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== Article Draft ==
== Article Draft ==
{{Short description|Species of butterfly}}
{{Speciesbox
|image=Celastrina_echo-dorsal.jpg
|image_caption=''C. echo cinerea'' upperside
|image2=Celastrina_echo-ventral.jpg
|image2_caption=''C. echo cinerea'' underside
| genus = Celastrina
| species = echo
| authority = (W. H. Edwards, 1864)
}}


'''''Celastrina echo''''', known generally as the '''echo azure''' or '''western azure''', is a species of [[Polyommatinae|blue]] in the [[butterfly]] family [[Lycaenidae]].<ref name=itis/><ref name=buglink/>
⚫
=== Lead ===
⚫
Celastrina Echo, also known as Western Azure, is a specie of blue butterfly under the family Lycaenidae. It is found in North America regions, originated in Canada. Celastrina echo have been observed in mostly western region of United States, including California, Oregon, Arizona, New Mexico, Montana<ref>{{Cite web |last=Webmaster |first=David Ratz |title=Western Azure - Montana Field Guide |url=https://fieldguide.mt.gov/speciesDetail.aspx?elcode=IILEPG0080 |access-date=2024-05-30 |website=fieldguide.mt.gov |language=en}}</ref>.


The MONA or Hodges number for ''Celastrina echo'' is 4363.2.<ref name=mpg/>
⚫
The taxonomic division of Celastrina Butterflies has been a very complicated issue. Some scientists like Layberry et al once recognized ''C. echo'' (Edwards 1864) as a subspecie of ''C.ladon'' (Cramer, 1780) in western Canada<ref name=":0">{{Cite journal |last=Schmidt |first=B. Christian |last2=Layberry |first2=Ross A. |date=2016-04-26 |title=What Azure blues occur in Canada? A re-assessment of Celastrina Tutt species (Lepidoptera, Lycaenidae) |url=https://zookeys.pensoft.net/article/7882/ |journal=ZooKeys |language=en |volume=584 |pages=135–164 |doi=10.3897/zookeys.584.7882 |issn=1313-2970}}</ref>. At 2001, Guppy & Shepard nominated the ''C. echo'' to the species level<ref>{{Cite journal |last=LaBar, Pelham, Kondla |first=Caitlin, Jonathan, Norbert |date=28 April 2022 |title=A new species of Celastrina from the northwestern United States and
[[File:Celastrina_echo-cinerea_puddling.jpg|thumb|[[mud-puddling]]]]

==Subspecies==
These four subspecies belong to the species ''Celastrina echo'':
* ''Celastrina echo cinerea'' <small>(W. H. Edwards, 1883)</small> (southwestern azure)
* ''Celastrina echo echo'' <small>(W. H. Edwards, 1864)</small> (Pacific azure)
* ''Celastrina echo nigrescens'' <small>(J. Fletcher, 1903)</small> (northwestern azure)
* ''Celastrina echo sidara'' <small>(Clench, 1944)</small> (Rocky Mountain azure)

==References==
{{Reflist|refs=
<ref name=itis>
{{Cite web| title=''Celastrina echo'' Report
| url=https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=777892
| website=Integrated Taxonomic Information System
| access-date=2019-09-24
}}</ref>

<ref name=mpg>
{{Cite web| title=North American Moth Photographers Group, ''Celastrina echo''
| url=http://mothphotographersgroup.msstate.edu/species.php?hodges=4363.2
| access-date=2019-09-24
}}</ref>

<ref name=buglink>
{{Cite web| title=''Celastrina echo'' species Information
| url=https://bugguide.net/node/view/41480
| website=BugGuide.net
| access-date=2019-09-24
}}</ref>
}}

==Further reading==
{{refbegin}}
* {{Cite report
| title = Annotated taxonomic checklist of the Lepidoptera of North America, North of Mexico
| date = 2016
| last1 = Pohl | first1 = Greg
| last2 = Patterson | first2 = Bob
| last3 = Pelham | first3 = Jonathan
| url = https://www.researchgate.net/publication/302570819
| doi = 10.13140/RG.2.1.2186.3287| doi-access = free
}}
{{refend}}

==External links==
{{refbegin}}
* {{Commons category-inline|Celastrina echo}}
{{refend}}

{{Taxonbar|from=Q15779554}}


⚫
=== General ===
⚫
Celastrina Echo (Edwards 1864), also known as Western Azure, is a specie of blue butterfly under the family Lycaenidae. It is found in North America regions, originated in Canada. Celastrina echo have been observed in mostly western region of United States, including California, Oregon, Arizona, New Mexico, Montana<ref>{{Cite web |last=Webmaster |first=David Ratz |title=Western Azure - Montana Field Guide |url=https://fieldguide.mt.gov/speciesDetail.aspx?elcode=IILEPG0080 |access-date=2024-05-30 |website=fieldguide.mt.gov |language=en}}</ref>.

⚫
The taxonomic division of Celastrina Butterflies has been a very complicated issue. Some scientists like Layberry et al once recognized ''C. echo'' as a subspecie of ''C.ladon'' (Cramer, 1780) in western Canada<ref name=":0">{{Cite journal |last=Schmidt |first=B. Christian |last2=Layberry |first2=Ross A. |date=2016-04-26 |title=What Azure blues occur in Canada? A re-assessment of Celastrina Tutt species (Lepidoptera, Lycaenidae) |url=https://zookeys.pensoft.net/article/7882/ |journal=ZooKeys |language=en |volume=584 |pages=135–164 |doi=10.3897/zookeys.584.7882 |issn=1313-2970}}</ref>. At 2001, Guppy & Shepard nominated the ''C. echo'' to the species level<ref>{{Cite journal |last=LaBar |first=Caitlin |last2=Pelham |first2=Jonathan |last3=Kondla |first3=Norbert |date=28 April 2022 |title=A new species of Celastrina from the northwestern United States and
southwestern Canada with a lectotype designation of Lycaena
southwestern Canada with a lectotype designation of Lycaena
pseudargiolus var. nigrescens Fletcher (Lycaenidae: Polyommatinae) |url=https://lepsurvey.carolinanature.com/report.html |journal=The Taxonomic Report |volume=10}}</ref>. Until now, many scientists agreed to put ''C.echo'' as a distinguished specie<ref name=":0" />.
pseudargiolus var. nigrescens Fletcher (Lycaenidae: Polyommatinae) |url=https://lepsurvey.carolinanature.com/ttr/ttr-10-3.pdf |journal=The Taxonomic Report |volume=10 |issue=3 |pages=1-24}}</ref>. Until now, many scientists agreed to put ''C.echo'' as a distinguished specie<ref name=":0" />.


=== Article body ===
=== Article body ===

==== Physical Identification and Relationship with Ants ====
The butterflies under the ''Lycaenidae'' family in general are usually flat as larvae. The adult individuals are usually small and have hairy, ringed antenna-like tails. Their wings are mostly bright glowing blue and green. A large portion of the ''Lycaenidae'' butterflies were found to have different associations with ants, mutualistic, parasitic, or predatory varied by specific species. <ref>{{Cite journal |last=Robbins |first=Robert K. |date=1981-11 |title=The "False Head" Hypothesis: Predation and Wing Pattern Variation of Lycaenid Butterflies |url=https://www.journals.uchicago.edu/doi/10.1086/283868 |journal=The American Naturalist |language=en |volume=118 |issue=5 |pages=770–775 |doi=10.1086/283868 |issn=0003-0147}}</ref><ref>{{Cite journal |last=Ueda |first=Shouhei |last2=Komatsu |first2=Takashi |last3=Itino |first3=Takao |last4=Arai |first4=Ryusuke |last5=Sakamoto |first5=Hironori |date=2016-11-03 |title=Host-ant specificity of endangered large blue butterflies (Phengaris spp., Lepidoptera: Lycaenidae) in Japan |url=https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5093462/ |journal=Scientific Reports |volume=6 |pages=36364 |doi=10.1038/srep36364 |issn=2045-2322 |pmc=5093462 |pmid=27808223}}</ref><ref name=":1">{{Cite journal |last=Pierce |first=Naomi E. |last2=Braby |first2=Michael F. |last3=Heath |first3=Alan |last4=Lohman |first4=David J. |last5=Mathew |first5=John |last6=Rand |first6=Douglas B. |last7=Travassos |first7=Mark A. |date=2002-01 |title=The Ecology and Evolution of Ant Association in the Lycaenidae (Lepidoptera) |url=https://www.annualreviews.org/doi/10.1146/annurev.ento.47.091201.145257 |journal=Annual Review of Entomology |language=en |volume=47 |issue=1 |pages=733–771 |doi=10.1146/annurev.ento.47.091201.145257 |issn=0066-4170}}</ref>

''C. echo'' in particular is usually in pale blue-grey with small black spots or dashes. Based on the limited observations reported and the life history records of the other blue butterfly (Polyommatini), ''C. echo''<nowiki/>'s known life history starts with the eggs laid on the flower buds of its host plants. Attended and protected by the ants, ''C. echo''<nowiki/>'s larvae feed on the flower buds and flowers while ants receive sugar-rich honeydew from them throughout the larval lifespan. ''C. echo'' then may pupate within ant nests <ref name=":2">{{Cite web |title=Echo Azure Celastrina echo (W.H. Edwards, 1864) {{!}} Butterflies and Moths of North America |url=https://www.butterfliesandmoths.org/species/Celastrina-echo |access-date=2024-06-10 |website=www.butterfliesandmoths.org}}</ref>. Some ''Lycaenidae'' species may become parasites or even predator within the ant nest. The specific relationships to ants in the later life stage of C. echo are not quite clear. <ref name=":1" /> <ref name=":2" />

==== Host Plants, Habitat, and Flight Period: ====
The ''C. echo'' larval foodplants are mainly composed of ''Ceanothus'' (California wild lilac). ''Spiraea'' (Holodiscus), ''Aesculus'' (California buckeye), ''Rubus'' (blackberries), and some legumes. <ref>{{Cite book |last=Orsak |first=Larry J. |title=The butterflies of Orange County, California |date=1977 |publisher=Center for Pathobiology, University of California, Irvine |isbn=978-0-9601418-1-4 |series=Research series - Museum of Systematic Biology ; no. 4 |location=Irvine}}</ref><ref name=":2" /> The adults' host plants are also very diverse. They majorly feed on slower nectars coming from plants including the larval host plants listed above, ''Heteromeles'' (Toyon), ''Rhamnus'' (cascara), and several others. <ref name=":3">{{Cite web |last=Webmaster |first=David Ratz |title=Western Azure - Montana Field Guide |url=https://fieldguide.mt.gov/speciesDetail.aspx?elcode=IILEPG0080 |access-date=2024-06-10 |website=fieldguide.mt.gov |language=en}}</ref> <ref name=":2" />

''C. echo'' were often observed in woodlands, shrublands, and near mountain streams: places that have woody host plants they feed on.<ref name=":3" /> Depending on the moisture and relative condition of the larval hostplants, ''C. echo'' may fly in 1-2 or maybe more broods from early spring to the fall (around February to late June, July, or even till October if there is no severe cold weather). <ref name=":2" /><ref name=":3" />


==== Subspecies ====
==== Subspecies ====
Line 22: Line 98:


=== References ===
=== References ===
[[Category:Wikipedia Student Program]]

Latest revision as of 06:44, 10 June 2024

Article Draft

[edit]

Xinyuy22/Celastrina echo
C. echo cinerea upperside
C. echo cinerea underside
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Lycaenidae
Genus: Celastrina
Species:
C. echo
Binomial name
Celastrina echo
(W. H. Edwards, 1864)

Celastrina echo, known generally as the echo azure or western azure, is a species of blue in the butterfly family Lycaenidae.[1][2]

The MONA or Hodges number for Celastrina echo is 4363.2.[3]

mud-puddling

Subspecies

[edit]

These four subspecies belong to the species Celastrina echo:

  • Celastrina echo cinerea (W. H. Edwards, 1883) (southwestern azure)
  • Celastrina echo echo (W. H. Edwards, 1864) (Pacific azure)
  • Celastrina echo nigrescens (J. Fletcher, 1903) (northwestern azure)
  • Celastrina echo sidara (Clench, 1944) (Rocky Mountain azure)

References

[edit]
  1. ^ "Celastrina echo Report". Integrated Taxonomic Information System. Retrieved 2019-09-24.
  2. ^ "North American Moth Photographers Group, Celastrina echo". Retrieved 2019-09-24.

Further reading

[edit]
[edit]


General

[edit]

Celastrina Echo (Edwards 1864), also known as Western Azure, is a specie of blue butterfly under the family Lycaenidae. It is found in North America regions, originated in Canada. Celastrina echo have been observed in mostly western region of United States, including California, Oregon, Arizona, New Mexico, Montana[1].

The taxonomic division of Celastrina Butterflies has been a very complicated issue. Some scientists like Layberry et al once recognized C. echo as a subspecie of C.ladon (Cramer, 1780) in western Canada[2]. At 2001, Guppy & Shepard nominated the C. echo to the species level[3]. Until now, many scientists agreed to put C.echo as a distinguished specie[2].

Article body

[edit]

Physical Identification and Relationship with Ants

[edit]

The butterflies under the Lycaenidae family in general are usually flat as larvae. The adult individuals are usually small and have hairy, ringed antenna-like tails. Their wings are mostly bright glowing blue and green. A large portion of the Lycaenidae butterflies were found to have different associations with ants, mutualistic, parasitic, or predatory varied by specific species. [4][5][6]

C. echo in particular is usually in pale blue-grey with small black spots or dashes. Based on the limited observations reported and the life history records of the other blue butterfly (Polyommatini), C. echo's known life history starts with the eggs laid on the flower buds of its host plants. Attended and protected by the ants, C. echo's larvae feed on the flower buds and flowers while ants receive sugar-rich honeydew from them throughout the larval lifespan. C. echo then may pupate within ant nests [7]. Some Lycaenidae species may become parasites or even predator within the ant nest. The specific relationships to ants in the later life stage of C. echo are not quite clear. [6] [7]

Host Plants, Habitat, and Flight Period:

[edit]

The C. echo larval foodplants are mainly composed of Ceanothus (California wild lilac). Spiraea (Holodiscus), Aesculus (California buckeye), Rubus (blackberries), and some legumes. [8][7] The adults' host plants are also very diverse. They majorly feed on slower nectars coming from plants including the larval host plants listed above, Heteromeles (Toyon), Rhamnus (cascara), and several others. [9] [7]

C. echo were often observed in woodlands, shrublands, and near mountain streams: places that have woody host plants they feed on.[9] Depending on the moisture and relative condition of the larval hostplants, C. echo may fly in 1-2 or maybe more broods from early spring to the fall (around February to late June, July, or even till October if there is no severe cold weather). [7][9]

Subspecies

[edit]

There are five recorded subspecies belong to the species Celastrina echo[10]:

  • Celastrina echo echo (W. H. Edwards, 1864), also known as Pacific Azure;
  • Celastrina echo gozora (Boisduval, 1870), also known as Mexican Azure[11];
  • Celastrina echo cinerea (W. H. Edwards, 1883), also known as Southwestern Azure;
  • Celastrina echo nigrescens (J. Fletcher, 1903), also known as Northwestern Azure;
  • Celastrina echo sidara (Clench, 1944), also known as Rocky Mountain Azure;

References

[edit]
  1. ^ Webmaster, David Ratz. "Western Azure - Montana Field Guide". fieldguide.mt.gov. Retrieved 2024-05-30.
  2. ^ a b Schmidt, B. Christian; Layberry, Ross A. (2016-04-26). "What Azure blues occur in Canada? A re-assessment of Celastrina Tutt species (Lepidoptera, Lycaenidae)". ZooKeys. 584: 135–164. doi:10.3897/zookeys.584.7882. ISSN 1313-2970.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  3. ^ LaBar, Caitlin; Pelham, Jonathan; Kondla, Norbert (28 April 2022). "A new species of Celastrina from the northwestern United States and southwestern Canada with a lectotype designation of Lycaena pseudargiolus var. nigrescens Fletcher (Lycaenidae: Polyommatinae)" (PDF). The Taxonomic Report. 10 (3): 1–24. {{cite journal}}: line feed character in |title= at position 68 (help)
  4. ^ Robbins, Robert K. (1981-11). "The "False Head" Hypothesis: Predation and Wing Pattern Variation of Lycaenid Butterflies". The American Naturalist. 118 (5): 770–775. doi:10.1086/283868. ISSN 0003-0147. {{cite journal}}: Check date values in: |date= (help)
  5. ^ Ueda, Shouhei; Komatsu, Takashi; Itino, Takao; Arai, Ryusuke; Sakamoto, Hironori (2016-11-03). "Host-ant specificity of endangered large blue butterflies (Phengaris spp., Lepidoptera: Lycaenidae) in Japan". Scientific Reports. 6: 36364. doi:10.1038/srep36364. ISSN 2045-2322. PMC 5093462. PMID 27808223.
  6. ^ a b Pierce, Naomi E.; Braby, Michael F.; Heath, Alan; Lohman, David J.; Mathew, John; Rand, Douglas B.; Travassos, Mark A. (2002-01). "The Ecology and Evolution of Ant Association in the Lycaenidae (Lepidoptera)". Annual Review of Entomology. 47 (1): 733–771. doi:10.1146/annurev.ento.47.091201.145257. ISSN 0066-4170. {{cite journal}}: Check date values in: |date= (help)
  7. ^ a b c d e "Echo Azure Celastrina echo (W.H. Edwards, 1864) | Butterflies and Moths of North America". www.butterfliesandmoths.org. Retrieved 2024-06-10.
  8. ^ Orsak, Larry J. (1977). The butterflies of Orange County, California. Research series - Museum of Systematic Biology ; no. 4. Irvine: Center for Pathobiology, University of California, Irvine. ISBN 978-0-9601418-1-4.
  9. ^ a b c Webmaster, David Ratz. "Western Azure - Montana Field Guide". fieldguide.mt.gov. Retrieved 2024-06-10.
  10. ^ "ITIS - Report: Celastrina echo". www.itis.gov. Retrieved 2024-05-30.
  11. ^ "Celastrina echo thumbnails". www.butterfliesofamerica.com. Retrieved 2024-05-30.
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