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Coordinates: 57°30′N 11°00′W / 57.500°N 11.000°W / 57.500; -11.000[1]
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{{Good article}}
{{short description|A guyot in the Rockall Trough in the northeast Atlantic}}
{{Use mdy dates|date=February 2020}}
{{short description|Guyot in the Rockall Trough in the northeast Atlantic}}
{{Infobox Seamount
{{Infobox Seamount
| name = Anton Dohrn Seamount
| name = Anton Dohrn Seamount
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{{Use British English|date = February 2020}}
{{Use British English|date = February 2020}}
The '''Anton Dohrn Seamount''' is a [[guyot]] in the [[Rockall Trough]] in the northeast [[Atlantic]]. It is {{convert|1.8|km}} high and is topped with pinnacles, one of which reaches a depth of {{convert|530|m}}. Away from the flat top upon which the pinnacles rest, the slopes fall off steeply into the Rockall Trough and a moat in the sediment that surround the seamount.
The '''Anton Dohrn Seamount''' is a [[guyot]] in the [[Rockall Trough]] in the northeast [[Atlantic]]. It is {{convert|1.8|km}} high and is topped with pinnacles, one of which reaches a depth of {{convert|530|m}}. Away from the flat top upon which the pinnacles rest, the slopes fall off steeply into the Rockall Trough and a moat in the sediment that surrounds the seamount.


It appears to be a [[volcano]] formed by [[basalt]]ic [[lava]] and [[tuff]]. It formed during the [[Cretaceous]] and [[Paleogene]] and was proposed to be a source for [[bentonite]] layers across the [[British Isles]]. After the Cretaceous, subsidence and erosion lowered its top until it sank below sea level. The seamount was discovered in 1958.
It appears to be a [[volcano]] formed by [[basalt]]ic [[lava]] and [[tuff]]. It formed during the [[Cretaceous]] and [[Paleogene]] and was proposed to be a source for [[bentonite]] layers across the [[British Isles]]. After the Cretaceous, subsidence and erosion lowered its top until it sank below sea level. The seamount was discovered in 1958.


Anton Dohrn Seamount hosts a diverse ecosystem characterized by [[Bioherm|reefs]] formed by [[Cold-water corals|cold water corals]], [[sponge]]s and [[xenophyophora]]ns, which themselves host a number of animals. It has been impacted by human fishery however.
Anton Dohrn Seamount hosts a diverse ecosystem characterized by [[Bioherm|reefs]] formed by [[Cold-water corals|cold water corals]], [[sponge]]s and [[xenophyophora]]ns, which themselves host a number of animals. It has been affected by human fishing operations, however.


== Name and research history ==
== Name and research history ==


Anton Dohrn Seamount is also known as Anton Dohrn Kuppe, a name used by German charts,{{sfn|Rogalla|1962|p=60}} and as Anton Dohrn bank.<ref name="Bott1983" /> It was discovered in 22 September 1958 by the survey vessel ''Gauss'' during the [[Polarfront programme]] and later surveyed in 18–19 April 1959 by the fishery research vessel {{ship|FFS|Anton Dohrn}}.{{sfn|Rogalla|1962|p=59}}
Anton Dohrn Seamount is also known as Anton Dohrn Kuppe, a name used by German charts,{{sfn|Rogalla|1962|p=60}} and as Anton Dohrn bank.<ref name="Bott1983" /> It was discovered on 22 September 1958 by the survey vessel ''Gauss'' during the [[Polarfront programme]] and later surveyed on 18–19 April 1959 by the fishery research vessel {{ship|FFS|Anton Dohrn}}.{{sfn|Rogalla|1962|p=59}}


== Geography and geomorphology ==
== Geography and geomorphology ==


Anton Dohrn Seamount is located in the northeast [[Atlantic Ocean]] west of [[Scotland]],{{sfn|Stewart|Davies|Long|Strömberg|2009|p=10}} approximately halfway between [[St Kilda, Scotland|St Kilda]] ([[Hebrides]]) and [[Rockall]],{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} about {{convert|155|km}} west of the former.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} It lies in the [[Rockall Trough]], an over {{convert|2000|m}} submarine depression of unclear origin. North-northeast lies the [[Rosemary Bank]] and [[Hebrides Terrace Seamount]] is found south-southeast from the seamount.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} The seamount is located inside the [[exclusive economic zone]] of the United Kingdom.<ref name="Morato2013" />
Anton Dohrn Seamount is located in the northeast [[Atlantic Ocean]] west of [[Scotland]],{{sfn|Stewart|Davies|Long|Strömberg|2009|p=10}} approximately halfway between [[St Kilda, Scotland|St Kilda]] ([[Hebrides]]) and [[Rockall]],{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} about {{convert|155|km}} west of the former.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} It lies in the [[Rockall Trough]], an over {{convert|2000|m}} deep submarine depression of unclear origin. North-northeast lies the [[Rosemary Bank]] and [[Hebrides Terrace Seamount]] is found south-southeast from the seamount.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} The seamount is located inside the [[exclusive economic zone]] of the United Kingdom.<ref name="Morato2013" />


Anton Dohrn Seamount is a {{convert|1.8|km}} high{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} and about {{convert|45|km}}{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}}-{{convert|40|km}} wide circular{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} [[guyot]]{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} with a flat top at {{convert|1100|-|530|km}} depth.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=31}} Flat-topped seamounts are unusual in the North Atlantic.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} The shallowest point of the seamount lies at about {{convert|530|m}} depth{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} and is formed by a pinnacle that protrudes from the {{circa}} {{convert|600|m}} deep summit platform.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} A {{convert|100|m}} thick layer of sediment covers the flat top{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=244}} and appears to be reworked by storms and sea currents.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} Mounds,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=34}} slope breaks and other volcanic pinnacles are located on the flat top.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=14}} The seamount tilts southeastward.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=130}}
Anton Dohrn Seamount is a {{convert|1.8|km}} high{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} and about {{convert|45|km}}{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}}{{en dash}}{{convert|40|km}} wide circular{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} [[guyot]]{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} with a flat top at {{convert|1100|-|530|m}} depth.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=31}} Flat-topped seamounts are unusual in the North Atlantic.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} The shallowest point of the seamount lies at about {{convert|530|m}} depth{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} and is formed by a pinnacle that protrudes from the {{circa}} {{convert|600|m}} deep summit platform.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} A {{convert|100|m}} thick layer of sediment covers the flat top{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=244}} and appears to be reworked by storms and sea currents.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} Mounds,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=34}} slope breaks and other volcanic pinnacles are located on the flat top.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=14}} The seamount tilts southeastward.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=130}}


Beyond the margin of the flat top, the slopes of Anton Dohrn Seamount drop down to {{convert|2400|m}} depth.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=130}} The steep slopes have been variously described as either lacking a sediment cover{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=239}} or featuring [[gravel]]ly sediments along with outcropping bedrock.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} There are [[cliff]]s, [[ridge]]s{{sfn|Stewart|Davies|Long|Strömberg|2009|p=31}} and [[rockfall]]s{{sfn|Stewart|Davies|Long|Strömberg|2009|p=34}} but no gullies or canyons.<ref name="Morato2013" /> [[Parasitic cone]]s lie on the northwestern slope. A moat surrounds the seamount{{sfn|Stewart|Davies|Long|Strömberg|2009|p=14}} and reaches depths of about {{convert|2300|m}}.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} It might have formed either through erosion of surrounding sediments by ocean currents or through [[isostatic]] [[subsidence]].<ref name="HeezenJohnson1963" />
Beyond the margin of the flat top, the slopes of Anton Dohrn Seamount drop down to {{convert|2400|m}} depth.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=130}} The steep slopes have been variously described as either lacking a sediment cover{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=239}} or featuring [[gravel]]ly sediments along with outcropping bedrock.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} There are [[cliff]]s, [[ridge]]s{{sfn|Stewart|Davies|Long|Strömberg|2009|p=31}} and [[rockfall]]s{{sfn|Stewart|Davies|Long|Strömberg|2009|p=34}} but no gullies or canyons.<ref name="Morato2013" /> [[Parasitic cone]]s lie on the northwestern slope. A moat surrounds the seamount{{sfn|Stewart|Davies|Long|Strömberg|2009|p=14}} and reaches depths of about {{convert|2300|m}}.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} It might have formed either through erosion of surrounding sediments by ocean currents or through [[isostatic depression|isostatic]] [[subsidence]].<ref name="HeezenJohnson1963" />


== Geology ==
== Geology ==
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The [[Crust (geology)|crust]] underneath Anton Dohrn Seamount is much thinner than underneath the [[British Isles]] and the [[Rockall Plateau]] east and west of the seamount, respectively, and the [[Mohorovičić discontinuity]] is located at a shallower depth.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} It may be either stretched [[continental crust]] or [[oceanic crust]], and is covered by sediments.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=239}} At Anton Dohrn Seamount it appears to be unusually shallow, perhaps due to the [[Iceland plume]]'s buoyancy. The Iceland plume has uplifted terrain as far as {{convert|1000|km}} from the plume.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} A {{convert|100|km}} long crustal [[lineament]] known as the Anton Dohrn Lineament crosses through the seamount; it may extend into Scotland and Rockall Bank<ref name="Roberts2019" /> and runs in northwest–southeast direction.<ref name="Stoker2018" />
The [[Crust (geology)|crust]] underneath Anton Dohrn Seamount is much thinner than underneath the [[British Isles]] and the [[Rockall Plateau]] east and west of the seamount, respectively, and the [[Mohorovičić discontinuity]] is located at a shallower depth.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=238}} It may be either stretched [[continental crust]] or [[oceanic crust]], and is covered by sediments.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=239}} At Anton Dohrn Seamount it appears to be unusually shallow, perhaps due to the [[Iceland plume]]'s buoyancy. The Iceland plume has uplifted terrain as far as {{convert|1000|km}} from the plume.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} A {{convert|100|km}} long crustal [[lineament]] known as the Anton Dohrn Lineament crosses through the seamount; it may extend into Scotland and Rockall Bank<ref name="Roberts2019" /> and runs in northwest–southeast direction.<ref name="Stoker2018" />


Anton Dohrn Seamount is probably formed mostly by [[basalt]]ic [[lava]]{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} and [[tuff]]s{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} which define a transitional to alkaline suite.<ref name="Dietrich1980" /> The rocks contain [[feldspar]] and [[olivine]] [[phenocryst]]s as well as [[plagioclase]]. They are covered with [[ferromanganese]] crusts{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=240}} and vesicles contain [[carbonate]]s, [[clay]] and [[zeolite]]s which formed through alteration.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=241}} [[Chalk]]s of [[Maastricht]]ian age,{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=240}} [[Eocene]] nearshore conglomerates{{sfn|Stoker|Weering|Svaerdborg|2001|p=411}} and [[Miocene]] muds and sands have also been recovered.{{sfn|Stoker|Weering|Svaerdborg|2001|p=431}} A [[granite]] rock has been dredged as well; it may be a [[dropstone]] from [[iceberg]]s{{sfn|Rogalla|1962|p=62}} and such exotic rocks have been found in other dredge samples.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}}
Anton Dohrn Seamount is probably formed mostly by [[basalt]]ic [[lava]]{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} and [[tuff]]s{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} which define a transitional to alkaline [[suite (geology)|suite]].<ref name="Dietrich1980" /> The rocks contain [[feldspar]] and [[olivine]] [[phenocryst]]s as well as [[plagioclase]]. They are covered with [[ferromanganese]] crusts{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=240}} and vesicles contain [[carbonate]]s, [[clay]] and [[zeolite]]s which formed through alteration.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=241}} [[Chalk]]s of [[Maastricht]]ian age,{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=240}} [[Eocene]] nearshore conglomerates{{sfn|Stoker|Weering|Svaerdborg|2001|p=411}} and [[Miocene]] muds and sands have also been recovered.{{sfn|Stoker|Weering|Svaerdborg|2001|p=431}} A [[granite]] rock has been dredged as well; it may be a [[dropstone]] from [[iceberg]]s{{sfn|Rogalla|1962|p=62}} and such exotic rocks have been found in other dredge samples.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}}


== Geologic history ==
== Geologic history ==


Anton Dohrn Seamount is a former volcano.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} [[Radiometric dating]] of volcanic rocks dredged from it has yielded ages of 70 ± 1, 62 ± 1, 47 ± 1 and 41 ± 1 million years ago,{{sfn|O'Connor|Stoffers|Wijbrans|Shannon|2000|p=955}} indicating episodic activity over 29 million years.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} Pulses of volcanic activity of similar age have been identified at other volcanoes in the region and may reflect fluctuations of the [[Iceland plume]].{{sfn|O'Connor|Stoffers|Wijbrans|Shannon|2000|p=957}} The onset of volcanic activity may have been the consequence of crustal extension in the region.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} The activity during the [[Cretaceous]] implies that [[rift]]ing in the North Atlantic was already underway at that time.<ref name="GordonBrooks2016" /> At that time, the Rockall Trough was at least {{convert|1|km}} deep.<ref name="Scrutton1986" />
Anton Dohrn Seamount is a former volcano.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} [[Radiometric dating]] of volcanic rocks dredged from it has yielded ages of 70 ± 1, 62 ± 1, 47 ± 1 and 41 ± 1 million years ago,{{sfn|O'Connor|Stoffers|Wijbrans|Shannon|2000|p=955}} indicating episodic activity over 29 million years.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}} Pulses of volcanic activity of similar age have been identified at other volcanoes in the region and may reflect fluctuations of the Iceland plume.{{sfn|O'Connor|Stoffers|Wijbrans|Shannon|2000|p=957}} The onset of volcanic activity may have been the consequence of crustal extension in the region.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} The activity during the [[Cretaceous]] implies that [[rift]]ing in the North Atlantic was already underway at that time.<ref name="GordonBrooks2016" /> At that time, the Rockall Trough was at least {{convert|1|km}} deep.<ref name="Scrutton1986" />


[[Xenolith]]s found in volcanic rocks indicate that at Anton Dohrn volcanic activity involved interactions between [[magma]] and sediments, resulting in [[phreatomagmatic]] eruptions that could have dispersed [[volcanic ash]] in the region.{{sfn|Pacey|1984|p=57}} This volcanic ash erupted by Anton Dohrn may be the source of post-[[Cenomanian]] [[bentonite]]s of the [[British Isles]]{{sfn|Pacey|1984|p=58}} but the age and composition of the bentonites do not support this theory.<ref name="Wray1999" /> The seamount was once proposed to be the source of [[Turonian]] [[tephra]] deposits in Western Europe before its [[Maastrichtian]] age was established.<ref name="Wray1998" />
[[Xenolith]]s found in volcanic rocks indicate that at Anton Dohrn volcanic activity involved interactions between [[magma]] and sediments, resulting in [[phreatomagmatic]] eruptions that could have dispersed [[volcanic ash]] in the region.{{sfn|Pacey|1984|p=57}} This volcanic ash erupted by Anton Dohrn may be the source of post-[[Cenomanian]] [[bentonite]]s of the [[British Isles]]{{sfn|Pacey|1984|p=58}} but the age and composition of the bentonites do not support this theory.<ref name="Wray1999" /> The seamount was once proposed to be the source of [[Turonian]] [[tephra]] deposits in Western Europe before its [[Maastrichtian]] age was established.<ref name="Wray1998" />


During the [[Cretaceous]] the seamount was about {{convert|2|km}} higher than present,{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} perhaps even reaching {{convert|2000|m}} height above sea level;{{sfn|Jones|Ramsay|Preston|Smith|1974|p=130}} presumably it was then eroded during the [[Paleocene]] when a wave of erosion took place in western [[Great Britain|Britain]] and stripped much of the volcanic centres of northwest Scotland.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} An episode of crustal [[subsidence]] in the [[Cretaceous]]-[[Oligocene]] also played a role in lowering Anton Dohrn Seamount.{{sfn|Jones|Ramsay|Preston|Smith|1974|pp=130-131}} The pinnacles on the seamount may be leftover volcanic conduits that resisted erosion.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} Sedimentation covered the seamount and its flanks in the [[Eocene]] and continued afterwards.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}}
During the [[Cretaceous]] the seamount was about {{convert|2|km}} higher than present,{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} perhaps even reaching {{convert|2000|m}} height above sea level;{{sfn|Jones|Ramsay|Preston|Smith|1974|p=130}} presumably it was then eroded during the [[Paleocene]] when a wave of erosion took place in western [[Great Britain|Britain]] and stripped much of the volcanic centres of northwest Scotland.{{sfn|Jones|Siddall|Thirlwall|Chroston|1994|p=245}} An episode of crustal [[subsidence]] in the [[Cretaceous]]-[[Oligocene]] also played a role in lowering Anton Dohrn Seamount.{{sfn|Jones|Ramsay|Preston|Smith|1974|pp=130{{endash}}131}} The pinnacles on the seamount may be leftover volcanic conduits that resisted erosion.{{sfn|Jones|Ramsay|Preston|Smith|1974|p=129}} Sedimentation covered the seamount and its flanks in the Eocene and continued afterwards.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=12}}


== Ecology ==
== Ecology ==


[[Barnacle]]s and [[brachiopod]]s grow on the top of the seamount, and [[echinoderm]]s,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=13}} [[coral]]s<ref name="Wilson1979" /> and [[cirripede]]s also occur there.<ref name="Gage1986" /> On the sandy or gravelly substrate [[serpulid]]s and [[sponge]]s are found.<ref name="Morato2013" /> The seamount may be a [[shark]] nursery.<ref name="Moore2013" /> Finally, the [[bivalve]] ''[[Xylophaga anselli]]'' was found at Anton Dohrn Seamount and the [[Hebrides]] slope.<ref name="Romano2014" />
[[Barnacle]]s and [[brachiopod]]s grow on the top of the seamount, and [[echinoderm]]s,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=13}} [[coral]]s<ref name="Wilson1979" /> and [[cirripede]]s also occur there.<ref name="Gage1986" /> On the sandy or gravelly substrate [[serpulid]]s and [[sponge]]s are found.<ref name="Morato2013" /> The seamount may be a [[shark]] nursery.<ref name="Moore2013" /> Finally, the [[bivalve]] ''[[Xylophaga anselli]]'' has been found at Anton Dohrn Seamount and the [[Hebrides]] slope.<ref name="Romano2014" />


A number of ecosystems have been found on Anton Dohrn Seamount, including [[coral garden]]s, [[Deep-water coral|cold water coral reefs]] and [[sponge]] and [[xenophyophore]] communities;{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=14-15}}{{sfn|Henry|Vad|Findlay|Murillo|2014|p=1}} this seamount is the first place in the United Kingdom where coral gardens have been discovered.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=24}} The sandy and cobbly terrain of the slopes with occasional bedrock outcrops is populated by reefs that grow on bedrock or on cobbles.{{sfn|Stewart|Davies|Long|Strömberg|2009|pp=38-46}} They mostly occur on the sides of the seamount, on mounds on the flat top{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=16}} and its margin,<ref name="Morato2013" /> perhaps for hydrodynamic reasons or because substrates favourable for the development of the reefs are found there.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=26-27}} There is a vertical stratification, with ''[[Lophelia]]'' found at shallower depths than ''[[Solenosmilia]]''.<ref name="HenryRoberts2014" /> Corals such as [[antipatharian]]s like ''[[Leiopathes]] sp.'', small [[bamboo coral]]s, large [[gorgonian]]s and [[soft coral]]s like as ''[[Anthomastus]] sp.'' have also been found at [[parasitic vent]]s.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} The [[cold water coral]] cover can become so thick that the underground disappears underneath it.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=30}}
A number of ecosystems have been found on Anton Dohrn Seamount, including [[coral garden]]s, [[Deep-water coral|cold water coral reefs]] and sponge and [[xenophyophore]] communities;{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=14—15}}{{sfn|Henry|Vad|Findlay|Murillo|2014|p=1}} this seamount is the first place in the United Kingdom where coral gardens have been discovered.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=24}} The sandy and cobbly terrain of the slopes with occasional bedrock outcrops is populated by reefs that grow on bedrock or on cobbles.{{sfn|Stewart|Davies|Long|Strömberg|2009|pp=38—46}} They mostly occur on the sides of the seamount, on mounds on the flat top{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=16}} and its margin,<ref name="Morato2013" /> perhaps for hydrodynamic reasons or because substrates favourable for the development of the reefs are found there.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=26—27}} There is a vertical stratification, with ''[[Lophelia]]'' found at shallower depths than ''[[Solenosmilia]]''.<ref name="HenryRoberts2014" /> Corals such as [[antipatharian]]s like ''[[Leiopathes]] sp.'', small [[bamboo coral]]s, large [[gorgonian]]s and [[soft coral]]s like as ''[[Anthomastus]] sp.'' have also been found at parasitic vents.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=9}} The cold water coral cover can become so thick that the underground disappears underneath it.{{sfn|Stewart|Davies|Long|Strömberg|2009|p=30}}


[[Dropframe camera]] surveys{{sfn|Henry|Vad|Findlay|Murillo|2014|p=1}} have seen [[anemone]]s, [[anthozoan]]s, [[ascidian]]s, the [[Asteroidea|asteroid]] (starfish) ''[[Henricia]] sp.'', [[bamboo coral]]s, [[Caryophylliidae|caryophyllids]], [[Cerianthidae|cerianthids]], [[antipatharian]] [[coral]]s with various shapes, the corals ''[[Desmophyllum dianthus]]'', ''[[Lophelia pertusa]]'' and ''[[Solenosmilia variabilis]]'', [[echinoderm]]s including [[Brisingida|brisingids]] and [[crinoid]]s, [[glass sponge]]s, [[gorgonian]]s, [[holothurian]]s, the [[ophiuroid]]s ''[[Ophiactis balli]]'' and ''[[Ophiomusium lymani]]'', the [[pencil urchin]] ''[[Cidaris cidaris]]'', [[pycnogonid]]s, the [[scleractinia]]n ''[[Madrepora oculata]]'', the [[seapen]] ''[[Pennatula phosphorea]]'', [[sea urchin]]s, [[sea whip]]s, [[serpulid]]s, [[soft coral]]s such as ''[[Gersemia]] sp.'' and ''[[Anthomastus]] sp.'', lobose, large and encrusting [[sponge]]s, [[Stylasteridae|stylasterids]] and [[xenophyophore]]s. [[Decapod]]s, fish including ''[[Lepidion eques]]'', the eel ''[[Synaphobranchus kaupi]]'' and [[squat lobster]]s ''[[Munida]] sp.'' have also been encountered.{{sfn|Stewart|Davies|Long|Strömberg|2009|pp=38-46}}{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=12-13}}<ref name="Copard2012" />
[[Dropframe camera]] surveys{{sfn|Henry|Vad|Findlay|Murillo|2014|p=1}} have seen [[anemone]]s, [[anthozoan]]s, [[ascidian]]s, the [[Asteroidea|asteroid]] (starfish) ''[[Henricia]] sp.'', bamboo corals, [[Caryophylliidae|caryophyllids]], [[Cerianthidae|cerianthids]], [[antipatharian]] [[coral]]s with various shapes, the corals ''[[Desmophyllum dianthus]]'', ''[[Lophelia pertusa]]'' and ''[[Solenosmilia variabilis]]'', [[echinoderm]]s including [[Brisingida|brisingids]] and [[crinoid]]s, [[glass sponge]]s, [[gorgonian]]s, [[holothurian]]s, the [[ophiuroid]]s ''[[Ophiactis balli]]'' and ''[[Ophiomusium lymani]]'', the [[pencil urchin]] ''[[Cidaris cidaris]]'', [[pycnogonid]]s, the [[scleractinia]]n ''[[Madrepora oculata]]'', the [[seapen]] ''[[Pennatula phosphorea]]'', [[sea urchin]]s, [[sea whip]]s, [[serpulid]]s, soft corals such as ''[[Gersemia]] sp.'' and ''[[Anthomastus]] sp.'', lobose, large and encrusting sponges, [[Stylasteridae|stylasterids]] and [[xenophyophore]]s. [[Decapoda|Decapods]], fish including ''[[Lepidion eques]]'', the eel ''[[Synaphobranchus kaupi]]'' and [[squat lobster]]s ''[[Munida]] sp.'' have also been encountered.{{sfn|Stewart|Davies|Long|Strömberg|2009|pp=38—46}}{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|pp=12—13}}<ref name="Copard2012" />


Seamounts are considered to be [[biodiversity hotspot]]s,{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=2}} and there are proposals to make Anton Dohrn Seamount a [[Special Area of Conservation]].<ref name="Morato2013" /> The region is considered to be "the cradle of deep-sea biology" as [[Victorian era|Victorian]]-era scientists sampled the regional fauna.<ref name="Morato2013" /> [[Ocean current]]s around Anton Dohrn Seamount are complicated and formed by various water masses.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} [[Internal tide]]s at the seamount appear to be important for its ecosystem.{{sfn|Henry|Vad|Findlay|Murillo|2014|p=5}}
Seamounts are considered to be [[biodiversity hotspot]]s,{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=2}} and there are proposals to make Anton Dohrn Seamount a [[Special Area of Conservation]].<ref name="Morato2013" /> The region is considered to be "the cradle of deep-sea biology" as [[Victorian era|Victorian]]-era scientists sampled the regional fauna.<ref name="Morato2013" /> [[Ocean current]]s around Anton Dohrn Seamount are complicated and formed by various water masses.{{sfn|Davies|Stewart|Narayanaswamy|Jacobs|2015|p=4}} [[Internal tide]]s at the seamount appear to be important for its ecosystem.{{sfn|Henry|Vad|Findlay|Murillo|2014|p=5}}


The seamount has been impacted by [[deep water fishing]].<ref name="Morato2013" /> Lost [[fishing gear]] and [[trawl mark]]s have been found on Anton Dohrn Seamount,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=74}} and animals found at its foot have ingested [[microplastic]]s.<ref name="Courtene-Jones2017" />
The seamount has been impacted by [[deep water fishing]].<ref name="Morato2013" /> Lost [[fishing gear]] and [[trawl mark]]s have been found on Anton Dohrn Seamount,{{sfn|Stewart|Davies|Long|Strömberg|2009|p=74}} and animals found at its foot have ingested [[microplastic]]s.<ref name="Courtene-Jones2017" /> In October 2020 the seamount was made part of the [[West of Scotland Marine Protected Area]] by the [[Scottish Government]] in attempt to protect the area's ecology.<ref name="BBC" /><ref name="JNCC" />


==References==
==References==
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<ref name="Roberts2019">{{cite journal |last1=Roberts |first1=Alan M. |last2=Alvey |first2=Andrew D. |last3=Kusznir |first3=Nick J. |title=Crustal structure and heat-flow history in the UK Rockall Basin, derived from backstripping and gravity-inversion analysis |journal=Petroleum Geoscience |date=May 1, 2019 |volume=25 |issue=2 |page=146 |doi=10.1144/petgeo2017-063 |s2cid=135171946 |language=en |issn=1354-0793|doi-access=free |bibcode=2019PetGe..25..131R }}</ref>
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<ref name="HenryRoberts2014">{{cite journal |last1=Henry |first1=Lea-Anne |last2=Roberts |first2=J. Murray |title=Recommendations for best practice in deep-sea habitat classification: Bullimore et al. as a case study |journal=ICES Journal of Marine Science |date=June 1, 2014 |volume=71 |issue=4 |pages=896–897 |doi=10.1093/icesjms/fst175 |language=en |issn=1054-3139|doi-access=free }}</ref>
<ref name="Courtene-Jones2017">{{cite journal |last1=Courtene-Jones |first1=Winnie |last2=Quinn |first2=Brian |last3=Gary |first3=Stefan F. |last4=Mogg |first4=Andrew O. M. |last5=Narayanaswamy |first5=Bhavani E. |title=Microplastic pollution identified in deep-sea water and ingested by benthic invertebrates in the Rockall Trough, North Atlantic Ocean |journal=Environmental Pollution |date=December 1, 2017 |volume=231 |issue=Pt 1 |pages=271–272 |doi=10.1016/j.envpol.2017.08.026 |pmid=28806692 |bibcode=2017EPoll.231..271C |s2cid=3355997 |language=en |issn=0269-7491 |url=https://myresearchspace.uws.ac.uk/ws/files/1868503/Accepted_Author_Manuscript.pdf |access-date=March 15, 2020 |archive-date=March 5, 2020 |archive-url=https://web.archive.org/web/20200305154248/https://myresearchspace.uws.ac.uk/ws/files/1868503/Accepted_Author_Manuscript.pdf |url-status=dead }}</ref>
<ref name="Morato2013">{{cite journal |last1=Morato |first1=T. |last2=Kvile |first2=K. Ø |last3=Taranto |first3=G. H. |last4=Tempera |first4=F. |last5=Narayanaswamy |first5=B. E. |last6=Hebbeln |first6=D. |last7=Menezes |first7=G. M. |last8=Wienberg |first8=C. |last9=Santos |first9=R. S. |last10=Pitcher |first10=T. J. |title=Seamount physiography and biology in the north-east Atlantic and Mediterranean Sea |journal=Biogeosciences |date=May 6, 2013 |volume=10 |issue=5 |page=3047 |doi=10.5194/bg-10-3039-2013 |bibcode=2013BGeo...10.3039M |language=English |issn=1726-4170|doi-access=free }}</ref>
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{{refend}}
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==External links==
==External links==
*[http://archive.jncc.gov.uk/default.aspx?page=6527 Anton Dohrn Seamount MPA]
*[https://jncc.gov.uk/our-work/anton-dohrn-seamount-mpa/ Anton Dohrn Seamount MPA]
*[https://web.archive.org/web/20090206163100/http://ngo.grida.no/wwfneap/Projects/Reports/Seamount_Report.pdf WWF report on seamounts of the Northeast Atlantic] ([[PDF]])
*[https://web.archive.org/web/20090206163100/http://ngo.grida.no/wwfneap/Projects/Reports/Seamount_Report.pdf WWF report on seamounts of the Northeast Atlantic] ([[PDF]])
*[https://www.bbc.co.uk/news/science-environment-36806038 BBC webpage report on the 2016 report on coral reef habitat]
*[https://www.bbc.co.uk/news/science-environment-36806038 BBC webpage report on the 2016 report on coral reef habitat]

Latest revision as of 15:03, 26 August 2024

Anton Dohrn Seamount
Anton Dohrn Seamount is located in Oceans around British Isles
Anton Dohrn Seamount
Anton Dohrn Seamount is located in North Atlantic
Anton Dohrn Seamount
Anton Dohrn Seamount (North Atlantic)
Summit depth600 metres
Height1,500 m
Location
LocationNorth Atlantic Ocean
Coordinates57°30′N 11°00′W / 57.500°N 11.000°W / 57.500; -11.000[1]
CountryUnited Kingdom (EEZ)
Geology
TypeGuyot
Last eruption~40 million years

The Anton Dohrn Seamount is a guyot in the Rockall Trough in the northeast Atlantic. It is 1.8 kilometres (1.1 mi) high and is topped with pinnacles, one of which reaches a depth of 530 metres (1,740 ft). Away from the flat top upon which the pinnacles rest, the slopes fall off steeply into the Rockall Trough and a moat in the sediment that surrounds the seamount.

It appears to be a volcano formed by basaltic lava and tuff. It formed during the Cretaceous and Paleogene and was proposed to be a source for bentonite layers across the British Isles. After the Cretaceous, subsidence and erosion lowered its top until it sank below sea level. The seamount was discovered in 1958.

Anton Dohrn Seamount hosts a diverse ecosystem characterized by reefs formed by cold water corals, sponges and xenophyophorans, which themselves host a number of animals. It has been affected by human fishing operations, however.

Name and research history

[edit]

Anton Dohrn Seamount is also known as Anton Dohrn Kuppe, a name used by German charts,[2] and as Anton Dohrn bank.[3] It was discovered on 22 September 1958 by the survey vessel Gauss during the Polarfront programme and later surveyed on 18–19 April 1959 by the fishery research vessel FFS Anton Dohrn.[4]

Geography and geomorphology

[edit]

Anton Dohrn Seamount is located in the northeast Atlantic Ocean west of Scotland,[5] approximately halfway between St Kilda (Hebrides) and Rockall,[6] about 155 kilometres (96 mi) west of the former.[7] It lies in the Rockall Trough, an over 2,000 metres (6,600 ft) deep submarine depression of unclear origin. North-northeast lies the Rosemary Bank and Hebrides Terrace Seamount is found south-southeast from the seamount.[6] The seamount is located inside the exclusive economic zone of the United Kingdom.[8]

Anton Dohrn Seamount is a 1.8 kilometres (1.1 mi) high[9] and about 45 kilometres (28 mi)[10]–40 kilometres (25 mi) wide circular[9] guyot[6] with a flat top at 1,100–530 metres (3,610–1,740 ft) depth.[11] Flat-topped seamounts are unusual in the North Atlantic.[12] The shallowest point of the seamount lies at about 530 metres (1,740 ft) depth[9] and is formed by a pinnacle that protrudes from the c. 600 metres (2,000 ft) deep summit platform.[10] A 100 metres (330 ft) thick layer of sediment covers the flat top[13] and appears to be reworked by storms and sea currents.[9] Mounds,[14] slope breaks and other volcanic pinnacles are located on the flat top.[15] The seamount tilts southeastward.[16]

Beyond the margin of the flat top, the slopes of Anton Dohrn Seamount drop down to 2,400 metres (7,900 ft) depth.[17] The steep slopes have been variously described as either lacking a sediment cover[1] or featuring gravelly sediments along with outcropping bedrock.[7] There are cliffs, ridges[11] and rockfalls[14] but no gullies or canyons.[8] Parasitic cones lie on the northwestern slope. A moat surrounds the seamount[15] and reaches depths of about 2,300 metres (7,500 ft).[10] It might have formed either through erosion of surrounding sediments by ocean currents or through isostatic subsidence.[18]

Geology

[edit]

The crust underneath Anton Dohrn Seamount is much thinner than underneath the British Isles and the Rockall Plateau east and west of the seamount, respectively, and the Mohorovičić discontinuity is located at a shallower depth.[6] It may be either stretched continental crust or oceanic crust, and is covered by sediments.[1] At Anton Dohrn Seamount it appears to be unusually shallow, perhaps due to the Iceland plume's buoyancy. The Iceland plume has uplifted terrain as far as 1,000 kilometres (620 mi) from the plume.[19] A 100 kilometres (62 mi) long crustal lineament known as the Anton Dohrn Lineament crosses through the seamount; it may extend into Scotland and Rockall Bank[20] and runs in northwest–southeast direction.[21]

Anton Dohrn Seamount is probably formed mostly by basaltic lava[10] and tuffs[12] which define a transitional to alkaline suite.[22] The rocks contain feldspar and olivine phenocrysts as well as plagioclase. They are covered with ferromanganese crusts[23] and vesicles contain carbonates, clay and zeolites which formed through alteration.[24] Chalks of Maastrichtian age,[23] Eocene nearshore conglomerates[25] and Miocene muds and sands have also been recovered.[26] A granite rock has been dredged as well; it may be a dropstone from icebergs[27] and such exotic rocks have been found in other dredge samples.[12]

Geologic history

[edit]

Anton Dohrn Seamount is a former volcano.[9] Radiometric dating of volcanic rocks dredged from it has yielded ages of 70 ± 1, 62 ± 1, 47 ± 1 and 41 ± 1 million years ago,[28] indicating episodic activity over 29 million years.[9] Pulses of volcanic activity of similar age have been identified at other volcanoes in the region and may reflect fluctuations of the Iceland plume.[29] The onset of volcanic activity may have been the consequence of crustal extension in the region.[12] The activity during the Cretaceous implies that rifting in the North Atlantic was already underway at that time.[30] At that time, the Rockall Trough was at least 1 kilometre (0.62 mi) deep.[31]

Xenoliths found in volcanic rocks indicate that at Anton Dohrn volcanic activity involved interactions between magma and sediments, resulting in phreatomagmatic eruptions that could have dispersed volcanic ash in the region.[32] This volcanic ash erupted by Anton Dohrn may be the source of post-Cenomanian bentonites of the British Isles[33] but the age and composition of the bentonites do not support this theory.[34] The seamount was once proposed to be the source of Turonian tephra deposits in Western Europe before its Maastrichtian age was established.[35]

During the Cretaceous the seamount was about 2 kilometres (1.2 mi) higher than present,[19] perhaps even reaching 2,000 metres (6,600 ft) height above sea level;[16] presumably it was then eroded during the Paleocene when a wave of erosion took place in western Britain and stripped much of the volcanic centres of northwest Scotland.[19] An episode of crustal subsidence in the Cretaceous-Oligocene also played a role in lowering Anton Dohrn Seamount.[36] The pinnacles on the seamount may be leftover volcanic conduits that resisted erosion.[12] Sedimentation covered the seamount and its flanks in the Eocene and continued afterwards.[9]

Ecology

[edit]

Barnacles and brachiopods grow on the top of the seamount, and echinoderms,[37] corals[38] and cirripedes also occur there.[39] On the sandy or gravelly substrate serpulids and sponges are found.[8] The seamount may be a shark nursery.[40] Finally, the bivalve Xylophaga anselli has been found at Anton Dohrn Seamount and the Hebrides slope.[41]

A number of ecosystems have been found on Anton Dohrn Seamount, including coral gardens, cold water coral reefs and sponge and xenophyophore communities;[42][43] this seamount is the first place in the United Kingdom where coral gardens have been discovered.[44] The sandy and cobbly terrain of the slopes with occasional bedrock outcrops is populated by reefs that grow on bedrock or on cobbles.[45] They mostly occur on the sides of the seamount, on mounds on the flat top[46] and its margin,[8] perhaps for hydrodynamic reasons or because substrates favourable for the development of the reefs are found there.[47] There is a vertical stratification, with Lophelia found at shallower depths than Solenosmilia.[48] Corals such as antipatharians like Leiopathes sp., small bamboo corals, large gorgonians and soft corals like as Anthomastus sp. have also been found at parasitic vents.[7] The cold water coral cover can become so thick that the underground disappears underneath it.[49]

Dropframe camera surveys[43] have seen anemones, anthozoans, ascidians, the asteroid (starfish) Henricia sp., bamboo corals, caryophyllids, cerianthids, antipatharian corals with various shapes, the corals Desmophyllum dianthus, Lophelia pertusa and Solenosmilia variabilis, echinoderms including brisingids and crinoids, glass sponges, gorgonians, holothurians, the ophiuroids Ophiactis balli and Ophiomusium lymani, the pencil urchin Cidaris cidaris, pycnogonids, the scleractinian Madrepora oculata, the seapen Pennatula phosphorea, sea urchins, sea whips, serpulids, soft corals such as Gersemia sp. and Anthomastus sp., lobose, large and encrusting sponges, stylasterids and xenophyophores. Decapods, fish including Lepidion eques, the eel Synaphobranchus kaupi and squat lobsters Munida sp. have also been encountered.[45][50][51]

Seamounts are considered to be biodiversity hotspots,[52] and there are proposals to make Anton Dohrn Seamount a Special Area of Conservation.[8] The region is considered to be "the cradle of deep-sea biology" as Victorian-era scientists sampled the regional fauna.[8] Ocean currents around Anton Dohrn Seamount are complicated and formed by various water masses.[10] Internal tides at the seamount appear to be important for its ecosystem.[53]

The seamount has been impacted by deep water fishing.[8] Lost fishing gear and trawl marks have been found on Anton Dohrn Seamount,[54] and animals found at its foot have ingested microplastics.[55] In October 2020 the seamount was made part of the West of Scotland Marine Protected Area by the Scottish Government in attempt to protect the area's ecology.[56][57]

References

[edit]
  1. ^ a b c Jones et al. 1994, p. 239.
  2. ^ Rogalla 1962, p. 60.
  3. ^ Bott, Martin H. P.; Saxov, Svend; Talwani, Manik; Thiede, Jörn, eds. (1983). Structure and Development of the Greenland-Scotland Ridge. Boston, MA: Springer US. p. 679. doi:10.1007/978-1-4613-3485-9. ISBN 978-1-4613-3487-3.
  4. ^ Rogalla 1962, p. 59.
  5. ^ Stewart et al. 2009, p. 10.
  6. ^ a b c d Jones et al. 1994, p. 238.
  7. ^ a b c Stewart et al. 2009, p. 9.
  8. ^ a b c d e f g Morato, T.; Kvile, K. Ø; Taranto, G. H.; Tempera, F.; Narayanaswamy, B. E.; Hebbeln, D.; Menezes, G. M.; Wienberg, C.; Santos, R. S.; Pitcher, T. J. (May 6, 2013). "Seamount physiography and biology in the north-east Atlantic and Mediterranean Sea". Biogeosciences. 10 (5): 3047. Bibcode:2013BGeo...10.3039M. doi:10.5194/bg-10-3039-2013. ISSN 1726-4170.
  9. ^ a b c d e f g Stewart et al. 2009, p. 12.
  10. ^ a b c d e Davies et al. 2015, p. 4.
  11. ^ a b Stewart et al. 2009, p. 31.
  12. ^ a b c d e Jones et al. 1974, p. 129.
  13. ^ Jones et al. 1994, p. 244.
  14. ^ a b Stewart et al. 2009, p. 34.
  15. ^ a b Stewart et al. 2009, p. 14.
  16. ^ a b Jones et al. 1974, p. 130.
  17. ^ Stewart et al. 2009, p. 130.
  18. ^ Heezen, Bruce C.; Johnson, G. Leonard (November 1, 1963). "A moated knoll in the Canary Passage". Deutsche Hydrografische Zeitschrift. 16 (6): 270. Bibcode:1963DeHyZ..16..269H. doi:10.1007/BF02025932. ISSN 1616-7228.
  19. ^ a b c Jones et al. 1994, p. 245.
  20. ^ Roberts, Alan M.; Alvey, Andrew D.; Kusznir, Nick J. (May 1, 2019). "Crustal structure and heat-flow history in the UK Rockall Basin, derived from backstripping and gravity-inversion analysis". Petroleum Geoscience. 25 (2): 146. Bibcode:2019PetGe..25..131R. doi:10.1144/petgeo2017-063. ISSN 1354-0793. S2CID 135171946.
  21. ^ Stoker, Martyn S.; Holford, Simon P.; Hillis, Richard R. (March 1, 2018). "A rift-to-drift record of vertical crustal motions in the Faroe–Shetland Basin, NW European margin: establishing constraints on NE Atlantic evolution". Journal of the Geological Society. 175 (2): 270. Bibcode:2018JGSoc.175..263S. doi:10.1144/jgs2017-076. ISSN 0016-7649. S2CID 133667534.
  22. ^ Dietrich, V. J; Jones, E. J. W (April 1, 1980). "Volcanic rocks from Rosemary Bank (Rockall Trough, NE Atlantic)". Marine Geology. 35 (4): 295. Bibcode:1980MGeol..35..287D. doi:10.1016/0025-3227(80)90122-X. ISSN 0025-3227.
  23. ^ a b Jones et al. 1994, p. 240.
  24. ^ Jones et al. 1994, p. 241.
  25. ^ Stoker, Weering & Svaerdborg 2001, p. 411.
  26. ^ Stoker, Weering & Svaerdborg 2001, p. 431.
  27. ^ Rogalla 1962, p. 62.
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