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The Proto-Uralic homeland is the earliest location in which the Proto-Uralic language was spoken, before its speakers dispersed geographically causing it to diverge into multiple languages. Various locations have been proposed and debated, although as of 2022 "scholarly consensus now gravitates towards a relatively recent provenance of the Uralic languages east of the Ural mountains".^[1]

Homeland hypotheses

Europe versus Siberia

It has been suggested that the Proto-Uralic homeland was located near the Ural Mountains, either on the European or the Siberian side. The main reason to suppose that there was a Siberian homeland has been the traditional taxonomic model that sees the Samoyedic branch as splitting off first. Because the present border between the Samoyedic and the Ugric branch is in Western Siberia, the original split was seen to have occurred there too.

However, a European homeland would be equally possible because the Ugric languages are known to have been spoken earlier on the European side of the Urals. In recent years, it has also been argued based on phonology that the oldest split was not between the Samoyedic and the Finno-Ugric but between the Finno-Permic and the Ugro-Samoyedic language groups.^[2] The lexical level is argued to be less reliable, and lexical innovativeness (a small number of shared cognates) can be confused because of the great age of the division. No new arguments for a Siberian homeland have been presented for a long time.

Both European and Siberian homeland proposals have been supported by palaeolinguistic evidence, but only those cases in which the semantic reconstructions are certain and valid. A Siberian homeland has been claimed based on two coniferous tree names in Proto-Uralic, but the trees (Abies sibirica and Pinus sibirica)^[3] have for a long time been present also in the far east of Europe. A European homeland is supported by words for 'bee', 'honey', 'elm' etc.^[4] They can be reconstructed already to Proto-Uralic, if Samoyedic is no longer seen as the first branch to split off.^[5]

More recently, loanword evidence has also been used to support a European homeland. Proto-Uralic has been seen as borrowing words from Proto-Indo-European,^[6]^[7] and the Proto-Indo-European homeland has rarely been located east of the Urals. Proto-Uralic even seems to have developed in close contact with Proto-Indo-Iranian,^[8] which is seen as having arisen in the Poltavka culture of the Caspian steppes before its spread to Asia.^[9]

Bryant et al. (2005) equated the Lyalovo culture [fi; ru] (ca. 5000–3650 BC) with the Proto-Uralic urheimat, and the following Volosovo culture (ca. 3650–1900 BC) with the Proto-Finno-Ugric urheimat.^[10] Two Finnish scholars believe that the culture of Lyalovo was the Proto-Uralic urheimat and that its inhabitants spread Uralic languages to north-eastern Europe.^[11] The Volosovo culture has been named the Bronze Age Successor Culture, a textile-ceramic culture that developed in the region between Upper Volga and lakes Ladoga and Onega. It was distinguished from other groups based on the traces of textile used for the production of ceramics, and spread southeast all the way to central Volga, south to the entire river valley of the Oka, southwest to the northern shore of the Daugava, and northwest of Fennoscandia to Karelia, Finland and northern Sweden and Norway.^[12] Known as the Seima-Turbino phenomenon, it was a culturally unified, extensive network of trade in copper and bronze. The traces of the Seima-Turbino phenomenon are found in a wide area that begins in Sweden and ends in the Altai Mountains.^[13]^[14]^[15]

However, Jaakko Häkkinen argues that the language of the Volosovo culture was not itself Uralic, but a Paleo-European substratum to Uralic, especially its westernmost branches, and identifies Proto-Uralic with the Garino-Bor culture instead.^[16]

The Volosovo region was invaded by the Abashevo cultural groups at about 2300 BC. The latter buried their deceased in kurgans, and they are thought to have spoken a form of Indo-European ancestral to the Indo-Iranian languages and to have influenced the Volosovian vocabulary by introducing Aryan (Indo-Iranian) loan words. The Abashevo contributed to the fact that livestock farming and small-scale farming began to be practiced in the southern parts of the forest zone of Taiga.^[17]^[18]

It has been hypothesized that Pre-Proto-Uralic was spoken in Asia, on the basis of typological similarity with the Altaic Sprachbund^[19] and hypothetical early contacts with the Yukaghir languages.^[20] Aikio (2014) agrees with Häkkinen (2012) that Uralic–Yukaghir is unsupported and implausible, and that common vocabulary shared by the two families is best explained as the result of borrowing from Uralic into Yukaghir. However, Aikio (2014) puts the date of borrowing much later, arguing that the loanwords he accepts as valid were borrowed from an early stage of Samoyedic (preceding Proto-Samoyedic; thus roughly in the 1st millennium BC) into Yukaghir, in the same general region between the Yenisei River and Lake Baikal.^[21]

Continuity theories

Archaeological continuity has long been used as the basis of an argument for linguistic continuity. The argument was advanced by Estonians Paul Ariste and Harri Moora in 1956.^[22] Just as long, this kind of argumentation has also been heavily criticised. The oldest version of the continuity theory can be called the moderate or shallow continuity theory. It claims that linguistic continuity in Estonia and Finland can be traced back to the arrival of Typical Combed Ware, about 6,000 years ago. This view became mainstream in the multidisciplinary Tvärminne symposium in 1980.^[23] At the time, there seemed to be no serious linguistic results to contradict this archaeological view.

The continuity argumentation in the Uralic studies gained greater visibility in the 1990s, when the next step of the continuity theory was popularised (even though this line of reasoning had occasionally received airing). In the radical or deep continuity theory, it is claimed that the linguistic continuity in Finland could be traced back to the Mesolithic initial colonization, beyond 10,000 years.^[24]^[25]

However, in Indo-European studies, J. P. Mallory had already thoroughly scrutinized the methodological weaknesses of the continuity argumentation in 1989.^[26] In Uralic studies, it was also soon noted that the same argument (archaeological continuity) was used to support contradicting views, which revealed the method's unreliability.^[27]^[28]^[29]^[30]

At the same time, new linguistic results appeared to contradict the continuity theories: the datings of Proto-Saami^[31]^[32] and Proto-Finnic^[33] and of Proto-Uralic (Kallio 2006; Häkkinen 2009)^[5]^[34] are both clearly younger than it was thought in the framework of the continuity theories.

Nowadays linguists rarely believe in the continuity theories because of their shown methodological flaws and their incompatibility with the new linguistic results, but some archaeologists and laymen may still advance such arguments.^[35]

Modern view

In the 21st century, linguistic arguments have placed the Proto-Uralic homeland possibly around the Kama River or, more generally, close to the Great Volga Bend and the Ural Mountains, although Petri Kallio, while agreeing with the placement of the homeland in Central Russia, prefers the Volga-Oka region further to the west.^[36] The expansion of Proto-Uralic has been dated to about 2000 BC (4000 years ago), and its earlier stages go back at least one or two millennia earlier. Either way, this is considerably later than the earlier views of the continuity theories, which would place Proto-Uralic deep into Europe.^[5]^[34] Asko Parpola associates the early Proto-Uralic language with the Neolithic Elshanka and Kama cultures, placing the ultimate homeland of Uralic languages to the Kama River valley. Proto-Uralic would later expand with the Seima-Turbino material culture. According to him, Uralic languages were transmitted by language shift from groups of hunters and fishers participating in the spread of the Seima-Turbino culture towards Siberia.^[37]

An alternative view has been presented by Juha Janhunen (2009), who argues for a homeland in southern or central Siberia, somewhere between the Ob and Yenisei river or near the Sayan mountains in the Russian–Mongolian border region.^[35]

This view has been corroborated by a number of scholars, including by a team around Grünthal et al. 2022. They presented evidence that the Proto-Uralic homeland was located east of the Urals, likely somewhere in Siberia, and spoken by local hunter-gatherer communities. The Finno-Ugric branch spread along rivers north- and westwards into the Volga region, while the Samoyedic branch headed north-and eastwards. The spread of Uralic languages may be in part due to the Seima-Turbino phenomenon, although no direct link between them can be made yet. The material culture technology of proto-Uralic-speakers can be described as "Neolithic", as it included pottery but no vocabulary for food production. They further concluded that Proto-Uralic must have stood out of contact with Proto-Indo-European and did not share any genealogical ancestry with Indo-European, arguing that "whether based on cognacy or loans the argument from lexical resemblances is flawed", invalidating previous arguments for a homeland in the Volga region. Last, they noted that a number of traits of Uralic are "distinctive in western Eurasia. A number of typological properties are eastern-looking overall, fitting comfortably into northeast Asia, Siberia, or the North Pacific Rim", and are rare or absent in Europe, and that the Uralic languages "must have expanded via language shift", while "the shifting population had minimal impact on Proto-Uralic grammar and vocabulary".^[38]^[39]

Rasmus G. Bjørn (2022), citing the previous findings, such as Grünthal et al. 2022, argues that the Proto-Uralic speakers likely resided in Southern Siberia, and may have been part of the local Okunev culture in the Altai region, following the proposals of Janhunen and Peyrot. He notes that contact between Uralic and the Indo-European branches of Tocharian and Indo-Iranian at the one hand, and between Uralic and Turkic at the other hand, support a homeland near the Altai Mountains. An association with the Seima-Turbino phenomenon and the dispersal of Uralic languages "fits the string-like distribution of the western Uralic languages as well as close contacts with the Andronovo (and preceding Sintashta) culture associated with speakers of Indo-Iranian. Proto-Samoyedic was then either left in or migrated to the area around the Minusinsk Basin". Bjørn attributes the "eastern lexical elements" to contact with Turkic, which radiated out from the Amur region and can be associated with Northeast Asian linguistic area.^[40]

Jaakko Häkkinen (2023) argues that the location of Early Proto-Uralic and more distant stages are temporally irrelevant for the location of the Late Proto-Uralic homeland. He further elaborates that the "very distant Pre-Proto-Uralic" may have been spoken far from the region where Late Proto-Uralic was spoken. Häkkinen argues that Late Proto-Uralic and the successive stages of disintegration happened in the Central Ural Region, based on certain tree names and Indo-Iranian loanword layers. According to him, the disintegration of the proto-language began soon after 2500 BCE, but the speech community remained in a narrow region for a long time, until the second millennium BCE, after which the actual Uralic expansion began. On this basis the early phase of the Seima-Turbino Network in Southwestern Siberia could not yet be associated to the Uralic languages, but perhaps the later stages in Europe could. In any case, according to him, the arrival of the Uralic language in the Central Ural Region would precede even the early stage of the Seima-Turbino network.^[41]

Evidence from population genetics

Vladimir Napolskikh, who studied the origins of the "earth-diver" creation myths, has concluded that a certain variety of those myths, which is found in the folklore of Uralic-speaking peoples and other N1, C3, and Q (Y-DNA) carrying populations, originated in Siberia.^[42]

Genetic data suggests that early Uralic-speakers may be associated with hunter-gatherers in Western and Southern Siberia. These specific Uralic-speaking hunter-gatherers are argued to have formed from an admixture event between Western and Eastern Eurasian sources, with the calculated date of admixture is inferred to be ~7–9 kya. The Western source can be attributed to local Ancient North Eurasians, while the Eastern sources can be linked to "Neo-Siberians" (Eastern Siberian hunter-gatherers).^[43]

A number of population genomic studies in 2018 and 2019 note that the spread of Uralic languages may be associated with observed "Siberian" geneflow (represented by Nganasans) into the Eastern Baltic region.^[44] The proto-Uralic languages may be associated with early "Neo-Siberian" expansions outgoing from Northeast Asia westwards (~6-11kya), which largely replaced previous "Paleosiberian groups", but predated the expansion of Turkic languages.^[45]^[46]

Rasmus G. Bjørn (2022) argues for a link between the Proto-Uralic-speakers and the pre-Indo-European Okunevo culture, as well as to the Tarim EMBA cluster. He argues that following the expansion of Indo-Europeans eastwards and Northeast Asian (putative Turkic) groups westwards meeting in the Altai region, the early Uralic-speaking groups dispersed along the Seima-Turbino route. He also notes a correspondence between this expansion route and the frequency of haplogroup N.^[1]

Peltola et al. 2023 reproduced the findings that modern Uralic-speaking populations display varying degrees of ancestry from a "Siberian" source maximized in modern day Nganasan people. Nganasans and a historical specimen from Bronze Age Southern Siberia (Krasnoyarsk_Krai_BA; kra001) were found to display the lowest f4 estimate for the eastern ancestry among Uralic-speaking populations, and represent a plausible source. In contrast, affinity for Ancient North Eurasian-rich ancestry (represented by Eastern Hunter-Gatherers) was not significant and only observed among Western Finno-Ugric speakers.^[48]

References

^ ^a ^b Bjørn, Rasmus G. (2022). "Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone". Evolutionary Human Sciences. 4: e23. doi:10.1017/ehs.2022.16. ISSN 2513-843X. PMC 10432883. PMID 37599704.
^ Häkkinen, Jaakko 2007: Kantauralin murteutuminen vokaalivastaavuuksien valossak."Kantauralin murteutuminen vokaalivastaavuuksien valossa". Archived from the original on 2012-05-23. Retrieved 2012-04-16.
^ Nichols, Johanna (2021-01-14). "The Origin and Dispersal of Uralic: Distributional Typological View". Annual Review of Linguistics. 7 (1): 351–369. doi:10.1146/annurev-linguistics-011619-030405. ISSN 2333-9683.
^ Sebestyén-Németh, Irene 1951–1952: Zur Frage des alten Wohngebietes der uralischen Völker.
^ ^a ^b ^c Häkkinen, Jaakko 2009: Kantauralin ajoitus ja paikannus: perustelut puntarissa. – Suomalais-Ugrilaisen Seuran Aikakauskirja 92, p. 9–56. http://www.sgr.fi/susa/92/hakkinen.pdf
^ Rédei, Károly 1986: Zu den indogermanisch-uralischen Sprachkontakten. (eds. Manfred Mayrhofer & Wolfgang U. Dressler.) Veröffentlichungen der Kommission für Linguistik und Kommunikationsforschung, Heft 16. Wien.
^ Koivulehto, Jorma 1991: Koivulehto, Jorma 1991: Uralische Evidenz für die Laryngaltheorie. Österreichische Akademie der Wissenschaften. Philosophisch-Historische Klasse. Sitzungsberichte, 566. Band. Wien 1991.
^ Häkkinen, Jaakko 2012: Uralic evidence for the Indo-European homeland.
^ Mallory, J. P. & Adams, D. Q. (editors) 1997: Encyclopedia of Indo-European Culture. London and Chicago: Fitzroy Dearborn Publishers. p. 439
^ Edwin Bryant, Laurie L. Patton, The Indo-Aryan Controversy: Evidence and Inference in Indian History, Routledge 2005, p. 127
^ Parpola, Asko & Carpelan, Christian: The Cultural Counterparts to Proto-Indo-European, Proto-Uralic and Proto-Aryan - Matching the dispersal and contact patterns in the linguistic and archaeological record, pp. 107–141, 2005
^ Koshmenko, MG: The Culture of Bronze Age Net Ware in Karelia. Fennoscandia Archaeologica, 1996, No. XIII, pp. 51-67. Helsinki: Finnish Archaeological Society.
^ Gimbutas, Marija: Bronze Age Cultures in Central and Eastern Europe. The Hague, The Netherlands: Walter de Gruyter, 1965. p. 610
^ Nordqvist, Kerkko & Herva, Vesa-Pekka & Ikäheimo, Janne & Lahelma, Antti: Early Copper Use in Neolithic North-Eastern Europe: An Overview. Estonian Journal of Archeology, 2012, 16th ed., No. 1, pp. 3–25. Tallinn, Tartu, Estonia: Estonian Academy of Sciences.
^ Korjakova, Ludmila & Epimakhov, Andrei: The Urals and Western Siberia in the Bronze and Iron Ages. Cambridge, England: Cambridge University Press, 2007.
^ Häkkinen, Jaakko (2012). "Early contacts between Uralic and Yukaghir" (PDF). Suomalais-Ugrilaisen Seuran Toimituksia − Mémoires de la Société Finno-Ougrienne (264). Helsinki: Finno-Ugric Society: 96. Retrieved 13 July 2013.
^ Parpola, Asko: "The Face Urns of Gandhra and the Nâsatya Cult", Migrations, trade and peoples, Part 3, pp.149-162
^ Antony, David W.: The Horse, The Wheel, and Language: The Bronze-Age Riders from the Eurasian Steppes Shaped the Modern World. New Jersey, USA: Princeton University Press, 2010.
^ Janhunen, Juha 2001: Indo-Uralic and Ural-Altaic: On the diachronic implications of areal typology. – Carpelan, Parpola & Koskikallio (editors): Early Contacts between Uralic and Indo-European: Linguistic and Archaeological Considerations, p. 207–220. Suomalais-Ugrilaisen Seuran Toimituksia 242.
^ Häkkinen, Jaakko 2012: Early contacts between Uralic and Yukaghir. Tiina Hyytiäinen, Lotta Jalava, Janne Saarikivi & Erika Sandman (editors): Per Urales ad Orientem Iter polyphonicum multilingue Festskrift tillägnad Juha Janhunen på hans sextioårsdag den 12 februari 2012. Suomalais-Ugrilaisen Seuran Toimituksia 264, p. 91–101. Helsinki: Suomalais-Ugrilainen Seura. http://www.sgr.fi/sust/sust264/sust264_hakkinenj.pdf
^ Aikio, Ante (2014). "The Uralic–Yukaghir lexical correspondences: genetic inheritance, language contact or chance resemblance?". Finnisch-Ugrische Forschungen. 2014 (62): 7–76. doi:10.33339/fuf.86078
^ Moora, Harri (editor) 1956: Eesti rahva etnilisest ajaloost. Tallinn.
^ Gallén, Jarl (editor) 1984: Suomen väestön esihistorialliset juuret. Tvärminnen symposiumi 17.–19.1.1980. Bidrag till kännedom av Finlands natur och folk, 131. Helsinki: Societas Scientiarum Fennica.
^ Nuñez, Milton G. 1987: A Model for the Early Settlement of Finland. – Fennoscandia Archaeologica 4.
^ Wiik, Kalevi 2002: Eurooppalaisten juuret. Jyväskylä: Atena.
^ Mallory, J. P. 1989: In Search of the Indo-Europeans. Language, Archaeology and Myth. London: Thames and Hudson.
^ Mallory, J. P. 2001: Uralics and Indo-Europeans: Problems of time and space. Carpelan et al. (edited): Early Contacts between Uralic and Indo-European: Linguistic and Archaeological Considerations, p. 345–366. Suomalais-Ugrilaisen Seuran Toimituksia 242.
^ Aikio, Ante; Aikio, Aslak (2001). "Heimovaelluksista jatkuvuuteen. Suomalaisen väestöhistorian tutkimuksen pirstoutuminen" (PDF). Muinaistutkija. 4. Helsinki: Suomen arkeologinen seura: 2–21. Archived from the original (PDF) on 2008-02-27. Retrieved 2008-01-07.
^ Häkkinen, Jaakko 2006: Studying the Uralic proto-language. http://www.alkuperasivusto.fi/Uralic.html [Translation from: Uralilaisen kantakielen tutkiminen. – Tieteessä tapahtuu 1 / 2006, p. 52–58.]
^ Häkkinen, Jaakko 2010: Jatkuvuusperustelut ja saamelaisen kielen leviäminen (OSA 1). – Muinaistutkija 1 / 2010, p. 19–36. http://www.alkuperasivusto.fi/Jatkuvuus1.pdf
^ Aikio, Ante 2004: An essay on substrate studies and the origin of Saami. – Irma Hyvärinen, Petri Kallio & Jarmo Korhonen (toim.): Etymologie, Entlehnungen und Entwicklungen. Festschrift für Jorma Koivulehto zum 70. Geburtstag, s. 5–34. Mémoires de la Société Néophilologique de Helsinki, LXIII. Helsinki: Uusfilologinen yhdistys ry.
^ Aikio, Ante 2006: On Germanic-Saami contacts and Saami prehistory. – Suomalais-Ugrilaisen Seuran Aikakauskirja 91, p. 9–55. Helsinki: Suomalais-Ugrilainen Seura. http://www.sgr.fi/susa/91/aikio.pdf
^ Saarikivi, Janne & Grünthal, Riho 2005: Itämerensuomalaisten kielten uralilainen tausta. Muuttuva muoto. Kirjoituksia Tapani Lehtisen 60-vuotispäivän kunniaksi, s. 111–146. Kieli 16.
^ ^a ^b Kallio, Petri 2006: Suomen kantakielten absoluuttista kronologiaa. – Virittäjä 1 / 2006, p. 2–25. http://www.kotikielenseura.fi/virittaja/hakemistot/jutut/2006_2.pdf
^ ^a ^b Janhunen, Juha (2009). "Proto-Uralic—what, where and when?" (PDF). In Ylikoski, Jussi (ed.). The Quasquicentennial of the Finno-Ugrian Society. Suomalais-Ugrilaisen Seuran Toimituksia 258. Helsinki: Société Finno-Ougrienne. ISBN 978-952-5667-11-0. ISSN 0355-0230.
^ Kallio, Petri (2015). "The language contact situation in prehistoric Northeastern Europe". In Mailhammer, Robert; Vennemann gen. Nierfeld, Theo; Olsen, Birgit Anette (eds.). The Linguistic Roots of Europe: Origin and Development of European Languages. Copenhagen: Museum Tusculanum Press (Copenhagen Studies in Indo-European 6). p. 86. ISBN 978-87-635-4209-8.
^ Парпола, Аско (2022-04-29). "Location of the Uralic proto-language in the Kama River Valley and the Uralic speakers' Expansion east and west with the 'Sejma-Turbino transcultural phenomenon' 2200-1900 BC". Археология Евразийских степей (2): 258–277. doi:10.24852/2587-6112.2022.2.258.277. hdl:10138/354948. ISSN 2618-9488. Alternative link to free pdf:https://www.researchgate.net/publication/360290015_Location_of_the_Uralic_proto-language_in_the_Kama_River_Valley_and_the_Uralic_speakers'_Expansion_east_and_west_with_the_'Sejma-Turbino_transcultural_phenomenon'_2200-1900_BC
^ Grünthal, Riho; Heyd, Volker; Holopainen, Sampsa; Janhunen, Juha; Khanina, Olga; Miestamo, Matti; Nichols, Johanna; Saarikivi, Janne; Sinnemäki, Kaius (2022-08-29). "Drastic demographic events triggered the Uralic spread". Diachronica. 39 (4): 490–524. doi:10.1075/dia.20038.gru. hdl:10138/347633. ISSN 0176-4225. S2CID 248059749.
^ Nichols, Johanna (2021-01-01). "The Origin and Dispersal of Uralic: Distributional Typological View". Annual Review of Linguistics. Rochester, NY. SSRN 3778495. PU had (and the modern Uralic languages mostly still have) a number of typological traits that are typical of the greater North Pacific Rim area and not of western Eurasia. These traits are shared with many Siberian languages but are (otherwise) rare in Europe. More precisely, the patterns, as types, are found throughout Siberia and nearby; in Uralic, the morphemes marking them are generally native to Uralic. Uralic languages also have a number of traits that define what I call the Inner Asian type. These traits are not all typical of the North Pacific Rim or Siberian linguistic populations but rather are hallmarks of a smaller set of families: prototypically, Tungusic, Turkic, Mongolic, and Uralic or at least eastern Uralic; to a lesser extent, Korean, Japonic, and Eskimo-Aleut.
^ Bjørn, Rasmus G. (2022). "Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone". Evolutionary Human Sciences. 4: e23. doi:10.1017/ehs.2022.16. ISSN 2513-843X. PMC 10432883. PMID 37599704. S2CID 248358873. Proto-Uralic was probably spoken in the Okunevo culture that constitutes the latest possible period for adoption of loanwords before the formation of the separate branches. The analysis also supports the proposition that an early stage of the Proto-Turkic language community was present around the Altai Mountains before 2000 BC, probably reflected in the eastern genetic component of Okunevo, and surely in the culturally transmitted loanwords of Indo-European provenance shared with early Uralic.
^ Häkkinen, Jaakko (2023). "On Locating Proto-Uralic". Finnisch-Ugrische Forschungen. 68 (68): 43–100. doi:10.33339/fuf.120910.
^ Napolskikh V. V. (Izhevsk, Russia). Earth-Diver Myth (А812) in northern Eurasia and North America: twenty years later.
^ Wong, Emily H.M. (2015). "Reconstructing genetic history of Siberian and Northeastern European populations". Genome Research. 27 (1): 1–14. doi:10.1101/gr.202945.115. PMC 5204334. PMID 27965293. Our results also provide insights for ongoing anthropological debates about the origin of indigenous populations in the Uralic region. Uralic people had been considered as mixed groups descended from the longtime admixture between Europeans and Asians. Our model of gene flows during the last 50,000 yr in Europe, East Asia, and Siberia (Fig. 8) provides support for Bunak's hypothesis since Western Siberians were associated with a distinct ancient lineage, possibly related to ANE people. At the same time, our models suggest a more complex scenario that included admixture with ancient Eastern Siberians who came into contact with proto Western Siberians ~7–9 kya.
^ Tambets, Kristiina; Metspalu, Mait; Lang, Valter; Villems, Richard; Kivisild, Toomas; Kriiska, Aivar; Thomas, Mark G.; Díez del Molino, David; Crema, Enrico Ryunosuke (2019). "The Arrival of Siberian Ancestry Connecting the Eastern Baltic to Uralic Speakers further East". Current Biology. 29 (10): 1701–1711.e16. doi:10.1016/j.cub.2019.04.026. ISSN 0960-9822. PMC 6544527. PMID 31080083.
^ Sikora, Martin; Pitulko, Vladimir V.; Sousa, Vitor C.; Allentoft, Morten E.; Vinner, Lasse; Rasmussen, Simon; Margaryan, Ashot; de Barros Damgaard, Peter; de la Fuente, Constanza; Renaud, Gabriel; Yang, Melinda A.; Fu, Qiaomei; Dupanloup, Isabelle; Giampoudakis, Konstantinos; Nogués-Bravo, David (June 2019). "The population history of northeastern Siberia since the Pleistocene". Nature. 570 (7760): 182–188. doi:10.1038/s41586-019-1279-z. hdl:1887/3198847. ISSN 1476-4687. Most modern Siberian speakers of Neosiberian languages genetically fall on an East- West cline between Europeans and Early East Asians. Taking Even speakers as representatives, the Neosiberian turnover from the south, which largely replaced Ancient Paleosiberian ancestry, can be associated with the northward spread of Tungusic and probably also Turkic and Mongolic. However, the expansions of Tungusic as well as Turkic and Mongolic are too recent to be associable with the earliest waves of Neosiberian ancestry, dated later than ~11 kya, but discernible in the Baikal region from at least 6 kya onwards. Therefore, this phase of the Neosiberian population turnover must initially have transmitted other languages or language families into Siberia, including possibly Uralic and Yukaghir.
^ Tambets, Kristiina; Yunusbayev, Bayazit; Hudjashov, Georgi; Ilumäe, Anne-Mai; Rootsi, Siiri; Honkola, Terhi; Vesakoski, Outi; Atkinson, Quentin; Skoglund, Pontus; Kushniarevich, Alena; Litvinov, Sergey; Reidla, Maere; Metspalu, Ene; Saag, Lehti; Rantanen, Timo (2018-09-21). "Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations". Genome Biology. 19 (1): 139. doi:10.1186/s13059-018-1522-1. ISSN 1474-760X. PMC 6151024. PMID 30241495.
^ Bjørn, Rasmus G. (January 2022). "Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone". Evolutionary Human Sciences. 4: e23. doi:10.1017/ehs.2022.16. ISSN 2513-843X. PMC 10432883. PMID 37599704.
^ Peltola, Sanni; Majander, Kerttu; Makarov, Nikolaj; Dobrovolskaya, Maria; Nordqvist, Kerkko; Salmela, Elina; Onkamo, Päivi (2023-01-09). "Genetic admixture and language shift in the medieval Volga-Oka interfluve". Current Biology. 33 (1): 174–182.e10. doi:10.1016/j.cub.2022.11.036. ISSN 0960-9822.

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+{{Short description|Location where the Proto-Uralic language originated}}
 [[File:Fenno-Ugrian_languages.png|thumb|upright=2|Current distribution of the Uralic languages|300px]]
+The '''Proto-Uralic homeland''' is the earliest location in which the [[Proto-Uralic language]] was spoken, before its speakers dispersed geographically causing it to diverge into multiple languages. Various locations have been proposed and debated, although as of 2022 "scholarly consensus now gravitates towards a relatively recent provenance of the Uralic languages east of the Ural mountains".<ref name=":0" />
-Various '''Proto-Uralic homeland hypotheses''' on the origin of the [[Uralic languages]] and the location ([[Urheimat]] or homeland) and period in which the [[Proto-Uralic language]] was spoken, have been advocated over the years.
 ==Homeland hypotheses==
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 It has been suggested that the Proto-Uralic homeland was located near the [[Ural Mountains]], either on the European or the Siberian side. The main reason to suppose that there was a Siberian homeland has been the traditional taxonomic model that sees the [[Samoyedic languages|Samoyedic]] branch as splitting off first. Because the present border between the Samoyedic and the [[Ugric languages|Ugric]] branch is in Western Siberia, the original split was seen to have occurred there too.
-However, because the Ugric languages are known to have been spoken earlier on the European side of the Urals, a European homeland would be equally possible. In recent years, it has also been argued on the basis of phonology that the oldest split was not between the Samoyedic and the [[Finno-Ugric]] but between the [[Finno-Permic]] and the Ugro-Samoyedic language groups.<ref>Häkkinen, Jaakko 2007: Kantauralin murteutuminen vokaalivastaavuuksien valossak.{{cite web |url=https://oa.doria.fi/handle/10024/7044 |title=Kantauralin murteutuminen vokaalivastaavuuksien valossa |access-date=2012-04-16 |url-status=dead |archive-url=https://archive.today/20120523223524/https://oa.doria.fi/handle/10024/7044 |archive-date=2012-05-23 }}</ref> The lexical level is argued to be less reliable, and lexical innovativeness (a small number of shared cognates) can be confused because of the great age of the division. For a long time, no new arguments for a Siberian homeland have been presented.
+However, a European homeland would be equally possible because the Ugric languages are known to have been spoken earlier on the European side of the Urals. In recent years, it has also been argued based on phonology that the oldest split was not between the Samoyedic and the [[Finno-Ugric]] but between the [[Finno-Permic]] and the Ugro-Samoyedic language groups.<ref>Häkkinen, Jaakko 2007: Kantauralin murteutuminen vokaalivastaavuuksien valossak.{{cite web |url=https://oa.doria.fi/handle/10024/7044 |title=Kantauralin murteutuminen vokaalivastaavuuksien valossa |access-date=2012-04-16 |url-status=dead |archive-url=https://archive.today/20120523223524/https://oa.doria.fi/handle/10024/7044 |archive-date=2012-05-23 }}</ref> The lexical level is argued to be less reliable, and lexical innovativeness (a small number of shared cognates) can be confused because of the great age of the division. No new arguments for a Siberian homeland have been presented for a long time.
-Both European and Siberian homeland proposals have been supported by palaeolinguistic evidence, but only those cases in which the semantic reconstructions are certain are valid. A Siberian homeland has been claimed on the basis of two coniferous tree names in Proto-Uralic, but the trees (''[[Abies sibirica]]'' and ''[[Pinus cembra]]''{{Clarify|reason=The second species named is actually native to the Alps and Carpathians.|date=May 2023}}) have for a long time been present also in the far east of Europe. A European homeland is supported by words for 'bee', 'honey', 'elm' etc.<ref>Sebestyén-Németh, Irene 1951–1952: Zur Frage des alten Wohngebietes der uralischen Völker.</ref> They can be reconstructed already to Proto-Uralic, if Samoyedic is no longer seen as the first branch to split off.<ref name="JH09">Häkkinen, Jaakko 2009: Kantauralin ajoitus ja paikannus: perustelut puntarissa. – Suomalais-Ugrilaisen Seuran Aikakauskirja 92, p. 9–56. http://www.sgr.fi/susa/92/hakkinen.pdf</ref>
+Both European and Siberian homeland proposals have been supported by palaeolinguistic evidence, but only those cases in which the semantic reconstructions are certain and valid. A Siberian homeland has been claimed based on two coniferous tree names in Proto-Uralic, but the trees (''[[Abies sibirica]]'' and ''[[Pinus sibirica]]'')<ref>{{Cite journal |last=Nichols |first=Johanna |date=2021-01-14 |title=The Origin and Dispersal of Uralic: Distributional Typological View |url=https://www.annualreviews.org/doi/10.1146/annurev-linguistics-011619-030405 |journal=Annual Review of Linguistics |language=en |volume=7 |issue=1 |pages=351–369 |doi=10.1146/annurev-linguistics-011619-030405 |issn=2333-9683}}</ref> have for a long time been present also in the far east of Europe. A European homeland is supported by words for 'bee', 'honey', 'elm' etc.<ref>Sebestyén-Németh, Irene 1951–1952: Zur Frage des alten Wohngebietes der uralischen Völker.</ref> They can be reconstructed already to Proto-Uralic, if Samoyedic is no longer seen as the first branch to split off.<ref name="JH09">Häkkinen, Jaakko 2009: Kantauralin ajoitus ja paikannus: perustelut puntarissa. – Suomalais-Ugrilaisen Seuran Aikakauskirja 92, p. 9–56. http://www.sgr.fi/susa/92/hakkinen.pdf</ref>
 More recently, [[loanword]] evidence has also been used to support a European homeland. Proto-Uralic has been seen as borrowing words from [[Proto-Indo-European language|Proto-Indo-European]],<ref>Rédei, Károly 1986: Zu den indogermanisch-uralischen Sprachkontakten. (eds. Manfred Mayrhofer & Wolfgang U. Dressler.) Veröffentlichungen der Kommission für Linguistik und Kommunikationsforschung, Heft 16. Wien.</ref><ref>Koivulehto, Jorma 1991: Koivulehto, Jorma 1991: Uralische Evidenz für die Laryngaltheorie. Österreichische Akademie der Wissenschaften. Philosophisch-Historische Klasse. Sitzungsberichte, 566. Band. Wien 1991.</ref> and the Proto-Indo-European homeland has rarely been located east of the Urals. Proto-Uralic even seems to have developed in close contact with [[Proto-Indo-Iranian language|Proto-Indo-Iranian]],<ref>Häkkinen, Jaakko 2012: [http://www.alkuperasivusto.fi/UralicEvidence.pdf ''Uralic evidence for the Indo-European homeland''].</ref> which is seen as having arisen in the [[Poltavka culture]] of the Caspian steppes before its spread to Asia.<ref>Mallory, J. P. & Adams, D. Q. (editors) 1997: Encyclopedia of Indo-European Culture. London and Chicago: Fitzroy Dearborn Publishers. p. 439</ref>
-The [[Lyalovo culture]] (ca. 5000–3650 BC) has been equated with the Proto-Uralic urheimat, and the following [[Volosovo culture]] (ca. 3650–1900 BC) with the Proto-Finno-Ugric urheimat.<ref>Edwin Bryant, Laurie L. Patton, ''The Indo-Aryan Controversy: Evidence and Inference in Indian History'', Routledge 2005, p. 127</ref> Some scholars believe that the culture of Lyalovo was in fact the Proto-Uralic urheimat and that its inhabitants spread Uralic languages to north-eastern Europe.<ref>Parpola, Asko & Carpelan, Christian: The Cultural Counterparts to Proto-Indo-European, Proto-Uralic and Proto-Aryan - Matching the dispersal and contact patterns in the linguistic and archaeological record'', pp. 107–141, 2005</ref> The Volosovo culture has been named the Bronze Age [[Successor culture|Successor Culture]], a textile-ceramic culture that developed in the region between Upper Volga and lakes Ladoga and Onega. It was distinguished from other groups based on the traces of textile used for the production of ceramics, and spread southeast all the way to central Volga, south to the entire river valley of the Oka, southwest to the northern shore of the Daugava, and northwest of Fennoscandia to Karelia, Finland and northern Sweden and Norway.<ref>Koshmenko, MG: The Culture of Bronze Age Net Ware in Karelia. Fennoscandia Archaeologica, 1996, No. XIII, pp. 51-67. Helsinki: Finnish Archaeological Society.</ref> Known as the [[Seima-Turbino phenomenon]], it was a culturally unified, extensive network of trade in copper and bronze. The traces of the Seima-Turbino phenomenon are found in a wide area that begins in Sweden and ends in the Altai Mountains.<ref>Gimbutas, Marija: ''Bronze Age Cultures in Central and Eastern Europe''. The Hague, The Netherlands: Walter de Gruyter, 1965. p. 610</ref><ref>Nordqvist, Kerkko & Herva, Vesa-Pekka & Ikäheimo, Janne & Lahelma, Antti: Early Copper Use in Neolithic North-Eastern Europe: An Overview. Estonian Journal of Archeology, 2012, 16th ed., No. 1, pp. 3–25. Tallinn, Tartu, Estonia: Estonian Academy of Sciences.</ref><ref>Korjakova, Ludmila & Epimakhov, Andrei: ''The Urals and Western Siberia in the Bronze and Iron Ages''. Cambridge, England: Cambridge University Press, 2007.</ref>
+Bryant et al. (2005) equated the {{ill|Lyalovo culture|fi|Ljalovon kulttuuri|ru|Льяловская культура}} (ca. 5000–3650 BC) with the Proto-Uralic urheimat, and the following [[Volosovo culture]] (ca. 3650–1900 BC) with the Proto-Finno-Ugric urheimat.<ref>Edwin Bryant, Laurie L. Patton, ''The Indo-Aryan Controversy: Evidence and Inference in Indian History'', Routledge 2005, p. 127</ref> Two Finnish scholars believe that the culture of Lyalovo was the Proto-Uralic urheimat and that its inhabitants spread Uralic languages to north-eastern Europe.<ref>Parpola, Asko & Carpelan, Christian: The Cultural Counterparts to Proto-Indo-European, Proto-Uralic and Proto-Aryan - Matching the dispersal and contact patterns in the linguistic and archaeological record'', pp. 107–141, 2005</ref> The Volosovo culture has been named the Bronze Age [[Successor culture|Successor Culture]], a textile-ceramic culture that developed in the region between Upper Volga and lakes Ladoga and Onega. It was distinguished from other groups based on the traces of textile used for the production of ceramics, and spread southeast all the way to central Volga, south to the entire river valley of the Oka, southwest to the northern shore of the Daugava, and northwest of Fennoscandia to Karelia, Finland and northern Sweden and Norway.<ref>Koshmenko, MG: The Culture of Bronze Age Net Ware in Karelia. Fennoscandia Archaeologica, 1996, No. XIII, pp. 51-67. Helsinki: Finnish Archaeological Society.</ref> Known as the [[Seima-Turbino phenomenon]], it was a culturally unified, extensive network of trade in copper and bronze. The traces of the Seima-Turbino phenomenon are found in a wide area that begins in Sweden and ends in the Altai Mountains.<ref>Gimbutas, Marija: ''Bronze Age Cultures in Central and Eastern Europe''. The Hague, The Netherlands: Walter de Gruyter, 1965. p. 610</ref><ref>Nordqvist, Kerkko & Herva, Vesa-Pekka & Ikäheimo, Janne & Lahelma, Antti: Early Copper Use in Neolithic North-Eastern Europe: An Overview. Estonian Journal of Archeology, 2012, 16th ed., No. 1, pp. 3–25. Tallinn, Tartu, Estonia: Estonian Academy of Sciences.</ref><ref>Korjakova, Ludmila & Epimakhov, Andrei: ''The Urals and Western Siberia in the Bronze and Iron Ages''. Cambridge, England: Cambridge University Press, 2007.</ref>
 However, Jaakko Häkkinen argues that the language of the Volosovo culture was not itself Uralic, but a Paleo-European substratum to Uralic, especially its westernmost branches, and identifies Proto-Uralic with the Garino-Bor culture instead.<ref>{{cite journal|last=Häkkinen|first=Jaakko|title=Early contacts between Uralic and Yukaghir|journal=Suomalais-Ugrilaisen Seuran Toimituksia − Mémoires de la Société Finno-Ougrienne|year=2012|issue=264|page=96|url=http://www.sgr.fi/sust/sust264/sust264_hakkinenj.pdf|access-date=13 July 2013|publisher=Finno-Ugric Society|location=Helsinki}}</ref>
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 The continuity argumentation in the Uralic studies gained greater visibility in the 1990s, when the next step of the continuity theory was popularised (even though this line of reasoning had occasionally received airing). In the radical or deep continuity theory, it is claimed that the linguistic continuity in Finland could be traced back to the [[Mesolithic]] initial colonization, beyond 10,000 years.<ref>Nuñez, Milton G. 1987: A Model for the Early Settlement of Finland. – Fennoscandia Archaeologica 4.</ref><ref>Wiik, Kalevi 2002: Eurooppalaisten juuret. Jyväskylä: Atena.</ref>
-However, in Indo-European studies, J. P. Mallory had already thoroughly scrutinized the methodological weaknesses of the continuity argumentation in 1989.<ref>Mallory, J. P. 1989: In Search of the Indo-Europeans. Language, Archaeology and Myth. London: Thames and Hudson.</ref> In Uralic studies, it was also soon noted that the same argument (archaeological continuity) was used to support contradicting views, which revealed the method's unreliability.<ref>Mallory, J. P. 2001: Uralics and Indo-Europeans: Problems of time and space. Carpelan et al. (edited): Early Contacts between Uralic and Indo-European: Linguistic and Archaeological Considerations, p. 345–366. Suomalais-Ugrilaisen Seuran Toimituksia 242.</ref><ref>Aikio, Ante & Aikio, Aslak 2001: Heimovaelluksista jatkuvuuteen. Suomalaisen väestöhistorian tutkimuksen pirstoutuminen. – Muinaistutkija 4/2001, p. 2–21. Helsinki: Suomen arkeologinen seura. {{cite web |url=http://cc.oulu.fi/~anaikio/Heimovaelluksista_jatkuvuuteen.pdf |title=Archived copy |access-date=2008-01-07 |url-status=dead |archive-url=https://web.archive.org/web/20080227095022/http://cc.oulu.fi/~anaikio/Heimovaelluksista_jatkuvuuteen.pdf |archive-date=2008-02-27 }}</ref><ref>Häkkinen, Jaakko 2006: Studying the Uralic proto-language. http://www.alkuperasivusto.fi/Uralic.html [Translation from: Uralilaisen kantakielen tutkiminen. – Tieteessä tapahtuu 1 / 2006, p. 52–58.]</ref><ref>Häkkinen, Jaakko 2010: Jatkuvuusperustelut ja saamelaisen kielen leviäminen (OSA 1). – Muinaistutkija 1 / 2010, p. 19–36. http://www.alkuperasivusto.fi/Jatkuvuus1.pdf</ref>
+However, in Indo-European studies, J. P. Mallory had already thoroughly scrutinized the methodological weaknesses of the continuity argumentation in 1989.<ref>Mallory, J. P. 1989: In Search of the Indo-Europeans. Language, Archaeology and Myth. London: Thames and Hudson.</ref> In Uralic studies, it was also soon noted that the same argument (archaeological continuity) was used to support contradicting views, which revealed the method's unreliability.<ref>Mallory, J. P. 2001: Uralics and Indo-Europeans: Problems of time and space. Carpelan et al. (edited): Early Contacts between Uralic and Indo-European: Linguistic and Archaeological Considerations, p. 345–366. Suomalais-Ugrilaisen Seuran Toimituksia 242.</ref><ref>{{cite journal |author1=Aikio, Ante |author2=Aikio, Aslak |date=2001 |title=Heimovaelluksista jatkuvuuteen. Suomalaisen väestöhistorian tutkimuksen pirstoutuminen |journal=Muinaistutkija |volume=4 |page=2–21 |location=Helsinki |publisher=Suomen arkeologinen seura |url=http://cc.oulu.fi/~anaikio/Heimovaelluksista_jatkuvuuteen.pdf |access-date=2008-01-07 |url-status=dead |archive-url=https://web.archive.org/web/20080227095022/http://cc.oulu.fi/~anaikio/Heimovaelluksista_jatkuvuuteen.pdf |archive-date=2008-02-27 }}</ref><ref>Häkkinen, Jaakko 2006: Studying the Uralic proto-language. http://www.alkuperasivusto.fi/Uralic.html [Translation from: Uralilaisen kantakielen tutkiminen. – Tieteessä tapahtuu 1 / 2006, p. 52–58.]</ref><ref>Häkkinen, Jaakko 2010: Jatkuvuusperustelut ja saamelaisen kielen leviäminen (OSA 1). – Muinaistutkija 1 / 2010, p. 19–36. http://www.alkuperasivusto.fi/Jatkuvuus1.pdf</ref>
 At the same time, new linguistic results appeared to contradict the continuity theories: the datings of [[Proto-Samic language|Proto-Saami]]<ref>Aikio, Ante 2004: An essay on substrate studies and the origin of Saami. – Irma Hyvärinen, Petri Kallio & Jarmo Korhonen (toim.): Etymologie, Entlehnungen und Entwicklungen. Festschrift für Jorma Koivulehto zum 70. Geburtstag, s. 5–34. Mémoires de la Société Néophilologique de Helsinki, LXIII. Helsinki: Uusfilologinen yhdistys ry.</ref><ref>Aikio, Ante 2006: On Germanic-Saami contacts and Saami prehistory. – Suomalais-Ugrilaisen Seuran Aikakauskirja 91, p. 9–55. Helsinki: Suomalais-Ugrilainen Seura. http://www.sgr.fi/susa/91/aikio.pdf</ref> and Proto-Finnic<ref>Saarikivi, Janne & Grünthal, Riho 2005: Itämerensuomalaisten kielten uralilainen tausta. Muuttuva muoto. Kirjoituksia Tapani Lehtisen 60-vuotispäivän kunniaksi, s. 111–146. Kieli 16.</ref> and of Proto-Uralic (Kallio 2006; Häkkinen 2009)<ref name="JH09"/><ref name="PK06">Kallio, Petri 2006: Suomen kantakielten absoluuttista kronologiaa. – Virittäjä 1 / 2006, p. 2–25.
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 ==Modern view==
-In the 21st century, linguistic arguments have placed the Proto-Uralic homeland possibly around the [[Kama River]] or, more generally, close to the Great [[Volga]] Bend and the [[Ural Mountains]], although Petri Kallio, while agreeing with the placement of the homeland in Central Russia, prefers the Volga-Oka region further to the west.<ref>{{cite book |last1=Kallio |first1=Petri |editor1-last=Mailhammer |editor1-first=Robert |editor2-last=Vennemann gen. Nierfeld |editor2-first=Theo |editor3-last=Olsen |editor3-first=Birgit Anette |title=The Linguistic Roots of Europe: Origin and Development of European Languages |date=2015 |publisher=Museum Tusculanum Press (Copenhagen Studies in Indo-European 6) |location=Copenhagen |isbn=978-87-635-4209-8 |page=86 |chapter-url=https://www.academia.edu/20252178 |language=en |chapter=The language contact situation in prehistoric Northeastern Europe}}</ref> The expansion of Proto-Uralic has been dated to about 2000 BC (4000 years ago), and its earlier stages go back at least one or two millennia earlier. Either way, this is considerably later than the earlier views of the continuity theories, which would place Proto-Uralic deep into Europe.<ref name="JH09"/><ref name="PK06"/> [[Asko Parpola]] associates the early [[Proto-Uralic language]] with the Neolithic [[Elshanka culture|Elshanka]] and [[Kama culture]]s, placing the ultimate homeland of Uralic languages to the [[Kama (river)|Kama River valley]]. Proto-Uralic would later expand with the [[Seima-Turbino phenomenon|Seima-Turbino material culture]]. According to him, Uralic languages were transmitted by language shift from groups of hunters and fishers participating in the spread of the Seima-Turbino culture towards Siberia.<ref>{{Cite journal |last=Парпола |first=Аско |date=2022-04-29 |title=Location of the Uralic proto-language in the Kama River Valley and the Uralic speakers' Expansion east and west with the 'Sejma-Turbino transcultural phenomenon’ 2200-1900 BC |url=https://evrazstep.ru/index.php/aes/article/view/724 |journal=Археология Евразийских степей |language=en |issue=2 |pages=258–277 |doi=10.24852/2587-6112.2022.2.258.277 |issn=2618-9488 |quote=Alternative link to free pdf:https://www.researchgate.net/publication/360290015_Location_of_the_Uralic_proto-language_in_the_Kama_River_Valley_and_the_Uralic_speakers'_Expansion_east_and_west_with_the_'Sejma-Turbino_transcultural_phenomenon'_2200-1900_BC}}</ref>
+In the 21st century, linguistic arguments have placed the Proto-Uralic homeland possibly around the [[Kama River]] or, more generally, close to the [[Samara Bend|Great Volga Bend]] and the [[Ural Mountains]], although Petri Kallio, while agreeing with the placement of the homeland in Central Russia, prefers the Volga-Oka region further to the west.<ref>{{cite book |last1=Kallio |first1=Petri |editor1-last=Mailhammer |editor1-first=Robert |editor2-last=Vennemann gen. Nierfeld |editor2-first=Theo |editor3-last=Olsen |editor3-first=Birgit Anette |title=The Linguistic Roots of Europe: Origin and Development of European Languages |date=2015 |publisher=Museum Tusculanum Press (Copenhagen Studies in Indo-European 6) |location=Copenhagen |isbn=978-87-635-4209-8 |page=86 |chapter-url=https://www.academia.edu/20252178 |language=en |chapter=The language contact situation in prehistoric Northeastern Europe}}</ref> The expansion of Proto-Uralic has been dated to about 2000 BC (4000 years ago), and its earlier stages go back at least one or two millennia earlier. Either way, this is considerably later than the earlier views of the continuity theories, which would place Proto-Uralic deep into Europe.<ref name="JH09"/><ref name="PK06"/> [[Asko Parpola]] associates the early [[Proto-Uralic language]] with the Neolithic [[Elshanka culture|Elshanka]] and [[Kama culture]]s, placing the ultimate homeland of Uralic languages to the [[Kama (river)|Kama River valley]]. Proto-Uralic would later expand with the [[Seima-Turbino phenomenon|Seima-Turbino material culture]]. According to him, Uralic languages were transmitted by language shift from groups of hunters and fishers participating in the spread of the Seima-Turbino culture towards Siberia.<ref>{{Cite journal |last=Парпола |first=Аско |date=2022-04-29 |title=Location of the Uralic proto-language in the Kama River Valley and the Uralic speakers' Expansion east and west with the 'Sejma-Turbino transcultural phenomenon' 2200-1900 BC |url=https://evrazstep.ru/index.php/aes/article/view/724 |journal=Археология Евразийских степей |language=en |issue=2 |pages=258–277 |doi=10.24852/2587-6112.2022.2.258.277 |issn=2618-9488 |quote=Alternative link to free pdf:https://www.researchgate.net/publication/360290015_Location_of_the_Uralic_proto-language_in_the_Kama_River_Valley_and_the_Uralic_speakers'_Expansion_east_and_west_with_the_'Sejma-Turbino_transcultural_phenomenon'_2200-1900_BC|hdl=10138/354948 |hdl-access=free }}</ref>
+An alternative view has been presented by [[Juha Janhunen]] (2009), who argues for a homeland in southern or central [[Siberia]], somewhere between the [[Ob River|Ob]] and [[Yenisei River|Yenisei river]] or near the [[Sayan Mountains|Sayan mountains]] in the [[Russia]]n–[[Mongolia]]n border region.<ref name="Janhunen2009"/>
-An alternative view has been presented by [[Juha Janhunen]] (2009), who argues for a homeland in eastern or central [[Siberia]], somewhere between the [[Ob River|Ob]] and [[Yenisei River|Yenisei river]] or near the [[Sayan Mountains|Sayan mountains]] in the [[Russia]]n–[[Mongolia]]n border region.<ref name="Janhunen2009"/> This view has been corroborated by a number of scholars, including recently by a team around Grünthal et al. 2022. They presented evidence that the Proto-Uralic homeland was located East of the Urals, likely somewhere in [[Siberia]] and spoken by local hunter-gatherers. The [[Finno-Ugric languages|Finno-Ugric branch]] spread along rivers north- and westwards into the [[Volga region]], while the [[Samoyedic languages|Samoyedic branch]] headed eastwards. The spread of Uralic languages may be in part due to the [[Seima-Turbino phenomenon]], although no direct link between them can be made yet. The material culture technology of proto-Uralic-speakers can be described as "Neolithic", as it included pottery but no vocabulary for food production. They further concluded that Proto-Uralic must have stood out of contact with Proto-Indo-European and did not share any genealogical ancestry with Indo-European, arguing that "''whether based on cognacy or loans the argument from lexical resemblances is flawed''", invalidating previous arguments for a homeland in the Volga region. Lastly they noted that a number of traits of Uralic are "''distinctive in western Eurasia. A number of typological properties are eastern-looking overall, fitting comfortably into northeast Asia, Siberia, or the North Pacific Rim''", but also with the "''Northeast Asian linguistic area''" (also known as "Altaic areal grouping"), which are rare or absent in Europe, and that the Uralic languages "''must have expanded via language shift''", while "''the shifting population had minimal impact on Proto-Uralic grammar and vocabulary''".<ref>{{Cite journal|url=https://researchportal.helsinki.fi/en/publications/drastic-demographic-events-triggered-the-uralic-spread|title=Drastic demographic events triggered the Uralic spread|journal=Diachronica|year=2022|last1=Grünthal|first1=Riho|volume=39 |issue=4 |pages=490–524 |doi=10.1075/dia.20038.gru|hdl=10138/347633 |s2cid=248059749|hdl-access=free}}</ref><ref>{{Cite journal |last=Nichols |first=Johanna |date=2021-01-01 |title=The Origin and Dispersal of Uralic: Distributional Typological View |url=https://papers.ssrn.com/abstract=3778495 |journal= |language=en |location=Rochester, NY |quote=PU had (and the modern Uralic languages mostly still have) a number of typological traits that are typical of the greater North Pacific Rim area and not of western Eurasia. These traits are shared with many Siberian languages but are (otherwise) rare in Europe. More precisely, the patterns, as types, are found throughout Siberia and nearby; in Uralic, the morphemes marking them are generally native to Uralic. Uralic languages also have a number of traits that define what I call the Inner Asian type. These traits are not all typical of the North Pacific Rim or Siberian linguistic populations but rather are hallmarks of a smaller set of families: prototypically, Tungusic, Turkic, Mongolic, and Uralic or at least eastern Uralic; to a lesser extent, Korean, Japonic, and Eskimo-Aleut.}}</ref>
+This view has been corroborated by a number of scholars, including by a team around Grünthal et al. 2022. They presented evidence that the Proto-Uralic homeland was located east of the Urals, likely somewhere in [[Siberia]], and spoken by local hunter-gatherer communities. The [[Finno-Ugric languages|Finno-Ugric branch]] spread along rivers north- and westwards into the [[Volga region]], while the [[Samoyedic languages|Samoyedic branch]] headed north-and eastwards. The spread of Uralic languages may be in part due to the [[Seima-Turbino phenomenon]], although no direct link between them can be made yet. The material culture technology of proto-Uralic-speakers can be described as "Neolithic", as it included pottery but no vocabulary for food production. They further concluded that Proto-Uralic must have stood out of contact with Proto-Indo-European and did not share any genealogical ancestry with Indo-European, arguing that "whether based on cognacy or loans the argument from lexical resemblances is flawed", invalidating previous arguments for a homeland in the Volga region. Last, they noted that a number of traits of Uralic are "distinctive in western Eurasia. A number of typological properties are eastern-looking overall, fitting comfortably into northeast Asia, Siberia, or the North Pacific Rim", and are rare or absent in Europe, and that the Uralic languages "must have expanded via language shift", while "the shifting population had minimal impact on Proto-Uralic grammar and vocabulary".<ref>{{Cite journal |last1=Grünthal |first1=Riho |last2=Heyd |first2=Volker |last3=Holopainen |first3=Sampsa |last4=Janhunen |first4=Juha |last5=Khanina |first5=Olga |last6=Miestamo |first6=Matti |last7=Nichols |first7=Johanna |last8=Saarikivi |first8=Janne |last9=Sinnemäki |first9=Kaius |date=2022-08-29 |title=Drastic demographic events triggered the Uralic spread |url=https://researchportal.helsinki.fi/en/publications/drastic-demographic-events-triggered-the-uralic-spread |journal=Diachronica |language=English |volume=39 |issue=4 |pages=490–524 |doi=10.1075/dia.20038.gru |s2cid=248059749 |issn=0176-4225 |hdl=10138/347633 |hdl-access=free }}</ref><ref>{{Cite journal |last=Nichols |first=Johanna |date=2021-01-01 |title=The Origin and Dispersal of Uralic: Distributional Typological View |url=https://papers.ssrn.com/abstract=3778495 |journal=Annual Review of Linguistics |language=en |location=Rochester, NY |ssrn=3778495 |quote=PU had (and the modern Uralic languages mostly still have) a number of typological traits that are typical of the greater North Pacific Rim area and not of western Eurasia. These traits are shared with many Siberian languages but are (otherwise) rare in Europe. More precisely, the patterns, as types, are found throughout Siberia and nearby; in Uralic, the morphemes marking them are generally native to Uralic. Uralic languages also have a number of traits that define what I call the Inner Asian type. These traits are not all typical of the North Pacific Rim or Siberian linguistic populations but rather are hallmarks of a smaller set of families: prototypically, Tungusic, Turkic, Mongolic, and Uralic or at least eastern Uralic; to a lesser extent, Korean, Japonic, and Eskimo-Aleut.}}</ref>
-According to Bjørn Rasmus G. argues, based on linguistic influence from Uralic onto Tocharian, as well as loanwords from Indo-Iranian into Uralic, that the Proto-Uralic speakers likely resided in Southern Siberia, and may be associated with the local [[Okunev culture]] in the [[Altai Mountains|Altai region]].<ref>{{Cite journal |last=Bjørn |first=Rasmus G. |date=2022 |title=Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone |url=https://www.cambridge.org/core/journals/evolutionary-human-sciences/article/indoeuropean-loanwords-and-exchange-in-bronze-age-central-and-east-asia/91132871EAF737CD98BDFC43C16C761A |journal=Evolutionary Human Sciences |language=en |volume=4 |pages=e23 |doi=10.1017/ehs.2022.16 |issn=2513-843X |s2cid=248358873 |quote=Proto-Uralic was probably spoken in the Okunevo culture that constitutes the latest possible period for adoption of loanwords before the formation of the separate branches. The analysis also supports the proposition that an early stage of the Proto-Turkic language community was present around the Altai Mountains before 2000 BC, probably reflected in the eastern genetic component of Okunevo, and surely in the culturally transmitted loanwords of Indo-European provenance shared with early Uralic. |doi-access=free}}</ref>
+Rasmus G. Bjørn (2022), citing the previous findings, such as Grünthal et al. 2022, argues that the Proto-Uralic speakers likely resided in Southern Siberia, and may have been part of the local [[Okunev culture]] in the [[Altai Mountains|Altai region]], following the proposals of Janhunen and Peyrot. He notes that contact between Uralic and the Indo-European branches of Tocharian and Indo-Iranian at the one hand, and between Uralic and Turkic at the other hand, support a homeland near the [[Altai Mountains]]. An association with the [[Seima-Turbino phenomenon]] and the dispersal of Uralic languages "fits the string-like distribution of the western Uralic languages as well as close contacts with the Andronovo (and preceding Sintashta) culture associated with speakers of Indo-Iranian. Proto-Samoyedic was then either left in or migrated to the area around the [[Minusinsk basin|Minusinsk Basin]]". Bjørn attributes the "eastern lexical elements" to contact with [[Turkic languages|Turkic]], which radiated out from the Amur region and can be associated with Northeast Asian linguistic area.<ref>{{Cite journal |last=Bjørn |first=Rasmus G. |date=2022 |title=Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone |journal=Evolutionary Human Sciences |language=en |volume=4 |pages=e23 |doi=10.1017/ehs.2022.16 |issn=2513-843X |s2cid=248358873 |quote=Proto-Uralic was probably spoken in the Okunevo culture that constitutes the latest possible period for adoption of loanwords before the formation of the separate branches. The analysis also supports the proposition that an early stage of the Proto-Turkic language community was present around the Altai Mountains before 2000 BC, probably reflected in the eastern genetic component of Okunevo, and surely in the culturally transmitted loanwords of Indo-European provenance shared with early Uralic. |doi-access=free|pmid=37599704 |pmc=10432883 }}</ref>
+Jaakko Häkkinen (2023) argues that the location of Early Proto-Uralic and more distant stages are temporally irrelevant for the location of the Late Proto-Uralic homeland. He further elaborates that the "very distant Pre-Proto-Uralic" may have been spoken far from the region where Late Proto-Uralic was spoken. Häkkinen argues that Late Proto-Uralic and the successive stages of disintegration happened in the Central Ural Region, based on certain tree names and Indo-Iranian loanword layers. According to him, the disintegration of the proto-language began soon after 2500 BCE, but the speech community remained in a narrow region for a long time, until the second millennium BCE, after which the actual Uralic expansion began. On this basis the early phase of the Seima-Turbino Network in Southwestern Siberia could not yet be associated to the Uralic languages, but perhaps the later stages in Europe could. In any case, according to him, the arrival of the Uralic language in the Central Ural Region would precede even the early stage of the Seima-Turbino network.<ref>{{cite journal |last=Häkkinen |first=Jaakko |date=2023 |title=On Locating Proto-Uralic |journal=Finnisch-Ugrische Forschungen |volume=68 |issue=68 |pages=43–100 |doi=10.33339/fuf.120910|doi-access=free }}</ref>
 ==Evidence from population genetics==
-[[Vladimir Napolskikh]], who studied the origins of the "[[earth-diver]]" [[creation myth]]s, has concluded that a certain variety of those myths, which is found in the folklore of Uralic-speaking peoples and other N1, C3, and Q (Y-DNA) carrying populations, originated in [[Siberia]].<ref>Napolskikh V. V. (Izhevsk, Russia). [https://www.academia.edu/4918926/Diving_Bird_Myth_after_20_years_2012 Earth-Diver Myth (А812) in northern Eurasia and North America: twenty years later].</ref>
+[[Vladimir Napolskikh]], who studied the origins of the "[[earth-diver]]" [[creation myth]]s, has concluded that a certain variety of those myths, which is found in the folklore of [[Uralic peoples|Uralic-speaking peoples]] and other N1, C3, and Q (Y-DNA) carrying populations, originated in [[Siberia]].<ref>Napolskikh V. V. (Izhevsk, Russia). [https://www.academia.edu/4918926/Diving_Bird_Myth_after_20_years_2012 Earth-Diver Myth (А812) in northern Eurasia and North America: twenty years later].</ref>
+Genetic data suggests that early Uralic-speakers may be associated with hunter-gatherers in Western and Southern Siberia. These specific Uralic-speaking hunter-gatherers are argued to have formed from an admixture event between Western and Eastern Eurasian sources, with the calculated date of admixture is inferred to be ~7–9 kya. The Western source can be attributed to local [[Ancient North Eurasian|Ancient North Eurasians]], while the Eastern sources can be linked to "Neo-Siberians" (Eastern Siberian hunter-gatherers).<ref>{{Cite journal |last=Wong |first=Emily H.M. |date=2015 |title=Reconstructing genetic history of Siberian and Northeastern European populations |journal=Genome Research |volume=27 |issue=1 |pages=1–14 |doi=10.1101/gr.202945.115 |pmid=27965293 |pmc=5204334 |quote=Our results also provide insights for ongoing anthropological debates about the origin of indigenous populations in the Uralic region. Uralic people had been considered as mixed groups descended from the longtime admixture between Europeans and Asians. Our model of gene flows during the last 50,000 yr in Europe, East Asia, and Siberia (Fig. 8) provides support for Bunak's hypothesis since Western Siberians were associated with a distinct ancient lineage, possibly related to ANE people. At the same time, our models suggest a more complex scenario that included admixture with ancient Eastern Siberians who came into contact with proto Western Siberians ~7–9 kya.}}</ref>
+A number of population genomic studies in 2018 and 2019 note that the spread of Uralic languages may be associated with observed "Siberian" geneflow (represented by [[Nganasans]]) into the Eastern Baltic region.<ref>{{Cite journal |last1=Tambets |first1=Kristiina |last2=Metspalu |first2=Mait |last3=Lang |first3=Valter |last4=Villems |first4=Richard |last5=Kivisild |first5=Toomas |last6=Kriiska |first6=Aivar |last7=Thomas |first7=Mark G. |last8=Díez del Molino |first8=David |last9=Crema |first9=Enrico Ryunosuke |date=2019 |title=The Arrival of Siberian Ancestry Connecting the Eastern Baltic to Uralic Speakers further East |journal=Current Biology |language=en |volume=29 |issue=10 |pages=1701–1711.e16 |doi=10.1016/j.cub.2019.04.026 |issn=0960-9822 |pmc=6544527 |pmid=31080083}}</ref> The proto-Uralic languages may be associated with early "Neo-Siberian" expansions outgoing from [[Northeast Asia]] westwards (~6-11kya), which largely replaced previous "[[Paleosiberian languages|Paleosiberian groups]]", but predated the expansion of [[Turkic languages]].<ref>{{Cite journal |last1=Sikora |first1=Martin |last2=Pitulko |first2=Vladimir V. |last3=Sousa |first3=Vitor C. |last4=Allentoft |first4=Morten E. |last5=Vinner |first5=Lasse |last6=Rasmussen |first6=Simon |last7=Margaryan |first7=Ashot |last8=de Barros Damgaard |first8=Peter |last9=de la Fuente |first9=Constanza |last10=Renaud |first10=Gabriel |last11=Yang |first11=Melinda A. |last12=Fu |first12=Qiaomei |last13=Dupanloup |first13=Isabelle |last14=Giampoudakis |first14=Konstantinos |last15=Nogués-Bravo |first15=David |date=June 2019 |title=The population history of northeastern Siberia since the Pleistocene |url=https://www.nature.com/articles/s41586-019-1279-z |journal=Nature |language=en |volume=570 |issue=7760 |pages=182–188 |doi=10.1038/s41586-019-1279-z |issn=1476-4687 |quote=Most modern Siberian speakers of Neosiberian languages genetically fall on an East- West cline between Europeans and Early East Asians. Taking Even speakers as representatives, the Neosiberian turnover from the south, which largely replaced Ancient Paleosiberian ancestry, can be associated with the northward spread of Tungusic and probably also Turkic and Mongolic. However, the expansions of Tungusic as well as Turkic and Mongolic are too recent to be associable with the earliest waves of Neosiberian ancestry, dated later than ~11 kya, but discernible in the Baikal region from at least 6 kya onwards. Therefore, this phase of the Neosiberian population turnover must initially have transmitted other languages or language families into Siberia, including possibly Uralic and Yukaghir.|hdl=1887/3198847 |hdl-access=free }}</ref><ref>{{Cite journal |last1=Tambets |first1=Kristiina |last2=Yunusbayev |first2=Bayazit |last3=Hudjashov |first3=Georgi |last4=Ilumäe |first4=Anne-Mai |last5=Rootsi |first5=Siiri |last6=Honkola |first6=Terhi |last7=Vesakoski |first7=Outi |last8=Atkinson |first8=Quentin |last9=Skoglund |first9=Pontus |last10=Kushniarevich |first10=Alena |last11=Litvinov |first11=Sergey |last12=Reidla |first12=Maere |last13=Metspalu |first13=Ene |last14=Saag |first14=Lehti |last15=Rantanen |first15=Timo |date=2018-09-21 |title=Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations |journal=Genome Biology |volume=19 |issue=1 |pages=139 |doi=10.1186/s13059-018-1522-1 |issn=1474-760X |pmc=6151024 |pmid=30241495 |doi-access=free }}</ref>
-Genetic data suggests that early Uralic-speakers can be associated with hunter-gatherers in Western Siberia. These specific Uralic-speaking hunter-gatherers are argued to have formed from an admixture event between Western and Eastern Eurasian sources, with the calculated date of admixture is inferred to be ~7–9 kya.<ref>{{Cite journal |last=Wong |first=Emily H.M. |date=2015 |title=Reconstructing genetic history of Siberian and Northeastern European populations |journal=Genome Research |volume=27 |issue=1 |pages=1–14 |doi=10.1101/gr.202945.115 |pmid=27965293 |pmc=5204334 |quote=Our results also provide insights for ongoing anthropological debates about the origin of indigenous populations in the Uralic region. Uralic people had been considered as mixed groups descended from the longtime admixture between Europeans and Asians.  Our model of gene flows during the last 50,000 yr in Europe, East Asia, and Siberia (Fig. 8) provides support for Bunak's hypothesis since Western Siberians were associated with a distinct ancient lineage, possibly related to ANE people. At the same time, our models suggest a more complex scenario that included admixture with ancient Eastern Siberians who came into contact with proto Western Siberians ~7–9 kya.}}</ref>
-[[File:QpAdm-model_for_selected_Eurasian_populations.jpg|thumb|Estimated ancestry components among selected Eurasian populations, including Uralic-speaking groups. The yellow component represents Neo-Siberian ancestry (represented by [[Nganasan people|Nganasans]]), which is inferred to be associated with the spread of Uralic languages.<ref>{{Cite journal |last1=Jeong |first1=Choongwon |last2=Balanovsky |first2=Oleg |last3=Lukianova |first3=Elena |last4=Kahbatkyzy |first4=Nurzhibek |last5=Flegontov |first5=Pavel |last6=Zaporozhchenko |first6=Valery |last7=Immel |first7=Alexander |last8=Wang |first8=Chuan-Chao |last9=Ixan |first9=Olzhas |last10=Khussainova |first10=Elmira |last11=Bekmanov |first11=Bakhytzhan |last12=Zaibert |first12=Victor |last13=Lavryashina |first13=Maria |last14=Pocheshkhova |first14=Elvira |last15=Yusupov |first15=Yuldash |date=June 2019 |title=The genetic history of admixture across inner Eurasia |journal=Nature Ecology & Evolution |language=en |volume=3 |issue=6 |pages=966–976 |doi=10.1038/s41559-019-0878-2 |issn=2397-334X |pmc=6542712 |pmid=31036896}}</ref>]]
-A number of population genomic studies in 2018 and 2019 note that the spread of Uralic languages may be associated with observed "Eastern Siberian" geneflow (represented by [[Nganasans]]) into the Eastern Baltic region.<ref>{{Cite journal |last1=Tambets |first1=Kristiina |last2=Metspalu |first2=Mait |last3=Lang |first3=Valter |last4=Villems |first4=Richard |last5=Kivisild |first5=Toomas |last6=Kriiska |first6=Aivar |last7=Thomas |first7=Mark G. |last8=Díez del Molino |first8=David |last9=Crema |first9=Enrico Ryunosuke |date=2019 |title=The Arrival of Siberian Ancestry Connecting the Eastern Baltic to Uralic Speakers further East |journal=Current Biology |language=en |volume=29 |issue=10 |pages=1701–1711.e16 |doi=10.1016/j.cub.2019.04.026 |issn=0960-9822 |pmc=6544527 |pmid=31080083}}</ref> The proto-Uralic languages may be associated with early "Neo-Siberian" expansions outgoing from [[Northeast Asia]] westwards (~6-11kya), which largely replaced previous "[[Paleosiberian languages|Paleosiberian groups]]".<ref>{{Cite journal |last=Sikora |first=Martin |last2=Pitulko |first2=Vladimir V. |last3=Sousa |first3=Vitor C. |last4=Allentoft |first4=Morten E. |last5=Vinner |first5=Lasse |last6=Rasmussen |first6=Simon |last7=Margaryan |first7=Ashot |last8=de Barros Damgaard |first8=Peter |last9=de la Fuente |first9=Constanza |last10=Renaud |first10=Gabriel |last11=Yang |first11=Melinda A. |last12=Fu |first12=Qiaomei |last13=Dupanloup |first13=Isabelle |last14=Giampoudakis |first14=Konstantinos |last15=Nogués-Bravo |first15=David |date=June 2019 |title=The population history of northeastern Siberia since the Pleistocene |url=https://www.nature.com/articles/s41586-019-1279-z |journal=Nature |language=en |volume=570 |issue=7760 |pages=182–188 |doi=10.1038/s41586-019-1279-z |issn=1476-4687 |quote=Most modern Siberian speakers of Neosiberian languages genetically fall on an East- West cline between Europeans and Early East Asians. Taking Even speakers as representatives, the Neosiberian turnover from the south, which largely replaced Ancient Paleosiberian ancestry, can be associated with the northward spread of Tungusic and probably also Turkic and Mongolic. However, the expansions of Tungusic as well as Turkic and Mongolic are too recent to be associable with the earliest waves of Neosiberian ancestry, dated later than ~11 kya, but discernible in the Baikal region from at least 6 kya onwards. Therefore, this phase of the Neosiberian population turnover must initially have transmitted other languages or language families into Siberia, including possibly Uralic and Yukaghir.}}</ref><ref>{{Cite journal |last=Tambets |first=Kristiina |last2=Yunusbayev |first2=Bayazit |last3=Hudjashov |first3=Georgi |last4=Ilumäe |first4=Anne-Mai |last5=Rootsi |first5=Siiri |last6=Honkola |first6=Terhi |last7=Vesakoski |first7=Outi |last8=Atkinson |first8=Quentin |last9=Skoglund |first9=Pontus |last10=Kushniarevich |first10=Alena |last11=Litvinov |first11=Sergey |last12=Reidla |first12=Maere |last13=Metspalu |first13=Ene |last14=Saag |first14=Lehti |last15=Rantanen |first15=Timo |date=2018-09-21 |title=Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations |url=https://doi.org/10.1186/s13059-018-1522-1 |journal=Genome Biology |volume=19 |issue=1 |pages=139 |doi=10.1186/s13059-018-1522-1 |issn=1474-760X |pmc=6151024 |pmid=30241495}}</ref>
+[[File:Later_diffusion_and_contacts_in_Central_Asia.png|thumb|Cultural diffusion and contacts in Central Asia.<ref>{{cite journal |last1=Bjørn |first1=Rasmus G. |date=January 2022 |title=Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone |journal=Evolutionary Human Sciences |language=en |volume=4 |pages=e23 |doi=10.1017/ehs.2022.16 |issn=2513-843X|doi-access=free |pmid=37599704 |pmc=10432883 }}</ref>]]Rasmus G. Bjørn (2022) argues for a link between the Proto-Uralic-speakers and the pre-Indo-European [[Okunevo culture]], as well as to the [[Tarim mummies|Tarim EMBA cluster]]. He argues that following the expansion of Indo-Europeans eastwards and Northeast Asian (putative Turkic) groups westwards meeting in the Altai region, the early Uralic-speaking groups dispersed along the [[Seima-Turbino phenomenon|Seima-Turbino route]]. He also notes a correspondence between this expansion route and the frequency of haplogroup N.<ref name=":0">{{Cite journal |last=Bjørn |first=Rasmus G. |date=2022 |title=Indo-European loanwords and exchange in Bronze Age Central and East Asia: Six new perspectives on prehistoric exchange in the Eastern Steppe Zone |journal=Evolutionary Human Sciences |language=en |volume=4 |pages=e23 |doi=10.1017/ehs.2022.16 |issn=2513-843X|doi-access=free |pmid=37599704 |pmc=10432883 }}</ref>
+Peltola et al. 2023 reproduced the findings that modern Uralic-speaking populations display varying degrees of ancestry from a "Siberian" source maximized in modern day [[Nganasan people]]. Nganasans and a historical specimen from [[Bronze Age]] Southern Siberia (Krasnoyarsk_Krai_BA; kra001) were found to display the lowest f4 estimate for the eastern ancestry among Uralic-speaking populations, and represent a plausible source. In contrast, affinity for Ancient North Eurasian-rich ancestry (represented by [[Eastern Hunter-Gatherer|Eastern Hunter-Gatherers]]) was not significant and only observed among Western Finno-Ugric speakers.<ref>{{Cite journal |last1=Peltola |first1=Sanni |last2=Majander |first2=Kerttu |last3=Makarov |first3=Nikolaj |last4=Dobrovolskaya |first4=Maria |last5=Nordqvist |first5=Kerkko |last6=Salmela |first6=Elina |last7=Onkamo |first7=Päivi |date=2023-01-09 |title=Genetic admixture and language shift in the medieval Volga-Oka interfluve |journal=Current Biology |volume=33 |issue=1 |pages=174–182.e10 |doi=10.1016/j.cub.2022.11.036 |issn=0960-9822|doi-access=free }}</ref>
 ==See also==