}}
|
A member of [[Palaeodictyoptera]].
|
|-
|
''[[Sandiella|Sandiella herbigi]]''<ref>{{Cite journal|last1=Brauckmann |first1=C. |last2=Herd |first2=K. J. |last3=Leipner |first3=A. |title=First Eugereonidae (Insecta: Palaeodictyoptera) from the Pennsylvanian (Late Carboniferous) of the Piesberg site near Osnabrück, Germany |journal=PalZ |date=2023 |issue=4 |pages=861–866 |volume=97 |doi=10.1007/s12542-021-00578-3 |s2cid=244745023 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Brauckmann, Herd & Leipner
|
Carboniferous ([[Moscovian (Carboniferous)|Moscovian]])
|
|
{{Flag|Germany}}
|
A member of Palaeodictyoptera belonging to the family [[Eugereonidae]].
|
|-
|}
====Palaeodictyopteran research====
* A study on the anatomy of fossil specimens described as consecutive series of different larval stages of [[palaeodictyoptera]]n species ''[[Tchirkovaea]] guttata'' and ''[[Paimbia]] fenestrata'', evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).<ref>{{Cite journal|last1=Rosová |first1=K. |last2=Sinitshenkova |first2=N. D. |last3=Prokop |first3=J. |title=Evidence for wing development in the Late Palaeozoic Palaeodictyoptera revisited |year=2021 |journal=Arthropod Structure & Development |volume=63 |pages=Article 101061 |doi=10.1016/j.asd.2021.101061 |pmid=34098321 |s2cid=235369956 }}</ref>

==Clade Palaeoptera==
===Ephemeroptera===
====New Ephemeropteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Astraeoptera]]''<ref>{{Cite journal|last1=Brandão |first1=N. C. A. |last2=Bittencourt |first2=J. S. |last3=Calor |first3=A. R. |last4=Mendes |first4=M. |last5=Langer |first5=M. C. |title=The Ephemeroptera (Hexapoda, Insecta) from the Lower Cretaceous Crato Formation (NE Brazil): a new genus and species, and reassessment of ''Costalimella zucchii'' Zamboni, 2001 and ''Cratogenites corradiniae'' Martins-Neto, 1996 |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104923 |doi=10.1016/j.cretres.2021.104923 |bibcode=2021CrRes.12704923B |url=http://www.repositorio.ufc.br/handle/riufc/64788 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Brandão ''et al.''
|
Early Cretaceous
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A mayfly of uncertain phylogenetic placement.<br/> Genus includes new species ''A. cretacica''.
|
|-
|
''[[Burmella]]''<ref>{{Cite journal|last1=Godunko |first1=R. J. |last2=Martynov |first2=A. V. |last3=Staniczek |first3=A. H. |title=First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution |year=2021 |journal=ZooKeys |issue=1036 |pages=99–120 |doi=10.3897/zookeys.1036.66435 |doi-access=free |pmid=34040482 |pmc=8128840 }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Godunko, Martynov & Staniczek
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Vietnamellidae|vietnamellid]] [[mayfly]].<br/> The type species is ''B. paucivenosa''; genus also includes ''B. clypeata''.
|
|-
|
''[[Protoligoneuria|Protoligoneuria heloisae]]''<ref>{{Cite journal|last1=Storari |first1=A. P. |last2=Godunko |first2=R. J. |last3=Salles |first3=F. F. |last4=Saraiva |first4=A. A. F. |last5=Staniczek |first5=A. H. |last6=Rodrigues |first6=T. |year=2021 |title=An overview of the Hexagenitidae (Ephemeroptera) from the Crato Formation (Aptian, Lower Cretaceous) of Brazil, with the description of a new species |journal=Historical Biology |volume=34 |issue=5 |pages=875–884 |doi=10.1080/08912963.2021.1952196 |s2cid=237727305 }}</ref>
|
Sp. nov
|
In press
|
Storari ''et al.''
|
Early Cretaceous ([[Aptian]])
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A [[Hexagenitidae|hexagenitid]] mayfly.
|
|-
|
''[[Triassodotes|Triassodotes rasnitsyni]]''<ref>{{Cite journal|last=Sinitshenkova |first=N. D. |title=A new mayfly species ''Triassodotes rasnitsyni'' sp. nov. of the family Misthodotidae Tillyard, 1932 (Insecta; Ephemerida, Permoplectoptera) from the Triassic deposits of Kuzbass, Russia |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=429–432 |doi=10.11646/palaeoentomology.4.5.6 |s2cid=244195004 }}</ref>
|
Sp. nov
|
Valid
|
Sinitshenkova
|
Triassic
|
|
{{Flag|Russia}}
|
A [[Misthodotidae|misthodotid]] mayfly.
|
|-
|}

====Ephemeropteran research====
* Sroka ''et al.'' (2021) provide new information on the morphology of Permian stem-mayflies ''[[Misthodotes]] sharovi'' and ''M. zalesskyi'' from the Tshekarda locality ([[Kungurian]] Koshelevka Formation; [[Perm Krai]], [[Russia]]), and infer the life history traits of both the adult and larval stages of these insects.<ref>{{Cite journal|last1=Sroka |first1=P. |last2=Godunko |first2=R. J. |last3=Sinitshenkova |first3=N. D. |last4=Prokop |first4=J. |year=2021 |title=Life history, systematics and flight ability of the Early Permian stem-mayflies in the genus ''Misthodotes'' Sellards, 1909 (Insecta, Ephemerida, Permoplectoptera) |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 97 |doi=10.1186/s12862-021-01820-x |pmid=34024284 |pmc=8142488 |doi-access=free }}</ref>
* Possible underwater mayfly burrows are described from the [[Turonian]] [[Ferron Formation|Ferron Sandstone]] ([[Utah]], United States) by King ''et al.'' (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new [[Ichnotaxon|ichnospecies]] ''[[Glossifungites]] gingrasi''.<ref>{{Cite journal|last1=King |first1=M. R. |last2=La Croix |first2=A. D. |last3=Gates |first3=T. A. |last4=Anderson |first4=P. B. |last5=Zanno |first5=L. E. |title=''Glossifungites gingrasi'' n. isp., a probable subaqueous insect domicile from the Cretaceous Ferron Sandstone, Utah |year=2021 |journal=Journal of Paleontology |volume=95 |issue=3 |pages=427–439 |doi=10.1017/jpa.2020.115 |bibcode=2021JPal...95..427K |doi-access=free }}</ref>

===Odonatoptera===
====Odonata====
=====New Odonatan taxa=====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Chalcolestes tibetensis]]''<ref>{{Cite journal|last1=Xia |first1=G. |last2=Zheng |first2=D. |last3=Krieg-Jacquier |first3=R. |last4=Fan |first4=Q. |last5=Chen |first5=Y. |last6=Nel |first6=A. |title=The oldest-known Lestidae (Odonata) from the late Eocene of Tibet: palaeoclimatic implications |year=2021 |journal=Geological Magazine |volume=159 |issue=4 |pages=511–518 |doi=10.1017/S0016756821001102 |s2cid=242028737 }}</ref>
|
Sp. nov
|
In press
|
Xia ''et al.''
|
Late Eocene
|
Nima Basin
|
{{Flag|China}}
|
A species of ''[[Chalcolestes]]''.
|
|-
|
''[[Cretaphylolestes]]''<ref>{{Cite journal|last1=Huang |first1=D. |last2=Fu |first2=Y. |last3=Lian |first3=X. |last4=Gao |first4=J. |last5=Nel |first5=A. |title=The oldest malachite damselfly (Odonata: Synlestidae) from the Lower Cretaceous of China |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105023 |doi=10.1016/j.cretres.2021.105023 |s2cid=239681781 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
In press
|
Huang ''et al.''
|
Early Cretaceous ([[Aptian]])
|
[[Shouchang Formation]]
|
{{Flag|China}}
|
A damselfly belonging to the family [[Synlestidae]]. Genus includes new species ''C. cretacicus''.
|
|-
|
''[[Dakotaeschnidium]]''<ref>{{Cite journal|last=Nel |first=A. |title=Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=209–212 |doi=10.11646/palaeoentomology.4.3.5 |s2cid=237876331 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Nel
|
Late Cretaceous ([[Maastrichtian]])
|
[[Fox Hills Formation]]
|
{{Flag|United States}}<br/>({{flag|South Dakota}})
|
A dragonfly belonging to the family [[Aeschnidiidae]]. Genus includes new species ''D. spedeni''.
|
|-
|
''[[Dysagrion pruettae]]''<ref name=Cephalozygoptera2021>{{Cite journal|last1=Archibald |first1=S. B. |last2=Cannings |first2=R. A. |last3=Erickson |first3=R. J. |last4=Bybee |first4=S. M. |last5=Mathewes |first5=R. W. |title=The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America |year=2021 |journal=Zootaxa |volume=4934 |issue=1 |pages=zootaxa.4934.1.1 |doi=10.11646/zootaxa.4934.1.1 |pmid=33756770 |s2cid=232337536 }}</ref>
|
Sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
[[Eocene]] ([[Ypresian]])
|
[[Klondike Mountain Formation]]
|
{{Flag|United States}}<br/>({{flag|Washington}})
|
A member of the family [[Dysagrionidae]].
|
[[File:Dysagrion pruettae holotype SRIC SR 13-005-012 A.tif|thumb|center|upright|''[[Dysagrion pruettae]]'']]
|-
|
''[[Dysagrionites]]''<ref name=Cephalozygoptera2021 />
|
Gen. et sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
Eocene (Ypresian)
|
Klondike Mountain Formation
|
{{Flag|United States}}<br/>({{flag|Washington}})
|
A collective group name for any fossil species referable to the [[Dysagrioninae]] whose [[orthotaxon]] generic affinity is unclear. Includes new species ''D. delinei''.
|
|-
|
''[[Epilestes|Epilestes rasnitsyni]]''<ref>{{Cite journal|last1=Felker |first1=A. |last2=Vasilenko |first2=D. |title=A new species of the {{okina}}protozygopteranʼ damselfly (Odonata: Permagrionidae) from the Lower-Middle Permian of Russia |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=462–467 |doi=10.11646/palaeoentomology.4.5.12 |s2cid=240547006 }}</ref>
|
Sp. nov
|
Valid
|
Felker & Vasilenko
|
[[Permian]]
|
|
{{Flag|Russia}}
|
A damselfly belonging to the family [[Permagrionidae]].
|
|-
|
''[[Jingguaeshna]]''<ref>{{Cite journal|last1=Zheng |first1=D. |last2=Li |first2=S. |last3=Zhao |first3=Y. |last4=Zhang |first4=H. |title=A potential telephlebiid dragonfly (Odonata: Anisoptera: Aeshnoidea) from Miocene of Yunnan, southwestern China |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=227–242 |doi=10.11646/palaeoentomology.4.3.11 |s2cid=237825025 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Zheng ''et al.''
|
Miocene
|
|
{{Flag|China}}
|
A dragonfly belonging to the group [[Aeshnoidea]], possibly a member of the family [[Telephlebiidae]]. Genus includes new species ''J. taoae''.
|
|-
|
''[[Marado marado|Marado]]''<ref>{{cite journal |last=Petrulevičius |first=J. F. |year=2021 |title=First Coenagrionid Damselfly (Odonata: Zygoptera) from the Late Palaeocene of Northwestern Argentina |journal=Life: The Excitement of Biology |volume=9 |issue=1 |pages=6–15 |url=https://blaypublishers.files.wordpress.com/2021/12/petrulevicius_leb9131dec.21.pdf }}</ref><ref>{{Cite journal|last=Nel |first=A. |title=Revision of the damselfly ''Hesperagrion praevolans'' (Odonata, Zygoptera, Coenagrionidae) from the uppermost Eocene of Florissant (Colorado, USA) |year=2022 |journal=Palaeoentomology |volume=5 |issue=6 |pages=517–519 |doi=10.11646/palaeoentomology.5.6.2 |s2cid=255089799 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Petrulevičius
|
[[Paleocene]]
|
Maíz Gordo Formation
|
{{Flag|Argentina}}
|
A [[damselfly]] belonging to the family [[Coenagrionidae]]. The type species is ''M. marado''.
|
|-
|
''[[Moltenophlebia]]''<ref>{{Cite journal|last1=Deregnaucourt |first1=I. |last2=Bardin |first2=J. |last3=Anderson |first3=J. M. |last4=Béthoux |first4=O. |title=The wing venation of a new fossil species, reconstructed using geometric morphometrics, adds to the rare fossil record of Triassic Gondwanian Odonata |year=2021 |journal=Arthropod Structure & Development |volume=63 |pages=Article 101056 |doi=10.1016/j.asd.2021.101056 |pmid=33984598 |s2cid=234494778 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Deregnaucourt ''et al.''
|
[[Triassic]]
|
[[Molteno Formation]]
|
{{Flag|South Africa}}
|
A member of the [[Crown group#Stem groups|stem group]] of [[Odonata]], belonging to the group [[Zygophlebiida]] within the [[Triadophlebiomorpha]]. Genus includes new species ''M. lindae''.
|
|-
|
''[[Okanagrion]]''<ref name=Cephalozygoptera2021 />
|
Gen. et 8 sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
Eocene (Ypresian)
|
Klondike Mountain Formation
|
{{Flag|United States}}<br/>({{flag|Washington}})
|
A member of the family Dysagrionidae. The type species is ''O. threadgillae''; genus also includes new species ''O. hobani'', ''O. beardi'', ''O. lochmum'', ''O. angustum'', ''O. dorrellae'', ''O. liquetoalatum'' and ''O. worleyae''.
|
[[File:Okanagrion threadgillae holotype SRIC SR 98-12-10.tiff|thumb|center|upright|''[[Okanagrion threadgillae]]'']]
|-
|
''[[Okanopteryx]]''<ref name=Cephalozygoptera2021 />
|
Gen. et 3 sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
Eocene (Ypresian)
|
Tranquille Formation
|
{{Flag|Canada}}<br>({{Flag|British Columbia}})
|
A member of the family Dysagrionidae. The type species is ''O. macabeensis''; genus also includes new species ''O. fraseri'' and ''O. jeppesenorum''.
|
|-
|
''[[Phyonganpodagrion]]''<ref>{{Cite journal|last1=So |first1=K. S. |last2=Won |first2=C. G. |title=The first 'Megapodagrionidae' (Odonata, Zygoptera) from the Lower Cretaceous of Democratic People's Republic of Korea |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105054 |doi=10.1016/j.cretres.2021.105054 |s2cid=244226045 }}</ref>
|
Gen. et sp. nov
|
In press
|
So & Won
|
Early Cretaceous ([[Barremian]]–[[Aptian]])
|
[[Sinuiju Formation]]
|
{{Flag|North Korea}}
|
A damselfly in the family [[Megapodagrionidae]]. Genus includes new species ''P. ryonsangae''.
|
|-
|
''[[Protohemiphlebia]]''<ref>{{cite book |last1=Zheng |first1=D. |last2=Jarzembowski |first2=E. A. |last3=Zhuo |first3=D. |last4=Nel |first4=A. |year=2021 |chapter=Protohemiphlebiidae fam. nov., a stem hemiphlebioid damselfly from Cretaceous amber in Kachin |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |volume=521 |pages=171–184 |publisher=The Geological Society of London |doi=10.1144/SP521-2020-249 |s2cid=244087958 }}</ref>
|
Fam. et gen. et 2 sp. nov
|
In press
|
Zheng, Jarzembowski & Nel ''in'' Zheng ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Crown group#Stem groups|stem]] [[Hemiphlebioidea|hemiphlebioid]] damselfly belonging to the new family [[Protohemiphlebiidae]]. Genus includes new species ''P. zhangi'' and ''P. meiyingae''.
|
|-
|
''[[Pulchrairina]]''<ref>{{Cite journal|last1=Martynov |first1=A. V. |last2=Vasilenko |first2=D. V. |last3=Perkovsky |first3=E. E. |title=First Odonata from Upper Eocene Rovno amber (Ukraine) |journal=Historical Biology |year=2022 |volume=34 |issue=11 |pages=2182–2187 |doi=10.1080/08912963.2021.2005040 |s2cid=244914618 }}</ref>
|
Gen. et sp. nov
|
In press
|
Martynov, Vasilenko & Perkovsky
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A damselfly belonging to the group [[Coenagrionoidea]].<br/> Genus includes new species ''P. electra''.
|
|-
|
''[[Republica (damselfly)|Republica]]''<ref>{{Cite journal|last1=Archibald |first1=S. B. |last2=Cannings |first2=R. A. |year=2021 |title=A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U.S.A. |journal=Zootaxa |volume=4966 |issue=3 |pages=392–400 |doi=10.11646/zootaxa.4966.3.11 |pmid=34186607 |s2cid=235557114 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Archibald & Cannings
|
Eocene (Ypresian)
|
Klondike Mountain Formation
|
{{Flag|United States}}<br/>({{flag|Washington}})
|
A [[Euphaeidae|euphaeid]] damselfly.<br/> Type species is ''R. weatbrooki''.<br/>Not to be confused with the plant genus ''[[Republica (plant)|Republica]]'' also from the formation.
|
[[file:Republica weatbrooki holotype SR 06-59-08 img1.tif|thumb|center|upright|''[[Republica weatbrooki]]'']]
|-
|
''[[Stenodiafanus]]''<ref name=Cephalozygoptera2021 />
|
Gen. et sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
Eocene (Ypresian)
|
Klondike Mountain Formation
|
{{Flag|United States}}<br/>({{flag|Washington}})
|
A dysagrionid odonate.<br/> Type species is ''S. westersidei''.
|
[[File:Stenodiafanus westersidei holotype SR 06-01-42 A (part) img1.tif|thumb|center|upright|''[[Stenodiafanus westersidei]]'']]
|-
|
''[[Stenophlebia|Stenophlebia ryonsangensis]]''<ref>{{Cite journal|last1=Won |first1=C.G. |last2=So |first2=K.S. |last3=Kim |first3=H.C. |title=A new odonatan (Stenophlebiidae) from the Lower Cretaceous Sinuiju Formation of North Phyongan Province, the Democratic People's Republic of Korea |year=2021 |journal=Palaeoworld |volume=31 |pages=116–120 |doi=10.1016/j.palwor.2021.02.002| issn=1871-174X  |s2cid=233959783 }}</ref>
|
Sp. nov
|
In press
|
Won, So & Kim
|
Early Cretaceous
|
[[Sinuiju Formation]]
|
{{Flag|North Korea}}
|
A [[Stenophlebiidae|stenophlebiid odonate]].
|
|-
|
''[[Whetwhetaksa]]''<ref name=Cephalozygoptera2021 />
|
Fam. Nov., Gen. et sp. nov
|
Valid
|
Archibald & Cannings ''in'' Archibald ''et al.''
|
Eocene (Ypresian)
|
Klondike Mountain Formation
|
{{Flag|United States}}
|
The type genus of [[Whetwhetaksidae]], a new family of uncertain placement.<br/> The type species is ''W. millerae''.
|
[[File:Whetwhetaksa millerae holotype SRIC SR 06-01-38 img1.tiff|thumb|center|upright|''[[Whetwhetaksa millerae]]'']]
|-
|}

=====Odonate research=====
* Archibald ''et al.'' (2021) establish a new, extinct suborder of Odonata named [[Cephalozygoptera]], close to but distinct from damselflies, composed of the families Dysagrionidae, [[Sieblosiidae]] and possibly Whetwhetaksidae;<ref name=Cephalozygoptera2021 /> their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald ''et al.'' to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.<ref>{{Cite journal|last1=Nel |first1=A. |last2=Zheng |first2=D. |title=The recently proposed odonatan 'suborder' Cephalozygoptera: fact or fiction |year=2021 |journal=Palaeoentomology |volume=4 |issue=2 |pages=165–170 |doi=10.11646/palaeoentomology.4.2.5 |s2cid=235536486 }}</ref><ref>{{Cite journal|last1=Archibald |first1=S. B. |last2=Cannings |first2=R. A. |title=The head of Cephalozygoptera (Odonata) |year=2021 |journal=Zootaxa |volume=5047 |issue=1 |pages=97–100 |doi=10.11646/zootaxa.5047.1.10 |pmid=34811004 |s2cid=244256746 |doi-access=free }}</ref>

====†Protozygoptera====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Kennedya kedrovkensis]]''<ref name=PJ552Kennedyidae>{{Cite journal|last=Felker |first=A. S. |title=The first damselflies of the family Kennedyidae (Odonata: Protozygoptera) from the Permian-Triassic boundary deposits of the Kuznetsk Basin, Russia |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=166–173 |doi=10.1134/S0031030121020052 |s2cid=233746783 |url=https://elibrary.ru/item.asp?id=44595944 }}</ref>
|
Sp. nov
|
Valid
|
Felker
|
[[Early Triassic]]
|
|
{{Flag|Russia}}
|
A member of [[Protozygoptera]] belonging to the family [[Kennedyidae]].
|
|-
|
''[[Kennedya suchonensis]]''<ref>{{Cite journal |last=Felker |first=A. S. |title=New damselflies of the family Kennedyidae (Odonata: Protozygoptera) from the Upper Permian of the Vologda Region |year=2021 |journal=Paleontological Journal |volume=55 |issue=4 |pages=396–404 |doi=10.1134/S0031030121040067 |s2cid=237375821 |url=https://elibrary.ru/item.asp?id=46227216 }}</ref>
|
Sp. nov
|
Valid
|
Felker
|
Late Permian
|
|
{{Flag|Russia}}
|
|
|-
|
''[[Progoneura kemerovensis]]''<ref name=PJ552Kennedyidae />
|
Sp. nov
|
Valid
|
Felker
|
Early Triassic
|
|
{{Flag|Russia}}
|
A member of Protozygoptera belonging to the family Kennedyidae.
|
|-
|}
====Other odonatopterans====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Brunellopteron]]''<ref>{{Cite journal|last1=Béthoux |first1=O. |last2=Norrad |first2=R. E. |last3=Stimson |first3=M. R. |last4=King |first4=O. A. |last5=Allen |first5=L. F. |last6=Deregnaucourt |first6=I. |last7=Hinds |first7=S. J. |last8=Lewis |first8=J. H. |last9=Schneider |first9=J. W. |title=A unique, large-sized stem Odonata (Insecta) found in the early Pennsylvanian of New Brunswick (Canada) |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=207–221 |doi=10.5194/fr-24-207-2021 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Béthoux, Deregnaucourt & Norrad ''in'' Béthoux ''et al.''
|
[[Carboniferous]] ([[Moscovian (Carboniferous)|Moscovian]])
|
Sunbury Creek Formation
|
{{Flag|Canada}}<br />({{Flag|New Brunswick}})
|
A member of [[Neodonatoptera]] of uncertain phylogenetic placement. The type species is ''B. norradi''.
|
|-
|
''[[Osnabruggiala]]''<ref>{{Cite journal|last1=Zessin |first1=W. |last2=Brauckmann |first2=C. |last3=Gröning |first3=E. |title=A new insect (probably basal Odonatoptera) from the Pennsylvanian (Late Carboniferous) of the Piesberg Fossil-Lagerstätte, Osnabrück, Germany |year=2021 |journal=Palaeoentomology |volume=4 |issue=6 |pages=532–536 |doi=10.11646/palaeoentomology.4.6.2 |s2cid=245170114 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Zessin, Brauckmann & Gröning
|
Carboniferous ([[Pennsylvanian (geology)|Pennsylvanian]])
|
Osnabrück Formation
|
{{Flag|Germany}}
|
Probably a [[Basal (phylogenetics)|basal]] member of [[Odonatoptera]]. The type species is ''O. seppelti''.
|
|-
|}
==Clade †Paoliidea==
===†Paoliida===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Avionblattinopsis]]''<ref>{{Cite journal|last1=Quispe |first1=L. |last2=Roques |first2=P. |last3=Garrouste |first3=R. |last4=Nel |first4=A. |year=2021 |title=Carboniferous Blattinopsidae: revision of ''Klebsiella'' and new genus and species from Avion (Insecta, Paoliida) |journal=Historical Biology |volume=34 |issue=3 |pages=383–389 |doi=10.1080/08912963.2021.1916817 |s2cid=234869567 |url=https://hal.sorbonne-universite.fr/hal-03232048/file/Quispe%20et%20al.%20-%202021%20-%20Carboniferous%20Blattinopsidae%20revision%20of%20Klebsiel.pdf }}</ref>
|
Gen. et sp. nov
|
In press
|
Quispe ''et al.''
|
Carboniferous (Moscovian)
|
|
{{Flag|France}}
|
A [[Blattinopsidae|blattinopsid]] [[Paoliida]]n.<br/> The type species is ''A. oudardi''. 
| 
|-
|
''[[Blattinopsis|Blattinopsis indefinitus]]''<ref>{{Cite journal|last1=Rasnitsyn |first1=A. P. |last2=Aristov |first2=D. S. |title=New species of the genus ''Blattinopsis'' Giebel, 1867 (Insecta: Blattinopsida: Blattinopsidae) from the Permian of Taimyr Peninsula, Russia |year=2021 |journal=Far Eastern Entomologist |volume=437 |pages=6–9 |doi=10.25221/fee.437.2 |s2cid=239636286 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
[[Alexandr Rasnitsyn|Rasnitsyn]] & Aristov
|
Permian ([[Kungurian]])
|
|
{{Flag|Russia}}<br>({{Flag|Krasnoyarsk Krai}})
|
A Blattinopsidae Paoliidan
|
|-
|
''[[Glaphyrophlebia|Glaphyrophlebia borea]]''<ref name=PJ556Glaphyrophlebia>{{Cite journal|last1=Aristov |first1=D. S. |last2=Rasnitsyn |first2=A. P. |last3=Naugolnykh |first3=S. V. |title=New Blattinopsidae (Insecta: Blattinopsida) in the Permian of the Pechora Basin (Komi Republic, Russia) in the context of landscape and vegetation evolution |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=641–649 |doi=10.1134/S0031030121060022 |s2cid=245009855 |url=https://www.elibrary.ru/item.asp?id=46621795 }}</ref>
|
Sp. nov
|
Valid
|
Aristov, Rasnitsyn & Naugolnykh
|
Permian
|
[[Inta Formation]]
|
{{Flag|Russia}}<br>({{Flag|Komi Republic}})
|
A Blattinopsid.
|
|-
|
''Glaphyrophlebia iva''<ref>{{Cite journal|last1=Aristov |first1=D. S. |last2=Rasnitsyn |first2=A. P. |title=New and little known Blattinopsidae (Insecta: Blattinopsida) from the Middle Permian of Russia |year=2021 |journal=Paleontological Journal |volume=55 |issue=3 |pages=288–293 |doi=10.1134/S0031030121030059 |s2cid=235966366 |url=https://elibrary.ru/item.asp?id=45690942 }}</ref>
|
Sp. nov
|
Valid
|
Aristov & Rasnitsyn
|
Middle Permian
|
|
{{Flag|Russia}}
|
A member of the family Blattinopsidae.
|
|-
|
''Glaphyrophlebia komia''<ref name=PJ556Glaphyrophlebia />
|
Sp. nov
|
Valid
|
Aristov, Rasnitsyn & Naugolnykh
|
Permian
|
Inta Formation
|
{{Flag|Russia}}<br>({{Flag|Komi Republic}})
|
A member of the family Blattinopsidae.
|
|-
|
''Glaphyrophlebia vorkutensis''<ref name=PJ556Glaphyrophlebia />
|
Sp. nov
|
Valid
|
Aristov, Rasnitsyn & Naugolnykh
|
Permian
|
Inta Formation
|
{{Flag|Russia}}<br>({{Flag|Komi Republic}})
|
A member of the family Blattinopsidae.
|
|-
|}

==Clade Paraneoptera==
===Hemipterans===
====New Hemipteran taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Acrotiara]]''<ref name=Acrotiarini>{{Cite journal|last1=Luo |first1=Y. |last2=Bourgoin |first2=T. |last3=Szwedo |first3=J. |last4=Feng |first4=J.-N. |title=Acrotiarini trib. nov., in the Cixiidae (Insecta, Hemiptera, Fulgoromorpha) from mid-Cretaceous amber of northern Myanmar, with new insights in the classification of the family |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104959 |doi=10.1016/j.cretres.2021.104959 |bibcode=2021CrRes.12804959L |doi-access=free }}</ref>
|
Trib. et gen. et sp. nov
|
In press
|
Bourgoin & Luo ''in'' Luo ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Cixiidae]]; the type genus of the new tribe [[Acrotiarini]]. The type species is ''A. multigranulata'' Luo & Bourgoin.
|
|-
|
''[[Aneurus damzeni]]''<ref>{{Cite journal|last1=Heiss |first1=E. |last2=Kairišs |first2=K. |last3=Bukejs |first3=A. |title=The invisible becomes visible: X-ray micro-CT reconstruction of ''Aneurus damzeni'' sp. nov. (Hemiptera: Heteroptera: Aradidae) from Eocene Baltic amber |year=2021 |journal=Zootaxa |volume=4950 |issue=1 |pages=189–195 |doi=10.11646/zootaxa.4950.1.11 |pmid=33903327 |s2cid=233409823 }}</ref>
|
Sp. nov
|
Valid
|
Heiss ''in'' Heiss, Kairišs & Bukejs
|
Eocene
|
Baltic amber
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
A species of ''[[Aneurus]]''.
|
|-
|
''[[Aphelicophontes]]''<ref>{{Cite journal|last1=Swanson |first1=D. R. |last2=Heads |first2=S. W. |last3=Taylor |first3=S. J. |last4=Wang |first4=Y. |year=2021 |title=A new remarkably preserved fossil assassin bug (Insecta, Heteroptera, Reduviidae) from the Eocene Green River Formation of Colorado |journal=Papers in Palaeontology |volume=7 |issue=3 |pages=1459–1478 |doi=10.1002/spp2.1349 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Swanson ''et al.''
|
[[Eocene]]
|
[[Green River Formation]]
|
{{Flag|United States}}
|
A member of the family [[Reduviidae]] belonging to the subfamily [[Harpactorinae]]. Genus includes new species ''A. danjuddi''.
|
|-
|
''[[Baltiomiris]]''<ref>{{Cite journal|last1=Kim |first1=J. |last2=Chérot |first2=F. |last3=Jung |first3=S. |year=2021 |title=A new genus and species of the subfamily Mirinae (Hemiptera: Cimicomorpha: Miridae) from Eocene Baltic amber with a new generic combination for ''Calocoris antennatus'' Statz in Statz & Wagner, 1950 |journal=Zootaxa |volume=5057 |issue=4 |pages=571–576 |doi=10.11646/zootaxa.5057.4.7 |pmid=34811191 |s2cid=239959240 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kim, Chérot & Jung
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Miridae]]. The type species is ''B. herczeki''.
|
|-
|
''[[Barremixius]]''<ref name=Acrotiarini />
|
Gen. et comb. nov
|
In press
|
Bourgoin & Szwedo ''in'' Luo ''et al.''
|
Early Cretaceous
|
[[Weald Clay]]
|
{{Flag|United Kingdom}}
|
A member of the family Cixiidae; a new genus for ''"[[Cixius]]" petrinus'' Fennah.
|
|-
|
''[[Beaconiella|Beaconiella cincta]]''<ref name=Triknightia>{{Cite journal|last=Shcherbakov |first=D. E. |title=New Curvicubitidae and Paraknightiidae (Homoptera: Cicadomorpha) from the Triassic of Central Asia |year=2021 |journal=Russian Entomological Journal |volume=30 |issue=2 |pages=129–134 |doi=10.15298/rusentj.30.2.02 |s2cid=242458292 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Shcherbakov
|
[[Triassic]] ([[Ladinian]]–[[Carnian]])
|
[[Madygen Formation]]
|
{{Flag|Kyrgyzstan}}
|
A member of [[Cicadomorpha]] belonging to the superfamily [[Pereborioidea]] and the family [[Curvicubitidae]].
|
|-
|
''Beaconiella pulchra''<ref name=Triknightia />
|
Sp. nov
|
Valid
|
Shcherbakov
|
Triassic (Ladinian–Carnian)
|
Madygen Formation
|
{{Flag|Kyrgyzstan}}
|
A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
|
|-
|
''[[Burmacader|Burmacader bicoloripennis]]''<ref>{{Cite journal|last1=Souma |first1=J. |last2=Yamamoto |first2=S. |last3=Takahashi |first3=Y. |title=A new species of the fossil lace bug genus ''Burmacader'' (Hemiptera: Heteroptera: Tingidae) from mid-Cretaceous Kachin amber from northern Myanmar |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104911 |doi=10.1016/j.cretres.2021.104911|issn=0195-6671 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Souma, Yamamoto & Takahashi
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Tingidae]].
|
|-
|
''Burmacader grandis''<ref>{{Cite journal|last1=Heiss |first1=E. |last2=Guilbert |first2=E. |title=A new species of ''Burmacader'' Heiss & Guilbert, 2013 from Cretaceous Burmese amber (Hemiptera, Heteroptera, Tingidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=30–33 |doi=10.11646/palaeoentomology.4.1.6 |s2cid=233966351 }}</ref>
|
Sp. nov
|
Valid
|
Heiss & Guilbert
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Tingidae]].
|
|-
|
''[[Burmala]]''<ref>{{Cite journal|last1=Liu |first1=G. |last2=Wang |first2=X. |last3=Zhuo |first3=D. |last4=Chen |first4=J. |title=A new non-jumping plant-louse (Hemiptera, Sternorrhyncha, Malmopsyllidae) in mid-Cretaceous Kachin amber, northern Myanmar |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104816 |doi=10.1016/j.cretres.2021.104816 |bibcode=2021CrRes.12404816L |s2cid=233619062 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Liu ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Malmopsyllidae]]. Genus includes new species ''B. liaoyaoi''.
|
|-
|
''[[Burmaprosbole]]''<ref>{{Cite journal|last1=Qiao |first1=D. |last2=Zhang |first2=W. |last3=Zhang |first3=X. |last4=Ren |first4=D. |last5=Yao |first5=Y. |title=New genus and species of Tettigarctidae (Hemiptera: Cicadomorpha: Cicadoidea) from the mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104900 |doi=10.1016/j.cretres.2021.104900 |bibcode=2021CrRes.12604900Q }}</ref>
|
Gen. et sp. nov
|
Valid
|
Qiao ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Tettigarctidae]]. Genus includes new species ''B. nervosa''.
|
|-
|
''[[Burmavianaida]]''<ref>{{Cite journal|last1=Souma |first1=J. |last2=Yamamoto |first2=S. |last3=Takahashi |first3=Y. |title=Discovery of the Smallest Lace Bug from Mid-Cretaceous of Northern Myanmar Supports the Hypothesis of a Miniaturization Phenomenon of Insects in Kachin Amber (Hemiptera: Heteroptera: Tingidae) |year=2021 |journal=Taxonomy |volume=1 |issue=2 |pages=60–68 |doi=10.3390/taxonomy1020007 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Souma, Yamamoto & Takahashi
|
Cretaceous (Albian or Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Tingidae. The type species is ''B. anomalocapitata''.
|
|-
|
''[[Cretamystilus]]''<ref>{{Cite journal|last1=Kim |first1=J. |last2=Lim |first2=H. |last3=Jung |first3=S. |title=A first member of bamboo-feeding lineage plant bug (Hemiptera: Heteroptera: Miridae: Mecistoscelini) in mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=121 |pages=Article 104741 |doi=10.1016/j.cretres.2020.104741 |bibcode=2021CrRes.12104741K |s2cid=233567365 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kim, Lim & Jung
|
[[Late Cretaceous]] ([[Cenomanian]])
|
[[Burmese amber]]
|
{{Flag|Myanmar}}
|
A member of the family [[Miridae]] belonging to the tribe [[Mecistoscelini]]. Genus includes new species ''C. herczeki''.
|
|-
|
''[[Cretodorus|Cretodorus rostellatus]]''<ref>{{Cite journal|last1=Zhang |first1=X. |last2=Yao |first2=Y. |last3=Ren |first3=D. |last4=Pang |first4=H. |title=A new species of ''Cretodorus'' (Hemiptera: Fulgoromorpha: Fulgoroidea: Mimarachnidae) from Upper Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104988 |doi=10.1016/j.cretres.2021.104988 |bibcode=2021CrRes.12804988Z |s2cid=238849433 }}</ref>
|
Sp. nov
|
In press
|
Zhang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Mimarachnidae]].
|
|-
|
''[[Cretomultinervis]]''<ref>{{Cite journal|last1=Fu |first1=Y. |last2=Huang |first2=D. |title=A new sinoalid froghopper in mid-Cretaceous amber from northern Myanmar (Hemiptera, Cicadomorpha, Sinoalidae) |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104841 |doi=10.1016/j.cretres.2021.104841 |bibcode=2021CrRes.12504841F }}</ref>
|
Gen. et sp. nov
|
Valid
|
Fu & Huang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Sinoalidae]]. Genus includes new species ''C. burmensis''.
|
|-
|
''[[Xiaochibangus|Dachibangus hui]]''<ref name=Multistria>{{Cite journal|last1=Zhang |first1=X. |last2=Yao |first2=Y. |last3=Ren |first3=D. |last4=Pang |first4=H. |last5=Chen |first5=H. |title=New mimarachnids (Hemiptera, Fulgoromorpha, Fulgoroidea) in mid-Cretaceous Burmese amber |year=2021 |journal=ZooKeys |issue=1057 |pages=37–48 |doi=10.3897/zookeys.1057.66434 |pmid=34539197 |pmc=8410752 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Zhang, Yao & Pang ''in'' Zhang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Mimarachnidae. Originally described as a species of ''[[Dachibangus]]'', but subsequently transferred to the genus ''Xiaochibangus''.<ref>{{Cite journal|last1=Jiang |first1=H. |last2=Chen |first2=J. |last3=Szwedo |first3=J. |title=A new ''Jaculistilus'' species of Mimarachnidae (Hemiptera: Fulgoromorpha) from mid-Cretaceous Kachin amber of northern Myanmar, with geometric morphometric analysis of the mimarachnid genera |year=2022 |journal=Cretaceous Research |volume=141 |at=105368 |doi=10.1016/j.cretres.2022.105368 |s2cid=252413881 }}</ref>
|
|-
|
''[[Delphitiara]]''<ref name=Acrotiarini />
|
Gen. et sp. nov
|
In press
|
Bourgoin & Luo ''in'' Luo ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is ''D. tibiocoronata'' Luo & Bourgoin.
|
|-
|
''[[Dorytocus|Dorytocus jiaxiaoae]]''<ref>{{Cite journal|last1=Song |first1=Z.-S. |last2=Zhang |first2=C.-L. |last3=Xi |first3=H.-Y. |last4=Szwedo |first4=J. |last5=Bourgoin |first5=T. |title=First record of adult Dorytocidae — ''Dorytocus jiaxiaoae'' Song, Szwedo & Bourgoin sp. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea) from mid-Cretaceous Kachin amber |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104863 |doi=10.1016/j.cretres.2021.104863 |bibcode=2021CrRes.12504863S }}</ref>
|
Sp. nov
|
Valid
|
Song, Szwedo & Bourgoin ''in'' Song ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[planthopper]].
|
|-
|
''[[Duyana]]''<ref>{{Cite journal|last1=Chen |first1=J. |last2=Jiang |first2=H. |last3=Shang |first3=T. |last4=Zhang |first4=L. |last5=Zheng |first5=Y. |last6=Zhang |first6=Q. |last7=Zhang |first7=H. |title=The second leafhopper of the subfamily Ledrinae (Hemiptera, Cicadellidae) in mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=122 |pages=Article 104763 |doi=10.1016/j.cretres.2021.104763 |bibcode=2021CrRes.12204763C |s2cid=233579718 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Chen ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[leafhopper]] belonging to the subfamily [[Ledrinae]]. Genus includes new species ''D. electronica''.
|
|-
|
''[[Electromyiomma|Electromyiomma herczeki]]''<ref>{{Cite journal|last1=Kim |first1=J. |last2=Jung |first2=S. |year=2021 |title=A new species of extinct genus ''Electromyiomma'' Popov & Herczek (Hemiptera: Miridae: Isometopinae) from Eocene Baltic amber, with the key to ''Electromyiomma'' species |journal=Zootaxa |volume=4927 |issue=2 |pages=289–293 |doi=10.11646/zootaxa.4927.2.9 |pmid=33756714 |s2cid=232339534 }}</ref>
|
Sp. nov
|
Valid
|
Kim & Jung
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Miridae]].
|
|-
|
''[[Eospinosus]]''<ref>{{cite journal |last1=Wedmann |first1=S. |last2=Kment |first2=P. |last3=Campos |first3=L. A. |last4=Hörnschemeyer |first4=T. |year=2021 |title=Bizarre morphology in extinct Eocene bugs (Heteroptera: Pentatomidae) |journal=Royal Society Open Science |volume=8 |issue=12 |pages=Article ID 211466 |doi=10.1098/rsos.211466 |pmid=34909219 |pmc=8652274 |bibcode=2021RSOS....811466W |doi-access=free }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Wedmann ''et al.''
|
Eocene
|
[[Messel pit]]
|
{{Flag|Germany}}<br>
{{Flag|United States}}<br>({{Flag|Colorado}})
|
A member of the family [[Pentatomidae]]. The type species is ''E. peterkulkai''; genus also includes ''E. greenriverensis''.
|
|-
|
''[[Ferriantenna]]''<ref>{{Cite journal|last1=Cumming |first1=R. T. |last2=Le Tirant |first2=S. |title=Drawing the Excalibur bug from the stone: adding credibility to the double-edged sword hypothesis of coreid evolution (Hemiptera, Coreidae) |year=2021 |journal=ZooKeys |issue=1043 |pages=117–131 |doi=10.3897/zookeys.1043.67730 |doi-access=free |pmid=34163298 |pmc=8217075 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cumming & Le Tirant
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Coreidae]] belonging to the subfamily [[Coreinae]]. The type species is ''F. excalibur''.
|
|-
|
''[[Hallex martinsnetoi]]''<ref>{{Cite journal|last1=Monferran |first1=M. D. |last2=de Lima |first2=F. J. |last3=Kellner |first3=A. W. A. |last4=Sayão |first4=J. M. |last5=Filho |first5=E. B. S. |last6=Saraiva |first6=A. Á. F. |last7=Sabater |first7=L. M. |last8=Bantim |first8=R. A. M. |title=An unusual preservation of a new species of Cicadomorpha (Auchenorrhyncha) from the Crato formation, Lower Cretaceous of Brazil |journal=Historical Biology |year=2022 |volume=34 |issue=7 |pages=1166–1174 |doi=10.1080/08912963.2021.1966776 |s2cid=244239311 }}</ref>
|
Sp. nov
|
In press
|
Monferran ''et al.''
|
[[Early Cretaceous]]
|
[[Crato Formation]]
|
{{Flag|Brazil}}
|
A [[leafhopper]].
|
|-
|
''[[Hypsotsijilia]]''<ref>{{Cite journal|last1=Cifuentes-Ruiz |first1=P. |last2=Brailovsky |first2=H. |last3=Serrano-Sánchez |first3=M. L. |last4=Vega |first4=F. J. |year=2021 |title=First fossil Schizopteridae (Hemiptera: Heteroptera) in Miocene amber from Chiapas, Mexico |journal=Boletín de la Sociedad Geológica Mexicana |volume=73 |issue=3 |pages=A170121 |doi=10.18268/BSGM2021v73n3a170121 |s2cid=245031228 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cifuentes-Ruiz & Brailovsky ''in'' Cifuentes-Ruiz ''et al.''
|
Early [[Miocene]]
|
[[La Quinta Formation, Mexico|La Quinta Formation]]<br>([[Mexican amber]])
|
{{Flag|Mexico}}
|
A member of the family [[Schizopteridae]]. The type species is ''H. bretoni''.
|
|-
|
''[[Inoderbe]]''<ref>{{Cite journal|last1=Shcherbakov |first1=D. E. |last2=Emeljanov |first2=A.F. |title=Paradoxical derbid-like planthopper (Homoptera: Fulgoroidea) from Cretaceous Burmese amber |year=2021 |journal=Russian Entomological Journal |volume=30 |issue=2 |pages=135–139 |doi=10.15298/rusentj.30.2.03 |s2cid=237815990 |doi-access=free }}</ref>
|
Fam. et gen. et sp. nov
|
Valid
|
Shcherbakov & Emeljanov
|
Cretaceous (Albian–Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[planthopper]] belonging to the new family [[Inoderbidae]]. The type species is ''I. rapunzel''.
|
|-
|
''[[Juroala|Juroala pulchra]]''<ref>{{Cite journal|last1=Fu |first1=Y.Z. |last2=Huang |first2=D.Y. |title=A new species of Sinoalidae from the topmost Late Jurassic Daohugou Bed (Hemiptera, Cicadomorpha) |year=2021 |journal=Palaeoentomology |volume=4 |issue=1 |pages=34–38 |doi=10.11646/palaeoentomology.4.1.7 |s2cid=233954621 }}</ref>
|
Sp. nov
|
Valid
|
Fu & Huang
|
Late Jurassic
|
[[Daohugou Beds]]
|
{{Flag|China}}
|
A member of the family [[Sinoalidae]].
|
|-
|
''[[Karatauella]]''<ref>{{Cite journal|last=Doweld |first=A. B. |title=''Karatauella'', a new generic replacement name for fossil ''Carpenterella'' Becker-Migdisova, 1968 (Insecta: Hemiptera: Protopsyllidiidae ''incertae sedis'') |year=2021 |journal=Zootaxa |volume=5047 |issue=1 |pages=95–96 |doi=10.11646/zootaxa.5047.1.9 |pmid=34811005 |s2cid=244258647 }}</ref>
|
Nom. nov
|
Valid
|
Doweld
|
[[Jurassic]] ([[Bathonian]]-[[Oxfordian (stage)|Oxfordian]])
|
|
{{Flag|Kazakhstan}}
|
A member of the family [[Protopsyllidiidae]]; a replacement name for ''Carpenterella'' Becker-Migdisova (1968).
|
|-
|
''[[Latidorsum]]''<ref>{{Cite journal|last1=Wang |first1=Y. |last2=Tang |first2=H. |last3=Ren |first3=D. |last4=Yao |first4=Y. |title=New Fossil Tingidae (Hemiptera: Heteroptera) from the Mid-Cretaceous of Myanmar, with Remarks on the Phylogenetic Relationships within the Family |year=2021 |journal=Insects |volume=12 |issue=10 |pages=Article 887 |doi=10.3390/insects12100887 |pmid=34680658 |pmc=8540450 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
|
Wang, Tang & Yao ''in'' Wang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Tingidae. The type species is ''L. carinbifarium''.
|
|-
|
''Microtingis''<ref>{{Cite journal|last1=Heiss |first1=E. |last2=Golub |first2=V. B. |title=A new genus and species of Tingidae from Cretaceous Burmese amber (Hemiptera, Heteroptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=2 |pages=137–140 |doi=10.11646/palaeoentomology.4.2.2 |s2cid=235539280 }}</ref>
|
Gen. et sp. nov
|
[[Synonym (taxonomy)|Junior synonym]]
|
Heiss & Golub
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Tingidae. Genus includes new species ''M. leptosoma''. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of ''Burmavianaida anomalocapitata''.<ref>{{Cite journal|last1=Heiss |first1=E. |last2=Golub |first2=V. B. |title=New synonymy in Burmese amber Tingidae (Hemiptera, Heteroptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=202 |doi=10.11646/palaeoentomology.4.3.2 |s2cid=237896242 }}</ref>
|
|-
|
''[[Multistria]]''<ref name=Multistria />
|
Gen. et sp. nov
|
Valid
|
Zhang, Yao & Pang ''in'' Zhang ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Mimarachnidae. The type species is ''M. orthotropa''.
|
|-
|
''[[Paleocader|Paleocader balticus]]''<ref name=Palaeoentomology44Paleocader>{{Cite journal|last1=Heiss |first1=E. |last2=Golub |first2=V. B. |title=Three new species of the fossil Cantacaderinae genus ''Paleocader'' Froeschner, 1996 from the Baltic and Rovno amber (Hemiptera, Heteroptera, Tingidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=4 |pages=360–366 |doi=10.11646/palaeoentomology.4.4.11 |s2cid=239623453 }}</ref>
|
Sp. nov
|
Valid
|
Heiss & Golub
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family [[Tingidae]].
|
|-
|
''Paleocader pulchellus''<ref name=Palaeoentomology44Paleocader />
|
Sp. nov
|
Valid
|
Heiss & Golub
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A member of the family Tingidae.
|
|-
|
''Paleocader rovnensis''<ref name=Palaeoentomology44Paleocader />
|
Sp. nov
|
Valid
|
Heiss & Golub
|
Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Tingidae.
|
|-
|
''[[Parasinalda|Parasinalda sukachevae]]''<ref>{{Cite journal|last1=Golub |first1=V. B. |last2=Perkovsky |first2=E. E. |last3=Vasilenko |first3=D. V. |year=2021 |title=A new fossil species of the genus ''Parasinalda'' Heiss & Golub (Hemiptera: Heteroptera: Tingidae) from Upper Eocene Rovno amber |journal=Zootaxa |volume=5027 |issue=2 |pages=290–296 |doi=10.11646/zootaxa.5027.2.9|issn=1175-5326 |pmid=34811229 |s2cid=239703141 }}</ref>
|
Sp. nov
|
Valid
|
Golub, Perkovsky & Vasilenko
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A member of the family Tingidae.
|
|-
|
''[[Pentacarinus]]''<ref name=Acrotiarini />
|
Gen. et sp. nov
|
In press
|
Bourgoin & Luo ''in'' Luo ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is ''P. kachinensis'' Luo & Bourgoin.
|
|-
|
''[[Pseudocaulisoculus]]''<ref>{{Cite journal|last1=Kóbor |first1=P. |last2=Roca-Cusachs |first2=M. |title=''Pseudocaulisoculus longicornis'': description of a new yuripopovinid true bug from mid-Cretaceous Burmese amber (Heteroptera: Coreoidea: Yuripopovinidae) |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104849 |doi=10.1016/j.cretres.2021.104849 |bibcode=2021CrRes.12504849K |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kóbor & Roca-Cusachs
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Yuripopovinidae]]. Genus includes new species ''P. longicornis''.
|
|-
|
''[[Pubivetanthocoris]]''<ref>{{Cite journal|last1=Tang |first1=H. |last2=Wang |first2=Y. |last3=Ren |first3=D. |last4=Yao |first4=Y. |title=The first record of vetanthocorids (Hemiptera: Heteroptera: Vetanthocoridae) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=131 |pages=Article 105090 |doi=10.1016/j.cretres.2021.105090| issn=0195-6671 |s2cid=244081314 }}</ref>
|
Gen. et sp. nov
|
In press
|
Tang, Wang & Yao ''in'' Tang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family [[Vetanthocoridae]]. Genus includes new species ''P. carinalis''.
|
|-
|
''[[Pyrrhocoris rottensis|"Pyrrhocoris" rottensis]]''<ref>{{Cite journal|last=Kment |first=P. |year=2021 |title=A review of the fossil taxa originally assigned to Pyrrhocoridae (Hemiptera: Heteroptera) |journal=Zootaxa |volume=5081 |issue=4 |pages=579–586 |doi=10.11646/zootaxa.5081.4.8 |pmid=35390991 |s2cid=245135992 }}</ref>
|
Nom. nov
|
Valid
|
Kment
|
Late [[Oligocene]]
|
|
{{Flag|Germany}}
|
A member of [[Lygaeoidea]] of uncertain phylogenetic placement; a replacement name for ''Pyrrhocoris tibialis'' Statz & Wagner (1950).
|
|-
|
''[[Rovnoxestus]]''<ref>{{Cite journal|last1=Dietrich |first1=C. H. |last2=Dmitriev |first2=D. A. |last3=Perkovsky |first3=E. E. |title=A new leafhopper (Hemiptera: Cicadellidae) from late Eocene Rovno amber described based on an adult and associated last-instar nymph from Perebrody (Ukraine) |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=445–452 |doi=10.11646/palaeoentomology.4.5.10 |s2cid=240525234 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Dietrich, Dmitriev & Perkovsky
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A [[leafhopper]]. Genus includes new species ''R. rasnitsyni''.
|
|-
|
''[[Saltissus|Saltissus fennahi]]''<ref>{{Cite journal|last1=Luo |first1=C. |last2=Liu |first2=X. |last3=Jiang |first3=T. |last4=Song |first4=Z. |last5=Jarzembowski |first5=E. A. |title=The first mimarachnid planthopper (Hemiptera: Fulgoromorpha: Mimarachnidae: ''Saltissus'') from the Wealden (Lower Cretaceous) of southern England |journal=Historical Biology |year=2022 |volume=34 |issue=10 |pages=2031–2036 |doi=10.1080/08912963.2021.1998035 |s2cid=243487825 }}</ref>
|
Sp. nov
|
In press
|
Luo, Liu & Jarzembowski ''in'' Luo ''et al.''
|
[[Early Cretaceous]]
|
[[Weald Clay]]
|
{{Flag|United Kingdom}}
|
A member of the family Mimarachnidae.
|
|-
|
''[[Scytinoptera|Scytinoptera tongchuanensis]]''<ref>{{Cite journal|last1=Zhang |first1=Q.-Q. |last2=Zheng |first2=D.-R. |last3=Teng |first3=X. |last4=Zhang |first4=H.-C. |title=New Scytinopteridae (Hemiptera: Scytinopteroidea) from the middle Triassic Tongchuan Entomofauna of NW China |journal=Historical Biology |year=2022 |volume=34 |issue=11 |pages=2259–2264 |doi=10.1080/08912963.2021.2010194 |s2cid=244916999 }}</ref>
|
Sp. nov
|
In press
|
Zhang, Zheng & Zhang ''in'' Zhang ''et al.''
|
[[Middle Triassic]]
|
Tongchuan Formation
|
{{Flag|China}}
|
A member of the family [[Scytinopteridae]].
|
|-
|
''[[Sinogranulus]]''<ref>{{Cite journal|last1=Zhang |first1=Q.-Q. |last2=Chen |first2=J. |last3=Zhang |first3=H.-C. |year=2021 |title=New Granulidae (Hemiptera: Scytinopteroidea) from the Middle Triassic Tongchuan Formation of NW China |journal=Historical Biology |volume=34 |pages=152–157 |doi=10.1080/08912963.2021.1903892 |s2cid=234836364 }}</ref>
|
Gen. et sp. nov
|
In press
|
Zhang, Chen & Zhang
|
[[Middle Triassic]]
|
[[Tongchuan Formation]]
|
{{Flag|China}}
|
A member of [[Scytinopteroidea]] belonging to the family [[Granulidae]]. Genus includes new species ''S. qishuiheensis''.
| 
|-
|
''[[Spinonympha]]''<ref>{{Cite journal|last1=Luo |first1=C. |last2=Wang |first2=B. |last3=Jarzembowski |first3=E. A. |title=A Bizarre Planthopper Nymph (Hemiptera: Fulgoroidea) from Mid-Cretaceous Kachin Amber |year=2021 |journal=Insects |volume=12 |issue=4 |pages=Article 318 |doi=10.3390/insects12040318 |pmid=33918264 |pmc=8066557 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Luo, Wang & Jarzembowski
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[planthopper]] of uncertain phylogenetic placement. The type species is ''S. shcherbakovi''.
|
|-
|
''[[Szeiinia]]''<ref>{{Cite journal|last1=Zhang |first1=Q. |last2=Jiang |first2=T. |last3=Szwedo |first3=J. |last4=Zhang |first4=H. |title=A new family of Triassic planthoppers (Hemiptera: Fulgoromorpha: Fulgoroidea) from the Shaanxi Province of China |year=2021 |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=45 |issue=1 |pages=86–90 |doi=10.1080/03115518.2021.1919206 |bibcode=2021Alch...45...86Z |s2cid=235748056 }}</ref>
|
Fam. nov., Gen. et sp. nov
|
Valid
|
Zhang ''et al.''
|
[[Late Triassic]]
|
[[Yanchang Formation]]
|
{{Flag|China}}
|
A [[planthopper]] in the new family [[Szeiiniidae]]. Genus includes new species ''S. huanglongensis''
|
|-
|
''[[Tennentsia|Tennentsia orientalis]]''<ref>{{Cite journal|last1=Fu |first1=Y. |last2=Azar |first2=D. |last3=Huang |first3=D. |year=2021 |title=The first Dysmorphoptilidae from the Middle Triassic of China (Hemiptera: Cicadomorpha) |journal=Historical Biology |volume=33 |issue=12 |pages=3506–3512 |doi=10.1080/08912963.2021.1874374 |s2cid=233710366 }}</ref>
|
Sp. nov
|
In press
|
Fu, Azar & Huang
|
[[Middle Triassic]]
|
[[Yanchang Formation]]
|
{{Flag|China}}
|
A member of [[Cicadomorpha]] belonging to the family [[Dysmorphoptilidae]].
|
|-
|
''[[Triknightia]]''<ref name=Triknightia />
|
Gen. et sp. nov
|
Valid
|
Shcherbakov
|
Triassic (Ladinian–Carnian)
|
Madygen Formation
|
{{Flag|Kyrgyzstan}}
|
A member of Cicadomorpha belonging to the superfamily [[Scytinopteroidea]] and the family [[Paraknightiidae]]. The type species is ''T. mira''.
|
|-
|
''[[Tuthillia maculosa]]''<ref>{{Cite journal|last1=Poinar Jr. |first1=G. |last2=Brown |first2=A. E. |title=A new species of plant louse from mid-Tertiary Dominican amber (Hemiptera: Psylloidea) with adjacent plant leaves |year=2021 |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=302 |issue=2 |pages=139–146 |doi=10.1127/njgpa/2021/1024 |s2cid=243913929 }}</ref>
|
Sp. nov
|
Valid
|
Poinar & Brown
|
|
[[Dominican amber]]
|
{{Flag|Dominican Republic}}
|
A member of [[Psylloidea]].
|
|-
|
''[[Valdicopis]]''<ref>{{Cite journal|last1=Li |first1=Y. |last2=Chen |first2=J. |last3=Jarzembowski |first3=E. |title=The first true procercopid (Hemiptera, Cercopoidea) from England |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104933 |doi=10.1016/j.cretres.2021.104933 |bibcode=2021CrRes.12704933L }}</ref>
|
Gen. et sp. nov
|
Valid
|
Li, Chen & Jarzembowski
|
Early Cretaceous
|
|
{{Flag|United Kingdom}}
|
A member of the family [[Procercopidae]]. Genus includes new species ''V. tonyi''.
|
|-
|
''[[Youngeewa simillima]]''<ref>{{Cite journal|last1=Gębicki |first1=C. |last2=Walczak |first2=M. |last3=Zmarzły |first3=M. |title=''Youngeewa simillima'' sp. nov. (Hemiptera: Cicadellidae: Mileewinae) from Eocene Baltic Amber |year=2021 |journal=Bonn Zoological Bulletin |volume=70 |issue=2 |pages=351–357 |doi=10.20363/BZB-2021.70.2.351 |url=https://zoologicalbulletin.de/BzB_Volumes/Volume_70_2/351_gebicki_20211029.pdf }}</ref>
|
Sp. nov
|
Valid
|
Gębicki & Walczak ''in'' Gębicki, Walczak & Zmarzły
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[leafhopper]] belonging to the subfamily [[Mileewinae]].
|
|-
|}

====Hemipteran research====
* [[David Grimaldi (entomologist)|Grimaldi]] & Vea (2021) reinterpret ''[[Mesophthirus]] engeli'' as a [[scale insect]], and reject the interpretation of this insect as an [[ectoparasite]].<ref>{{Cite journal|last1=Grimaldi |first1=D. A. |last2=Vea |first2=I. M. |title=Insects with 100 million-year-old dinosaur feathers are not ectoparasites |year=2021 |journal=Nature Communications |volume=12 |issue=1 |pages=Article number 1469 |doi=10.1038/s41467-021-21751-x |doi-access=free |pmid=33674573 |pmc=7935990 |bibcode=2021NatCo..12.1469G }}</ref>
* Redescription of the [[holotype]] specimen of ''Paranoika placida'' is published by Moura-Júnior ''et al.'' (2021), who transfer this species to the genus ''[[Lethocerus]]'', and consider this species to be the [[Synonym (taxonomy)|senior synonym]] of ''Lethocerus vetus''.<ref>{{Cite journal|last1=Moura-Júnior |first1=D. A. |last2=Scheffler |first2=S. M. |last3=Nel |first3=A. |last4=Mejdalani |first4=G. |title=Paranoikidae Zamboni, Martins-Neto & Popov, 2002, a junior synonym of Belostomatidae Leach, 1815 (Hemiptera: Heteroptera): redescription of a giant water bug from Crato Formation, Lower Cretaceous of Brazil |year=2021 |journal=Cretaceous Research |volume=124 |pages=Article 104796 |doi=10.1016/j.cretres.2021.104796 |bibcode=2021CrRes.12404796M |s2cid=233701548 }}</ref>
* A [[Fulgoridae|fulgorid]] specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu ''et al.'' (2021).<ref>{{Cite journal|last1=Xu |first1=X.-T. |last2=Deng |first2=W.-Y.-D. |last3=Zhou |first3=Z.-K. |last4=Wappler |first4=T. |last5=Su |first5=T. |title=The first Fulgoridae (Hemiptera: Fulgoromorpha) from the Eocene of the central Qinghai–Tibetan Plateau |year=2021 |journal=Fossil Record |volume=24 |issue=2 |pages=263–274 |doi=10.5194/fr-24-263-2021 |doi-access=free }}</ref>
* Revision of the fossil record of [[Fulgoromorpha]] and [[Cicadomorpha]] from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).<ref>{{cite book |last=Chen |first=J. |year=2021 |chapter=True hopper fossils (Fulgoromorpha and Cicadomorpha) in the Jurassic to Cretaceous of eastern Asia and their evolutionary implications |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |volume=521 |pages=61–80 |publisher=The Geological Society of London |doi=10.1144/SP521-2021-18 |s2cid=244098374 }}</ref>

===Psocodeans===
====New Psocodean taxa====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Burmesopsocus]]''<ref>{{Cite journal|last1=Yoshizawa |first1=K. |last2=Yamamoto |first2=S. |title=The earliest fossil record of the suborder Psocomorpha (Insecta: Psocodea) from mid-Cretaceous Burmese amber, with description of a new genus and species |year=2021 |journal=Insecta Matsumurana |series=New Series |volume=77 |pages=1–15 |hdl=2115/82942 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Yoshizawa ''in'' Yoshizawa & Yamamoto
|
Cretaceous (Albian–Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Homilopsocidea]] of uncertain phylogenetic placement. The type species is ''B. lienhardi''.
|
|-
|
''[[Cormopsocus|Cormopsocus baleoi]]''<ref>{{Cite journal|last1=Hakim |first1=M. |last2=Azar |first2=D. |last3=Huang |first3=D.-Y. |title=A new species of Cormopsocidae from Burmese amber (Psocodea; Trogiomorpha) |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=213–217 |doi=10.11646/palaeoentomology.4.3.6 |s2cid=237857111 }}</ref>
|
Sp. nov
|
Valid
|
Hakim, Azar & Huang
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Trogiomorpha]] belonging to the family [[Cormopsocidae]].
|
|-
|
''Cormopsocus neli''<ref>{{Cite journal|last1=Hakim |first1=M. |last2=Azar |first2=D. |last3=Fu |first3=Y.-Z. |last4=Cai |first4=C.-Y. |last5=Huang |first5=D.-Y. |title=A new cormopsocid from mid-Cretaceous Burmese amber (Psocodea: Trogiomorpha: Cormopsocidae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=2 |pages=178–185 |doi=10.11646/palaeoentomology.4.2.7 | issn=2624-2826  |s2cid=236620426 }}</ref>
|
Sp. nov
|
Valid
|
Hakim, Azar & Huang ''in'' Hakim ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Trogiomorpha belonging to the family Cormopsocidae.
|
|-
|
''[[Empherium]]''<ref name=Empherium>{{Cite journal|last1=Hakim |first1=M. |last2=Huang |first2=D.-Y. |last3=Azar |first3=D. |title=New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae) |year=2021 |journal=Palaeoentomology |volume=4 |issue=2 |pages=196–198 |doi=10.11646/palaeoentomology.4.2.8 |s2cid=235564657 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Hakim, Huang & Azar
|
Late Cretaceous
|
|
{{Flag|Russia}}
|
A member of Trogiomorpha belonging to the family [[Empheriidae]]. Genus includes new species ''E. rasnitsyni''.
|
|-
|
''[[Eolepinotus|Eolepinotus zherikhini]]''<ref name=Empherium />
|
Sp. nov
|
Valid
|
Hakim, Huang & Azar
|
Late Cretaceous
|
|
{{Flag|Russia}}
|
A member of Trogiomorpha belonging to the family [[Trogiidae]].
|
|-
|
''[[Heliadesdakruon]]''<ref>{{Cite journal|last1=Cumming |first1=R. T. |last2=Le Tirant |first2=S. |title=Review of the Cretaceous †Archaeatropidae and †Empheriidae and description of a new genus and species from Burmese amber (Psocoptera) |year=2021 |journal=Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire |volume=9 |issue=16 |pages=1–11 |url=https://www.researchgate.net/publication/351351269 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cumming & Le Tirant
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of [[Trogiomorpha]] belonging to the family [[Archaeatropidae]]. The type species is ''H. morganae''.
|
|-
|
''[[Longiglabellus]]''<ref>{{Cite journal|last1=Wang |first1=Q. |last2=Li |first2=S. |last3=Ren |first3=D. |last4=Yao |first4=Y. |title=New genus and species of †Cormopsocidae (Psocodae: Trogiomorpha) from mid-Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104992 |doi=10.1016/j.cretres.2021.104992 |bibcode=2021CrRes.12804992W }}</ref>
|
Gen. et 2 sp. nov
|
In press
|
Wang, Li & Yao ''in'' Wang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Cormopsocidae. Genus includes new species ''L. edentatus'' and ''L. pedhyalinus''.
|
|-
|
''[[Palaeosiamoglaris|Palaeosiamoglaris hammanaensis]]''<ref>{{Cite journal|last1=Hakim |first1=M. |last2=Huang |first2=D.-Y. |last3=Azar |first3=D. |title=Earliest record of Prionoglarididae from the Lower Cretaceous Lebanese amber (Psocodea; Trogiomorpha) |year=2021 |journal=Cretaceous Research |volume=132 |pages=Article 105121 |doi=10.1016/j.cretres.2021.105121 |s2cid=245304011 }}</ref>
|
Sp. nov
|
In press
|
Hakim, Huang & Azar
|
Early Cretaceous (Barremian)
|
[[Lebanese amber]]
|
{{Flag|Lebanon}}
|
A member of [[Psocodea]] belonging to the family [[Prionoglarididae]].
|
|-
|
''Palaeosiamoglaris hkamtiensis''<ref name=JouaultetalCRPsocodea>{{Cite journal|last1=Jouault |first1=C. |last2=Yoshizawa |first2=K. |last3=Hakim |first3=M. |last4=Huang |first4=D. |last5=Nel |first5=A. |title=New psocids (Psocodea: Prionoglarididae, Psyllipsocidae) from Cretaceous Burmese amber deposits |year=2021 |journal=Cretaceous Research |volume=126 |pages=Article 104890 |doi=10.1016/j.cretres.2021.104890 |bibcode=2021CrRes.12604890J |url=https://hal-insu.archives-ouvertes.fr/insu-03227758/file/jouault-2021.pdf }}</ref>
|
Sp. nov
|
In press
|
Jouault ''et al.''
|
Early Cretaceous
|
Hkamti amber
|
{{Flag|Myanmar}}
|
A member of Psocodea belonging to the family Prionoglarididae.
|
|-
|
''[[Psyllipsocus myanmarensis]]''<ref name=JouaultetalCRPsocodea />
|
Sp. nov
|
In press
|
Jouault ''et al.''
|
Late Cretaceous (Cenomanian)
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Psocodea belonging to the family [[Psyllipsocidae]], a species of ''[[Psyllipsocus]]''.
|
|-
|
''[[Psyllipsocus yangi]]''<ref>{{Cite journal|last1=Liang |first1=F. |last2=Liu |first2=X. |year=2021 |title=A new species of ''Psyllipsocus'' (Psocodea: Trogiomorpha: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar |journal=Zootaxa |volume=5072 |issue=1 |pages=81–87 |doi=10.11646/zootaxa.5072.1.9 |pmid=35390882 |s2cid=244682740 }}</ref>
|
Sp. nov
|
Valid
|
Liang & Liu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of Psocodea belonging to the family Psyllipsocidae, a species of ''Psyllipsocus''.
|
|-
|
''[[Stimulopsocus]]''<ref>{{Cite journal|last1=Liang |first1=F. |last2=Liu |first2=X. |title=A new genus and species of the family Cormopsocidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Myanmar |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105049 |doi=10.1016/j.cretres.2021.105049 |s2cid=244225553 }}</ref>
|
Gen. et sp. nov
|
In press
|
Liang & Liu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A member of the family Cormopsocidae. Genus includes new species ''S. jiewenae''.
|
|-
|}

====Psocodean research====
* Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller ''et al.'' (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.<ref>{{Cite journal|last1=Kiesmüller |first1=C. |last2=Haug |first2=J. T. |last3=Müller |first3=P. |last4=Hörnig |first4=M. K. |title=Debris-carrying behaviour of bark lice immatures preserved in 100 million years old amber |year=2021 |journal=PalZ |volume=96 |issue=2 |pages=231–258 |doi=10.1007/s12542-021-00567-6 |doi-access=free }}</ref>

==Clade Perlidea==
===Dermapterans===
====New Dermapteran====
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Aglyptoderma]]''<ref>{{Cite journal|last1=Xiong |first1=S. |last2=Engel |first2=M. S. |last3=Xiao |first3=L. |last4=Ren |first4=D. |title=New eodermapteran earwigs (Dermaptera) from the Middle Jurassic Jiulongshan Formation of China |year=2021 |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=45 |issue=3 |pages=335–343 |doi=10.1080/03115518.2021.1965215 |bibcode=2021Alch...45..335X |s2cid=244103979 }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Xiong ''et al.''
|
[[Middle Jurassic]]
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A [[Semenoviolidae|semenoviolid]] [[eodermaptera]]n earwig.<br/> Genus includes the new species ''A. elongatum'' and ''A. cylindratum''.
|
|-
|
''[[Aneuroderma]]''<ref name=Aneuroderma>{{Cite journal|last1=Xiong |first1=S. |last2=Engel |first2=M. S. |last3=Xiao |first3=L. |last4=Ren |first4=D. |title=New archidermapteran earwigs (Dermaptera) from the Middle Jurassic of Inner Mongolia, China |year=2021 |journal=ZooKeys |issue=1065 |pages=125–139 |doi=10.3897/zookeys.1065.72720 |pmid=34759716 |pmc=8563706 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Xiong, Engel & Ren ''in'' Xiong ''et al.''
|
[[Middle Jurassic]]
|
[[Jiulongshan Formation]]
|
{{Flag|China}}
|
A member of the family [[Protodiplatyidae]]. The type species is ''A. oiodes''.
|
|-
|
''[[Dacryoderma]]''<ref>{{Cite journal|last=Engel |first=M. S. |title=A new genus of Early Jurassic earwigs from England (Dermaptera) |year=2021 |journal=Novitates Paleoentomologicae |volume=22 |issue=22 |pages=1–3 |doi=10.17161/np.22.15759 |s2cid=237950109 |doi-access=free }}</ref>
|
Gen. et comb. nov
|
Valid
|
[[Michael S. Engel|Engel]]
|
[[Early Jurassic]] ([[Sinemurian]])
|
[[Charmouth Mudstone Formation]]
|
{{Flag|United Kingdom}}
|
A [[Dermapteridae|dermapterid]] earwig.<br/>The type species is ''"Brevicula" teres'' Tihelka ([[2019 in paleoentomology|2019]]).
|
|-
|
''[[Eminepygia]]''<ref>{{Cite journal|last1=Chen |first1=X.-Y. |last2=Zhang |first2=H.-C. |title=''Eminepygia myanmarensis'' sp. nov., a new Earwig (Dermaptera) Nymph in Burmese Amber |year=2021 |journal=Zootaxa |volume=4952 |issue=3 |pages=571–579 |doi=10.11646/zootaxa.4952.3.9 |pmid=33903361 |s2cid=233411605 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Chen & Zhang
|
[[Cenomanian]]
|
[[Burmese amber]]
|
{{Flag|Myanmar}}
|
A [[Pygidicranidae|pygidicranid]] [[earwig]].<br/> Genus includes new species ''E. myanmarensis''.
|
|-
|
''[[Sinopalaeodermata|Sinopalaeodermata concavum]]''<ref name=Aneuroderma />
|
Sp. nov
|
Valid
|
Xiong, Engel & Ren ''in'' Xiong ''et al.''
|
Middle Jurassic
|
Jiulongshan Formation
|
{{Flag|China}}
|
A member of the family [[Dermapteridae]].
|
|-
|}

===Plecoptera===
====New Plecopteran taxa====

{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Balticonemoura]]''<ref>{{Cite journal|last=Chen |first=Z.-T. |year=2021 |title=''Balticonemoura bulbosus'' gen. et sp. nov., a new stonefly of Nemouridae (Insecta: Plecoptera) from Eocene Baltic amber |journal=Historical Biology |volume=34 |issue=3 |pages=421–424 |doi=10.1080/08912963.2021.1922399 |s2cid=235528609 }}</ref>
|
Gen. et sp. nov
|
In press
|
Chen
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea region)
|
A [[Nemouridae|nemourid]] [[Plecoptera|stonefly]].<br/> Genus includes new species ''B. bulbosus''. 
| 
|-
|
''[[Billoperla]]''<ref>{{Cite journal|last1=Chen |first1=Z.-T. |last2=Xu |first2=C.-P. |title=New perlid stonefly (Insecta: Plecoptera) from mid-Cretaceous Burmese amber |year=2021 |journal=Cretaceous Research |volume=127 |pages=Article 104954 |doi=10.1016/j.cretres.2021.104954 |bibcode=2021CrRes.12704954C }}</ref>
|
Gen. et sp. nov
|
In press
|
Chen & Xu
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Perlidae|perlid]] stonefly belonging to the family.<br/> Genus includes new species ''B. starki''.
|
|-
|
''[[Isoperla baltica]]''<ref>{{Cite journal|last1=Jouault |first1=C. |last2=Legendre |first2=F. |last3=Condamine |first3=F. L. |last4=Nel |first4=A. |title=A new stonefly species (Plecoptera: Perlodidae) from Eocene Baltic amber and questions on the wing venation potential for species diagnostic of fossil Plecoptera |year=2021 |journal=Palaeoentomology |volume=4 |issue=3 |pages=243–256 |doi=10.11646/palaeoentomology.4.3.12 |s2cid=237847715 |url=https://hal-insu.archives-ouvertes.fr/insu-03272797/file/Jouault_et_al_BA-Plecoptera.pdf }}</ref>
|
Sp. nov
|
Valid
|
Jouault ''et al.''
|
Eocene
|
Baltic amber
|
Europe (Baltic Sea regon)
|
A [[Perlodidae|perlodid]] stonefly.
|
|-
|}

==Other insects==
===New taxa===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Anakitala]]''<ref name=Anakitala>{{Cite journal|last=Aristov |first=D. S. |title=New eoblattid insects (Insecta: Eoblattida) from the Upper Permian of Russia |year=2021 |journal=Paleontological Journal |volume=55 |issue=1 |pages=72–75 |doi=10.1134/S0031030121010020 |s2cid=232070926 |url=https://www.elibrary.ru/item.asp?id=44492497 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Aristov
|
Late [[Permian]]
|
|
{{Flag|Russia}}
|
A [[Mesorthopteridae|mesorthopterid]] [[Eoblattida]]n [[Polyneoptera]]n.<br/> The type species is ''A. intermittendia''.
|
|-
|
''[[Baharellinus|Baharellinus elegans]]''<ref name=Anakitala />
|
Sp. nov
|
Valid
|
Aristov
|
Late Permian
|
|
{{Flag|Russia}}
|
A member of Polyneoptera belonging to the group Eoblattida and the family [[Blattogryllidae]].
|
|-
|
''[[Belmophenopterum|Belmophenopterum rasnitsyni]]''<ref>{{Cite journal|last1=Storozhenko |first1=S. Yu. |last2=Aristov |first2=D. S. |title=The latest record of the genus ''Belmophenopterum'' (Eoblattida: Mesorthopteridae) from the Middle Triassic of Kyrgyzstan |year=2021 |journal=Far Eastern Entomologist |volume=437 |pages=31–36 |doi=10.25221/fee.437.6 |s2cid=239678300 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Storozhenko & Aristov
|
[[Triassic]] ([[Ladinian]]-[[Carnian]])
|
[[Madygen Formation]]
|
{{Flag|Kyrgyzstan}}
|
A member of the family Mesorthopteridae.
|
|-
|
''[[Cabarzopterum]]''<ref name=Cabarzopterum>{{Cite journal|last1=Cui |first1=Y. |last2=Brauner |first2=S. |last3=Schneider |first3=J. W. |last4=Béthoux |first4=O. |title=Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities |year=2021 |journal=Journal of Paleontology |volume= 96|issue=2 |pages=355–374 |doi=10.1017/jpa.2021.101 |s2cid=243486826 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Cui ''et al.''
|
Early [[Permian]]
|
[[Goldlauter Formation]]
|
{{Flag|Germany}}
|
A member of [[Grylloblattida]] of uncertain phylogenetic placement. Genus includes new species ''C. magnificus''.
|
|-
|
''[[Delopterinus|Delopterinus alsenzensis]]''<ref name=NJGPA3003>{{Cite journal|last1=Nel |first1=A. |last2=Poschmann |first2=M. J. |title=Early Permian acercarian insects (Miomoptera, Palaeomanteidae) from the Saar-Nahe Basin (Rhineland-Palatinate, SW Germany) |year=2021 |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=300 |issue=3 |pages=267–277 |doi=10.1127/njgpa/2021/0989 |s2cid=236233140 }}</ref>
|
Sp. nov
|
Valid
|
Nel & Poschmann
|
Early [[Permian]]
|
[[Saar–Nahe Basin]]
|
{{Flag|Germany}}
|
A [[miomoptera]]n belonging to the family [[Palaeomanteidae]].
|
|-
|
''[[Delopterum|Delopterum novokshonovi]]''<ref name=NJGPA3003 />
|
Sp. nov
|
Valid
|
Nel & Poschmann
|
Early Permian
|
Saar–Nahe Basin
|
{{Flag|Germany}}
|
A miomopteran belonging to the family Palaeomanteidae.
|
|-
|
''[[Elmocladus]]''<ref>{{Cite journal|last1=Rasnitsyn |first1=A. P. |last2=Aristov |first2=D. S. |title=Review of the family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe |year=2021 |journal=Paleontological Journal |volume=55 |issue=2 |pages=151–156 |doi=10.1134/S003103012102012X |s2cid=233746323 |url=https://elibrary.ru/item.asp?id=44595942 }}</ref>
|
Gen. et comb. nov
|
Valid
|
Rasnitsyn & Aristov
|
Early [[Permian]]
|
|
{{Flag|United States}}
|
A member of Polyneoptera belonging to the group Eoblattida and the family [[Tococladidae]]. The type species is ''"[[Opisthocladus]]" strictus'' Carpenter (1976).
|
|-
|
''[[Galloembia]]''<ref>{{Cite journal |last1=Falières |first1=E. |last2=Engel |first2=M. S. |last3=Nel |first3=A. |title=Earliest occurrence of Embiidae: A new genus from earliest Eocene Oise amber (Insecta: Embiodea) |year=2021 |journal=Comptes Rendus Palevol |volume=20 |issue=38 |pages=799–805 |doi=10.5852/cr-palevol2021v20a38 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Falières, Engel & Nel
|
Earliest Eocene
|
Oise amber
|
{{Flag|France}}
|
A webspinner belonging to the family [[Embiidae]]. The type species is ''G. raholai''.
|
|-
|
''[[Gnethoda|Gnethoda odontophora]]''<ref name=Multivena>{{Cite journal|last1=Lai |first1=Z. |last2=Yang |first2=Q. |last3=Shi |first3=C. |last4=Zhang |first4=W. |last5=Ren |first5=D. |title=New webspinners (Insecta: Embioptera) from Upper Cretaceous amber of northern Myanmar |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105040 |doi=10.1016/j.cretres.2021.105040 |s2cid=239202878 }}</ref>
|
Sp. nov
|
In press
|
Lai ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A [[Embioptera|webspinner]] belonging to the family [[Clothodidae]].
|
|-
|
''[[Kamopanorpa|Kamopanorpa rasnitsyni]]''<ref name=Kamopanorpa>{{Cite journal|last1=Bashkuev |first1=A. S. |last2=Sukatsheva |first2=I. D. |title=New species of ''Kamopanorpa'' Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera) |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=453–461 |doi=10.11646/palaeoentomology.4.5.11 | issn=2624-2826  |s2cid=240525412 }}</ref>
|
Sp. nov
|
Valid
|
Bashkuev & Sukatsheva
|
Permian
|
Minusinsk Coal Basin
|
{{Flag|Russia}}
|
A member of the ([[Crown group#Stem groups|stem]]-[[trichoptera]]n or stem-[[amphiesmenoptera]]n) family [[Microptysmatidae]].
|
|-
|
''[[Kamopanorpa|Kamopanorpa rotunda]]''<ref name=Kamopanorpa />
|
Sp. nov
|
Valid
|
Bashkuev & Sukatsheva
|
Permian
|
Minusinsk Coal Basin
|
{{Flag|Russia}}
|
A member of the family Microptysmatidae.
|
|-
|
''[[Kamopanorpa|Kamopanorpa sivchikovi]]''<ref name=Kamopanorpa />
|
Sp. nov
|
Valid
|
Bashkuev & Sukatsheva
|
Permian
|
Minusinsk Coal Basin
|
{{Flag|Russia}}
|
A member of the family Microptysmatidae.
|
|-
|
''[[Liomopterites|Liomopterites sperbersbachensis]]''<ref name=Cabarzopterum />
|
Sp. nov
|
Valid
|
Cui ''et al.''
|
Early Permian
|
Goldlauter Formation
|
{{Flag|Germany}}
|
A member of Grylloblattida belonging to the family [[Liomopteridae]].
|
|-
|
''[[Liomopterum|Liomopterum fuscatum]]''<ref name=Cabarzopterum />
|
Sp. nov
|
Valid
|
Cui ''et al.''
|
Early Permian
|
Goldlauter Formation
|
{{Flag|Germany}}
|
A member of Grylloblattida belonging to the family Liomopteridae.
|
|-
|
''[[Madygembia]]''<ref>{{Cite journal|last1=Aristov |first1=D. S. |last2=Storozhenko |first2=S. Yu. |title=A new genus of the family Alexarasniidae (Insecta: Embiida) from the Triassic of Kyrgyzstan |year=2021 |journal=Palaeoentomology |volume=4 |issue=5 |pages=433–435 |doi=10.11646/palaeoentomology.4.5.7 |s2cid=240549301 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Aristov & Storozhenko
|
[[Triassic]] ([[Ladinian]] or [[Carnian]])
|
[[Madygen Formation]]
|
{{Flag|Kyrgyzstan}}
|
A webspinner belonging to the family [[Alexarasniidae]]. The type species is ''M. rasnitsyni''.
|
|-
|
''[[Magicivena]]''<ref>{{Cite journal|last1=Yang |first1=H. |last2=Shih |first2=C. |last3=Rasnitsyn |first3=A. P. |last4=Ren |first4=D. |last5=Gao |first5=T. |title=Early Cretaceous enigmatic insect group showing unique wing venations and antennal sensilla |year=2021 |journal=Papers in Palaeontology |volume= 8|doi=10.1002/spp2.1402 |s2cid=244172848 }}</ref>
|
Fam. et gen. et 3 sp. nov
|
Valid
|
Yang ''et al.''
|
Early Cretaceous
|
[[Yixian Formation]]
|
{{Flag|China}}
|
A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family [[Magicivenidae]]. The type species is ''M. sticta''; genus also includes ''M. elegans'' and ''M. antennalis''.
|
|-
|
''[[Multivena]]''<ref name=Multivena />
|
Gen. et sp. nov
|
In press
|
Lai ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
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A webspinner belonging to the family [[Sorellembiidae]]. Genus includes new species ''M. curvivena''.
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''[[Pictoborella]]''<ref name=Cabarzopterum />
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Gen. et sp. et comb. nov
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Valid
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Cui ''et al.''
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Early Permian
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{{Flag|Germany}}
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A member of Grylloblattida. Genus includes new species ''P. clara'', as well as ''"[[Oborella]]" germanica'' Prokop ''et al.'' (2021).
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''[[Rasnalexia]]''<ref>{{Cite journal|last=Gorochov |first=A. V. |title=New Family, Genus and Species of the Primitive Embioptera from the Triassic of Kyrgyzstan and Remarks on the Classification of this Order |year=2021 |journal=Entomological Review |volume=101 |issue=6 |pages=837–845 |doi=10.1134/S0013873821060087 |s2cid=245108349 }}</ref>
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Fam. et gen. et sp. nov
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Valid
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Gorochov
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[[Triassic]]
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{{Flag|Kyrgyzstan}}
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A member of [[Embioptera]] belonging to the new family [[Rasnalexiidae]]. Genus includes new species ''R. rasnitsyni''.
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''[[Uralioma|Uralioma thuringiensis]]''<ref name=Cabarzopterum />
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Sp. nov
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Valid
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Cui ''et al.''
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Early Permian
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Goldlauter Formation
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{{Flag|Germany}}
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A member of Grylloblattida belonging to the family Liomopteridae.
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===Other insect research===
* A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the [[Permian]] ([[Wordian]]) [[Pelitic Formation]] (France) by Logghe ''et al.'' (2021), who name a new [[ichnotaxon]] ''[[Phasmichnus]] radagasti''.<ref>{{Cite journal|last1=Logghe |first1=A. |last2=Nel |first2=A. |last3=Steyer |first3=J.-S. |last4=Ngô-Muller |first4=V. |last5=Pouillon |first5=J.-M. |last6=Garrouste |first6=R. |title=A twig-like insect stuck in the Permian mud indicates early origin of an ecological strategy in Hexapoda evolution |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 20774 |doi=10.1038/s41598-021-00110-2 |doi-access=free |pmid=34675246 |pmc=8531326 |bibcode=2021NatSR..1120774L }}</ref>
* A dragonfly-type [[odonatoptera]]n larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).<ref>{{Cite journal |last1=Haug |first1=J. T. |last2=Müller |first2=P. |last3=Haug |first3=C. |title=Fossil dragonfly-type larva with lateral abdominal protrusions and implications on the early evolution of Pterygota |year=2021 |journal=iScience |volume=24 |issue=10 |pages=Article 103162 |doi=10.1016/j.isci.2021.103162 |pmid=34646993 |pmc=8501664 |bibcode=2021iSci...24j3162H }}</ref>

==General research==
* Revision of the fossil record of insects from the [[Triassic]] of China is published by Zhang ''et al.'' (2021).<ref>{{cite book |last1=Zhang |first1=Q. |last2=Zheng |first2=D. |last3=Wang |first3=B. |last4=Zhang |first4=H. |year=2021 |chapter=Review of Triassic insects in China |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |series=Geological Society, London, Special Publications |volume=521 |pages=45–60 |publisher=The Geological Society of London |doi=10.1144/SP521-2021-121 |s2cid=244645537 }}</ref>
* A study on the diversity of insect damage types in fossil plants from the Cretaceous ([[Albian]] to [[Cenomanian]]) [[Dakota Formation]] (United States), evaluating their implications for the knowledge of the early evolution of angiosperm [[Florivore|florivory]] and associated pollination, is published by Xiao ''et al.'' (2021).<ref>{{cite journal |last1=Xiao |first1=L. |last2=Labandeira |first2=C. |last3=Dilcher |first3=D. |last4=Ren |first4=D. |year=2021 |title=Florivory of Early Cretaceous flowers by functionally diverse insects: implications for early angiosperm pollination |journal=Proceedings of the Royal Society B: Biological Sciences |volume=288 |issue=1953 |pages=Article ID 20210320 |doi=10.1098/rspb.2021.0320 |pmid=34132112 |pmc=8207559 }}</ref>

==References==
{{reflist|2}}

[[Category:2021 in paleontology]]
[[Category:Paleoentomology]]
[[Category:2021-related lists]]