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#REDIRECT [[Laughingthrushes]] |
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{{Taxobox |
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| name = Laughingthrush |
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| image = Masked Laughingthrush.jpg |
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| image_width = 250px |
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| image_caption = [[Masked Laughingthrush]] <br/>''Garrulax perspicillatus'' |
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| regnum = [[Animal]]ia |
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| phylum = [[Chordata]] |
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| classis = [[Aves]] |
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| ordo = [[Passeriformes]] |
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| familia = [[Timaliidae]] |
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| genus = '''''Garrulax''''' |
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| genus_authority = [[René Primevère Lesson|Lesson]], 1831 |
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| subdivision_ranks = Species |
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| subdivision = |
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Presently about 50, but see text |
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}} |
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{{R from misspelling}} |
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The '''Laughingthrushes''' are the genus '''''Garrulax''''' of the large [[Old World babbler]] family of [[passerine]] [[bird]]s. They occur in tropical [[Asia]], with the greatest number of species occurring in the [[Himalaya]] and southern [[China]]. |
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These are rangy, medium-sized, floppy-tailed landbirds with soft fluffy plumage. These birds have strong legs and are quite terrestrial. This group is not strongly [[bird migration|migratory]], and most [[species]] have short rounded wings, and a weak flight. |
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A few, like the [[Streaked Laughingthrush]] occur in fairly open habitats, but most are jungle species, difficult to observe in the dense vegetation they prefer. |
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Like other babblers, these are noisy birds, and the characteristic laughing calls are often the best indication that these birds are present. As with other babbler species, they frequently occur in groups of up to a dozen, and the rainforest species like the [[Ashy-headed Laughingthrush]] often occur in the mixed feeding flocks typical of tropical Asian jungle. |
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== Systematics == |
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As with some other babblers, this genus has turned out to be highly [[polyphyletic]]. Rather than forming a natural [[evolution]]ary group of closest relatives, the laughingthrushes are spread about the entire babbler family. This is quite apparent by their diverse [[morphology (biology)|morphology]] and presence of several groups with distinct color patterns, but the rarity of several species and the sheer number of [[taxa]] have hitherto prevented a thorough revision. Before this paraphyly was realized to be as dramatic as it is, it was generally assumed that the laughingthrushes were a [[clade]] comprising several [[subgenus|subgenera]]. |
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This has much confounded further research: most [[molecular phylogeny]] studies, for example, used only the [[White-crested Laughingthrush]] as an "example" of this "genus", because specimens were readily available. Morphological studies, on the other hand, had usually more material at their disposal, but these too were generally conducted under a false assumption of monophyly. A 2003 study,<ref>Cibois (2003)</ref> analyzing [[mtDNA]] [[cytochrome b|cytochrome ''b'']] and [[12S rRNA|12S]]/[[MT-RNR2|16S rRNA]] data of a number of laughingthrushes, did establish however that there is no such thing as a "representative" laughingthrush. Consequently most of the work regarding the evolution of this assemblage needs to be evaluated anew and if necessary even redone. Indeed, the taxon ''Garrulax'' would need to be restricted to the [[type species]] ([[Rufous-fronted Laughingthrush]]) and its closest relatives, but the species generally considered the most likely candidates have not been comprehensively studied in recent times.<ref>Cibois (2003), Pasquet ''et al.'' (2006)</ref> |
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=== Species list === |
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The list below uses the treatment of the laughingthrushes by ''[[Birds of South Asia. The Ripley Guide|Birds of South Asia]]''<ref>Rasmussen & Anderton (2005)</ref> and the ''[[Handbook of Birds of the World]]''<ref>Collar & Robson (2007)</ref> as a baseline, acknowledging the polyphyly of ''Garrulax'' and attempting to reorganise it into [[monophyletic]] genera. |
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This list is by no means the last word on the subject though, and it is very likely that some of the new groups will be revised, expanded or split as more species are studied in the future; additional laughingthrushes are presently elevated to species status and even hitherto unknown taxa are discovered at a rate of about 1-2 per year. Some proposed small or [[monotypic]] genera on the other hand might ultimately be merged with others for convenience if they turn out to be [[sister taxa]]. All in all, the resolution of the laughingthrush assemblage's [[taxonomy]] depends on a resolution of the Old World babblers' [[systematics]] in general. |
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[[File:A03 2880 640x427.jpg|thumb|right|[[White-crested Laughingthrush]]<br/>''Garrulax leucolophus'']] |
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* '''Genus ''[[Garrulax]]'' - |
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** [[Chinese Hwamei]], ''Garrulax canorus'' |
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** [[Taiwan Hwamei]], ''Garrulax taewanus'' |
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** [[White-necked Laughingthrush]], ''Garrulax strepitans'' |
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** [[White-crested Laughingthrush]], ''Garrulax leucolophus'' |
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** [[Sumatran Laughingthrush]], ''Garrulax bicolor'' |
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** [[Lesser Necklaced Laughingthrush]], ''Garrulax monileger'' |
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** [[Greater Necklaced Laughingthrush]], ''Garrulax pectoralis'' |
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** [[White-throated Laughingthrush]], ''Garrulax albogularis'' |
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** [[Rufous-crowned Laughingthrush]], ''Garrulax ruficeps'' |
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** [[Masked Laughingthrush]], ''Garrulax perspicillatus'' |
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** [[Rufous-fronted Laughingthrush]], ''Garrulax rufifrons'' |
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** [[Ashy-headed Laughingthrush]], ''Garrulax cinereifrons'' |
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** [[Sunda Laughingthrush]], ''Garrulax palliatus'' |
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** [[Grey Laughingthrush]], ''Garrulax maesi'' |
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** [[Rufous-cheeked Laughingthrush]], ''Garrulax castanotis'' |
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** [[Black-hooded Laughingthrush]], ''Garrulax milleti'' |
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*** Kon Tum Black-hooded Laughingthrush, ''Garrulax milleti sweeti'' |
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** [[Cambodian Laughingthrush]], ''Garrulax ferrarius'' |
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** [[Snowy-cheeked Laughingthrush]], ''Garrulax sukatschewi'' |
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** [[Moustached Laughingthrush]], ''Garrulax cineracea'' |
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** [[Rufous-chinned Laughingthrush]], ''Garrulax rufogularis'' |
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** [[Chestnut-eared Laughingthrush]], ''Garrulax konkakinhensis'' |
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** [[Barred Laughingthrush]], ''Garrulax lunulata'' |
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** [[White-speckled Laughingthrush]], ''Garrulax bieti'' |
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** [[Giant Laughingthrush]], ''Garrulax maxima'' |
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** [[Spotted Laughingthrush]], ''Garrulax ocellata'' |
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* '''Proposed genus ''[[Dryonastes]]''''' |
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** [[Grey-sided Laughingthrush]], ''Dryonastes caerulatus'' |
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** [[Rusty Laughingthrush]], ''Dryonastes poecilorhynchus'' |
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** [[Buffy Laughingthrush]], ''Dryonastes berthemyi'' - formerly in ''D. poecilorhynchus'' |
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** [[Black-throated Laughingthrush]], ''Dryonastes chinensis'' |
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** [[Chestnut-backed Laughingthrush]], ''Dryonastes nuchalis'' |
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** [[Rufous-vented Laughingthrush]], ''Dryonastes gularis'' |
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** [[Wynaad Laughingthrush]], ''Dryonastes delesserti'' |
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** [[Blue-crowned Laughingthrush]] or Courtois's Laughingbird, ''Dryonastes courtoisi'' - formerly in ''D. galbanus'' |
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** [[Yellow-throated Laughingthrush]], ''Dryonastes galbanus'' |
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** [[White-cheeked Laughingthrush]], ''Dryonastes vassali''[[File:Chestnut-capped Laughing Thrush.jpg|thumb|right|[[Spectacled Laughingthrush]]<br/>''Garrulax mitratus/Rhinocichla mitrata'']] |
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** [[Rufous-necked Laughingthrush]], ''Dryonastes ruficollis'' |
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* '''Proposed genus ''[[Melanocichla]]''''' |
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** [[Black Laughingthrush]], ''Melanocichla lugubris'' |
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** [[Bare-headed Laughingthrush]], ''Melanocichla calva'' - formerly in ''M. lugubris'' |
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* '''Proposed genus ''[[Rhinocichla]]''''' - [[Chestnut-capped Laughingthrush]]es |
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** [[Spectacled Laughingthrush]], ''Rhinocichla mitrata'' |
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** [[Chestnut-hooded Laughingthrush]], ''Rhinocichla treacheri'' - formerly in ''R. mitrata'' |
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* '''Proposed genus ''[[Grammatoptila]]''''' |
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** [[Striated Laughingthrush]], ''Grammatoptila striata'' |
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[[File:New Friend.jpg|thumb|right|[[Chinese Hwamei]]<br/>''Garrulax canorus/Leucodioptron canorum'']] |
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* '''Proposed genus ''[[Stactocichla]]''''' |
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** [[Spot-breasted Laughingthrush]], ''Stactocichla merulina'' |
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** [[Orange-breasted Laughingthrush]], ''Stactocichla annamensis'' - formerly in ''S. merulina'' |
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[[File:Streaked Laughingthrush I IMG 3892.jpg|thumb|right|[[Streaked Laughingthrush]]<br/>''Garrulax lineatus/Strophocincla lineata'']] |
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* '''Proposed genus ''[[Strophocincla]]''''' |
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** [[Black-chinned Laughingthrush]], Rufous-breasted Laughingthrush or Nilgiri Laughingthrush, ''Strophocincla cachinnans'' |
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** [[Kerala Laughingthrush]] or Grey-breasted Laughingthrush, ''Strophocincla fairbanki'' |
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** [[Bhutan Laughingthrush]], ''Strophocincla imbricata'' - formerly in ''S. lineata'' |
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** [[Streaked Laughingthrush]], ''Strophocincla lineata'' |
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** [[Striped Laughingthrush]], ''Strophocincla virgata'' |
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* '''Proposed genus ''[[Pterorhinus]]''''' |
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** [[White-browed Laughingthrush]], ''Pterorhinus sannio'' |
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** [[Plain Laughingthrush]], Pere David's Laughingthrush or Plain Laughingbird, ''Pterorhinus davidi'' |
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* '''''incertae sedis - Ianthocincla'' or ''Trochalopteron''?''' |
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** [[Brown-capped Laughingthrush]],''Trochalopteron/Ianthocincla austeni'' |
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[[File:Garrulax formosus - front-6.jpg|thumb|right|[[Red-winged Laughingthrush]]<br/>''Garrulax formosus/Trochalopteron formosum'']] |
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* '''Proposed genus ''[[Trochalopteron]]''''' |
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** [[Scaly Laughingthrush]], ''Trochalopteron subunicolor'' |
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** [[Blue-winged Laughingthrush]], ''Trochalopteron squamatum'' |
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** [[Brown-cheeked Laughingthrush]] or Prince Henry's Laughingthrush, ''Trochalopteron henrici'' |
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** [[Elliot's Laughingthrush]], ''Trochalopteron elliotii'' |
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** [[Variegated Laughingthrush]], ''Trochalopteron variegatum'' |
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** [[White-whiskered Laughingthrush]], ''Trochalopteron morrisonianum'' |
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** [[Black-faced Laughingthrush]], ''Trochalopteron affine'' |
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** [[Chestnut-crowned Laughingthrush]], ''Trochalopteron erythrocephalum'' |
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** [[Assam Laughingthrush]], ''Trochalopteron chrysopterum'' - formerly in ''T. erythrocephalum'' |
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** [[Silver-eared Laughingthrush]], ''Trochalopteron melanostigma'' - formerly in ''T. erythrocephalum'' |
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** [[Malayan Laughingthrush]], ''Trochalopteron peninsulae'' - formerly in ''T. erythrocephalum'' |
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** [[Golden-winged Laughingthrush]], ''Trochalopteron ngoclinhense'' |
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** [[Collared Laughingthrush]], ''Trochalopteron yersini'' |
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** [[Red-winged Laughingthrush]], ''Trochalopteron formosum'' |
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** [[Red-tailed Laughingthrush]], ''Trochalopteron milnei'' |
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== References == |
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<references/> |
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* Cibois, Alice (2003a): Mitochondrial DNA Phylogeny of Babblers (Timaliidae). ''[[Auk (journal)|Auk]]'' '''120'''(1): 1-20. <small>[[Digital Object Identifier|DOI]]: 10.1642/0004-8038(2003)120[0035:MDPOBT]2.0.CO;2</small> [http://findarticles.com/p/articles/mi_qa3793/is_200301/ai_n9225604 HTML fulltext without images] |
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* Collar, N.J. & Robson, Craig (2007): Family Timaliidae (Babblers). ''In:'' del Hoyo, Josep; Elliott, Andrew & Christie, D.A. (eds.): ''[[Handbook of Birds of the World]], Volume 12'' (Picathartes to Tits and Chickadees): 70-291. Lynx Edicions, Barcelona. |
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* Pasquet, Eric; Bourdon, Estelle; Kalyakin, Mikhail V. & Cibois, Alice (2006). The fulvettas (''Alcippe''), Timaliidae, Aves): a polyphyletic group. ''[[Zoologica Scripta]]'' '''35''', 559–566. <small>{{doi|10.1111/j.1463-6409.2006.00253.x}}</small> (HTML abstract) |
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* [[Pamela Rasmussen|Rasmussen, Pamela C.]] & Anderton, J.C. (2005): ''[[Birds of South Asia. The Ripley Guide]]''. Smithsonian Institution and Lynx Edicions, Washington DC and Barcelona. |
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== External links == |
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{{commons category|Garrulax}} |
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*[http://ibc.lynxeds.com/family/babblers-timaliidae Laughingthrush videos] on the Internet Bird Collection |
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[[Category:Garrulax|*]] |
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[[Category:Laughingthrushes]] |
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[[ca:Garrulax]] |
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[[fr:Garrulax]] |
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[[hu:Garrulax]] |
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[[nl:Garrulax]] |
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[[ja:ガビチョウ亜科]] |
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[[pl:Garrulax]] |
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[[pt:Garrulax]] |
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[[ru:Кустарницы]] |
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[[zh:噪鹛属]] |
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Latest revision as of 21:22, 14 May 2023
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