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{{Short description|Pennsylvanian-age geological unit}}
{{Infobox rockunit
{{Infobox rockunit
| name = {{pagename}}
| name = Allegheny Group
| image = Fossiliferous bituminous coal (Clarion Coal, Middle Pennsylvanian; Prattsville West Roadcut, Vinton County, Ohio, USA) 1.jpg
| image =
| caption =
| caption = [[Bituminous coal]] (Clarion Coal) from the Allegheny Group, Ohio
| type = sedimentary
| type = Sedimentary
| age = [[Pennsylvanian]]
| age = ([[Moscovian (Carboniferous)|Moscovian]]<nowiki>)</nowiki><br />~{{Fossil range|311|306.8}}
| period = Pennsylvanian
| prilithology = [[sandstone]], [[coal]]
| prilithology = [[Sandstone]], [[coal]]
| otherlithology =
| otherlithology =
| namedfor =
| namedfor =
| namedby = [[Henry Darwin Rogers|H. D. Rogers]], 1840<ref>Rogers, H.D., (1840), Fourth annual report of the Geological Survey of the State of Pennsylvania: Pennsylvania Geological Survey Annual Report, no. 4, 215 p. ([https://books.google.com/books?id=AJ44AQAAMAAJ])</ref>
| namedby =
| region = [[Appalachian Mountains]]
| region = [[Appalachian Mountains]]
| country =
| country = United States
| coordinates =
| coordinates =
| unitof =
| unitof =
| subunits =
| subunits =[[Freeport Formation]]<BR>
: [[Upper Freeport Coal]]
| underlies = [[Catskill Formation]]
: [[Upper Freeport Limestone Member]]
| overlies = [[Pottsville Formation]]
: [[Butler Sandstone Member]]
: [[Lower Freeport Coal]]
: [[Freeport Sandstone Member]]

[[Kittanning Formation]]<BR>
: [[Upper Kittanning Coal]]
: [[Johnstone Limestone Member]]
: [[Middle Kittanning Coal]]
: [[Lower Kittanning Coal]]
: [[Kittanning Sandstone Member]]
: [[Vanport Limestone Member]]
| underlies = [[Conemaugh Group]]
| overlies = [[Kanawha Formation]] and [[Pottsville Formation]]
| thickness =
| thickness =
| extent = Pennsylvania, Maryland, West Virginia
| extent = Pennsylvania, Maryland, West Virginia, Ohio
| area =
| area =
| map =
| map =
| map_caption =
| map_caption =
}}
}}
The [[Pennsylvanian]] '''Allegheny Formation''' is a mapped bedrock unit in western and central [[Pennsylvania]], western [[Maryland]] and [[West Virginia]], USA. It is a major [[coal]]-bearing unit in the [[Ridge-and-Valley Appalachians]] of the eastern [[United States]].


The '''Allegheny Group''', often termed the '''Allegheny Formation''',<ref name=":6">{{Cite journal |last1=Opluštil |first1=Stanislav |last2=Cleal |first2=Christopher J. |last3=Wang |first3=Jun |last4=Wan |first4=Mingli |date=2022 |title=Carboniferous macrofloral biostratigraphy: an overview |url=https://www.lyellcollection.org/doi/10.1144/SP512-2020-97 |journal=Geological Society, London, Special Publications |language=en |volume=512 |issue=1 |pages=813–863 |doi=10.1144/SP512-2020-97 |bibcode=2022GSLSP.512..813O |issn=0305-8719}}</ref> is a [[Pennsylvanian (geology)|Pennsylvanian]]-age geological unit in the [[Appalachian Plateau]]. It is a major [[coal]]-bearing unit in the eastern [[United States]], extending through western and central [[Pennsylvania]], western [[Maryland]] and [[West Virginia]], and southeastern [[Ohio]]. [[Fossil]]s of [[fish]]es such as ''[[Bandringa]]'' are known from the [[Kittaning Formation]],<ref name="Zangerl1969">R. Zangerl. (1969). ''Bandringa rayi'': A New Ctenacanthoid Shark form the Pennsylvanian Essex Fauna of Illinois. ''Fieldiana Geology'' 12:157-169</ref> which is part of the Allegheny Group.
==Description==

The [[Pennsylvanian]] Allegheny Formation includes rocks from the base of the Brooksville Coal to the top of the Upper Freeport Coal, and was defined to include all econmically significant coals in the upper Pennsylvanian sequence.<ref>The Geology of Pennsylvania, C. H. Shultz, ed., DCNR Special Publication 1, 1999. (Ch. 10)</ref> The formation consists of [[Cyclothems|cyclothemic]] sequences of coal, shale, limestone, sandstone, and clay. It contains six major coal zones, which, in [[Stratigraphy|stratigraphic]] order, are:
==Stratigraphy==
In Pennsylvania, the Allegheny Group includes rocks from the base of the Brooksville Coal to the top of the Upper Freeport Coal, and was defined to include all economically significant coals in the upper Pennsylvanian sequence.<ref>The Geology of Pennsylvania, C. H. Shultz, ed., DCNR Special Publication 1, 1999. (Ch. 10) [https://web.archive.org/web/20031230153457/http://www.dcnr.state.pa.us/topogeo/pub/specialpub.aspx]</ref> The unit consists of [[Cyclothems|cyclothemic]] sequences of coal, shale, limestone, sandstone, and clay. It contains six major coal zones, which, in [[Stratigraphy|stratigraphic]] order, are:
* Upper Freeport Coal
* Upper Freeport Coal
* Lower Freeport Coal
* Lower Freeport Coal
Line 31: Line 47:
* Middle Kittanning Coal
* Middle Kittanning Coal
* Lower Kittanning Coal
* Lower Kittanning Coal
* Brooksville Coal
* Brookville Coal

===Members===
Glen Richey (PA), Laurel Run (PA), Mineral Springs (PA), Millstone Run (PA), Clearfield Creek (PA); Clarion (OH, MD, PA, WV), Kittanning (PA), Freeport (PA, MD, OH, WV); Putnam Hill (OH, PA); Vanport (PA, MD, OH, WV); Butler (MD, PA), Worthington (MD, PA); Washingtonville (OH, PA, WV), Columbiana (OH)<ref>[http://ngmdb.usgs.gov/Geolex/NewUnits/unit_53.html GEOLEX database, Geologic Unit: Allegheny], retrieved 28 December 2010</ref>

=== Age ===
Relative age dating of the Allegheny places it about mid-way through the [[Pennsylvanian (geology)|Pennsylvanian]], the second subperiod of the [[Carboniferous]] period. The plant and conchostracan fossils in particular can be equated with European strata of the "Asturian" ([[Westphalian D]]) regional stage. In North America, the roughly equivalent regional stage is known as the [[Desmoinesian]].<ref name=":6" /> The Asturian has an estimated age of 310.7 to 307.5 million years ago ([[Megaannum|Ma]]), equivalent to the latter half of the global [[Moscovian (Carboniferous)|Moscovian]] stage.<ref name=":2">{{Cite journal |last1=Knight |first1=John A. |last2=Cleal |first2=Christopher J. |last3=Álvarez-Vázquez |first3=Carmen |date=2023-06-14 |title=The challenge of relating the Kasimovian to west European chronostratigraphy: a critical review of the Cantabrian and Barruelian substages of the Stephanian Stage |journal=Geological Society, London, Special Publications |language=en |volume=535 |issue=1 |pages=31–71 |doi=10.1144/SP535-2022-189 |issn=0305-8719|doi-access=free |bibcode=2023GSLSP.535..189K }}</ref>

== Notable sites ==
The Allegheny Group has the richest fauna of [[tetrapod]] fossils in the entire [[Appalachian Basin]], and practically all of these fossils are concentrated at the Linton site of Ohio.<ref name=":22">{{Cite journal |last1=Hook |first1=Robert W. |last2=Baird |first2=Donald |date=1986-06-19 |title=The Diamond Coal Mine of Linton, Ohio, and its Pennsylvanian-age vertebrates |url=http://www.tandfonline.com/doi/abs/10.1080/02724634.1986.10011609 |journal=Journal of Vertebrate Paleontology |language=en |volume=6 |issue=2 |pages=174–190 |bibcode=1986JVPal...6..174H |doi=10.1080/02724634.1986.10011609 |issn=0272-4634}}</ref><ref name=":0">{{Cite journal |last1=Hook |first1=Robert W. |last2=Baird |first2=Donald |date=1988 |title=An Overview of the Upper Carboniferous Fossil Deposit at Linton, Ohio |url=https://kb.osu.edu/bitstream/handle/1811/23240/V088N1_055.pdf |journal=Ohio Journal of Science |volume=88 |issue=1 |pages=55–60}}</ref>

[[Linton, Ohio|Linton]] was once a tiny village at the mouth of Yellow Creek ([[Saline Township, Jefferson County, Ohio|Saline Township]]) in [[Jefferson County, Ohio|Jefferson County]]. The nearby Diamond Coal Mine was active from 1855 to 1892, and again from 1917 to 1921, when the [[drift mine]] was permanently closed. The drift mine collapsed soon afterwards and the settlement diminished into obscurity. The main Diamond coal seam is likely equivalent to the Upper Freeport coal. [[John Strong Newberry|John S. Newberry]], from [[Columbia University]] and Chief Geologist of the Geological Survey of Ohio, was the first paleontologist to investigate the Linton area.<ref>{{Cite journal |last=Babcock |first=L. E. |date=2024 |title=Some vertebrate types (Chondrichthyes, Actinopterygii, Sarcopterygii, and Tetrapoda) from two Paleozoic Lagerstätten of Ohio, U.S.A. |journal=Journal of Vertebrate Paleontology |volume=44 |pages=1–12 |doi=10.1080/02724634.2024.2308621|doi-access=free }}</ref> Starting in 1856, he collected thousands of fossils from the mine. Under the stewardship of [[Edward Drinker Cope|E.D. Cope]], some of Newberry's fossils were transferred to the [[American Museum of Natural History]] (AMNH), and others to the Orton Geological Museum at The Ohio State University. Other 19th-century geologists who collected fossils from the site include [[Frank Howe Bradley]] (1865, on behalf of [[Yale University|Yale]]), R. N. Fearon (1883, [[Harvard University|Harvard]]), and Thomas Stock (1888, [[British Museum]] and the [[National Museum of Natural History]]). Fossil collection extended into the 20th century, with expeditions led by Jesse Hyde (AMNH), [[Alfred Romer|A.S. Romer]] ([[Field Museum of Natural History|Field Museum]]), Donald Baird (Harvard, [[Princeton University|Princeton]]), Richard Lund and David Hamilla ([[Carnegie Museum of Natural History|Carnegie]]). Over 7000 Linton fossils are now in the collections of at least 14 museums across three countries.<ref name=":22" /><ref name=":0" />

A second fossiliferous site is Five Points, a similar coal deposit in [[Beaver Township, Mahoning County, Ohio|Beaver Township]] of [[Mahoning County, Ohio|Mahoning County]]. The Five Points cannel coal mine was first publicized as a fossil site by Robert Hook and Donald Baird in 1994, who collected tetrapod fossils comparable to those found at Linton. However, the mine at Five Points was [[Mine reclamation|reclaimed]] around the same time, so the fossiliferous [[Spoil tip|spoil piles]] are no longer accessible for further collection.<ref name=":1">{{Cite journal |last=Sequeira |first=Sandra E. K. |date=1998 |title=The cranial morphology and taxonomy of the saurerpetontid Isodectes obtusus comb. nov. (Amphibia: Temnospondyli) from the Lower Permian of Texas |journal=Zoological Journal of the Linnean Society |language=en |volume=122 |issue=1–2 |pages=237–259 |doi=10.1111/j.1096-3642.1998.tb02531.x|doi-access=free }}</ref><ref name=":3" />

== Paleobiota ==
Tetrapod records from Hook & Baird (1986/1988)<ref name=":22" /><ref name=":0" /> unless stated otherwise:

=== Amniotes ===
{| class="wikitable" align="center"
! colspan="4" align="center" |'''[[Amniote|Amniotes]] of East Kirkton'''
|-
!Species
!Locality
!Notes
!Images
|-
|''[[Anthracodromeus|Anthracodromeus longipes]]''
|Linton
|A "[[Protorothyrididae|protorothyridid]]" [[Eureptilia|eureptile]]
|[[File:Anthracodromeus longipes.jpg|center|150x150px]]
|-
|''[[Archaeothyris]]'' sp.
|Linton
|A possible [[Ophiacodontidae|ophiacodontid]] [[synapsid]] based on rare fragments
|[[File:Archaeothyris BW.jpg|center|150x150px]]
|-
|''[[Carbonodraco|Carbonodraco lundi]]<ref>{{Cite journal |last1=Mann |first1=Arjan |last2=McDaniel |first2=Emily J. |last3=McColville |first3=Emily R. |last4=Maddin |first4=Hillary C. |year=2019 |title=''Carbonodraco lundi'' gen et sp. nov., the oldest parareptile, from Linton, Ohio, and new insights into the early radiation of reptiles |url= |journal=Royal Society Open Science |volume=6 |issue=11 |pages=191191 |bibcode=2019RSOS....691191M |doi=10.1098/rsos.191191 |pmc=6894558 |pmid=31827854}}</ref>''
|Linton
|An [[Acleistorhinidae|acleistorhinid]] [[Parareptilia|parareptile]] based on fossils previously referred to ''[[Cephalerpeton]]''
|[[File:Carbonodraco holotype diagram.jpg|center|150x150px]]
|-
|''[[Melanedaphodon|Malanedaphodon hovaneci]]''<ref name="Mann2023">{{cite journal |last1=Mann |first1=A. |last2=Henrici |first2=A. C. |last3=Sues |first3=H.-D. |last4=Pierce |first4=S. E. |title=A new Carboniferous edaphosaurid and the origin of herbivory in mammal forerunners |journal=Scientific Reports |date=2023 |volume=13 |issue=1 |pages=4459 |doi=10.1038/s41598-023-30626-8|pmid=37019927 |pmc=10076360 }}</ref>
|Linton
|A [[Edaphosauridae|edaphosaurid]] [[Synapsida|synapsid]]
|[[File:Melanedaphodon.jpg|center|150x150px]]
|}

=== Temnospondyls ===
{| class="wikitable" align="center"
! colspan="4" align="center" |'''[[Temnospondyli|Temnospondyls]] of East Kirkton'''
|-
!Species
!Locality
!Notes
!Images
|-
|''[[Adamanterpeton|Adamanterpeton ohioensis]]<ref name="MS98">{{cite journal |last=Milner |first=A.R. |author2=Sequeira, S.E.K. |year=1998 |title=A cochleosaurid temnospondyl amphibian from the Middle Pennsylvanian of Linton, Ohio, U.S.A. |journal=Zoological Journal of the Linnean Society |volume=122 |issue=1 |pages=261–290 |doi=10.1111/j.1096-3642.1998.tb02532.x |doi-access=}}</ref>''
|Linton
|A rare [[Cochleosauridae|cochleosaurid]] [[Edopoidea|edopoid]], previously referred to "''[[Gaudrya]]'' cf. ''latistoma''"
|
|-
|''[[Erpetosaurus|Erpetosaurus radiatus]]''
|Linton
|A common [[Eobrachyopidae|eobrachyopid]] [[Dvinosauria|dvinosaur]]
|
|-
|''[[Isodectes|Isodectes obtusus]]''<ref name=":1" />
|Linton, Five Points<ref name=":1" />
|An abundant eobrachyopid dvinosaur, previously known as ''Saurerpeton obtusum''
|[[File:Isodectes obtusus.jpg|center|150x150px]]
|-
|''[[Macrerpeton|Macrerpeton huxleyi]]''
|Linton
|A rare temnospondyl, possibly an edopoid or dissorophoid<ref>{{Cite journal |last1=Schoch |first1=Rainer R. |last2=Milner |first2=Andrew R. |date=2021 |title=Morphology and relationships of the temnospondyl Macrerpeton huxleyi from the Pennsylvanian of Linton, Ohio (USA) |url= |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |language=en |volume=299 |issue=1 |pages=77–98 |doi=10.1127/njgpa/2021/0956 |issn=0077-7749}}</ref>
|
|-
|''[[Palodromeus|Palodromeus bairdi]]''<ref name=":3">{{Cite journal |last1=Schoch |first1=Rainer R. |last2=Henrici |first2=Amy C. |last3=Hook |first3=Robert W. |date=2021 |title=A new dissorophoid temnospondyl from the Allegheny Group (late Carboniferous) of Five Points, Mahoning County, Ohio (USA) |url=https://www.cambridge.org/core/product/identifier/S0022336020001018/type/journal_article |journal=Journal of Paleontology |language=en |volume=95 |issue=3 |pages=638–651 |doi=10.1017/jpa.2020.101 |bibcode=2021JPal...95..638S |issn=0022-3360}}</ref>
|Five Points
|The earliest-braching [[Olsoniformes|olsoniform]] dissorophoid, known from a single skull
|
|-
|''[[Platyrhinops|Platyrhinops lyelli]]''
|Linton
|A common [[Amphibamiformes|amphibamiform]] dissorophoid, previously considered a species of ''[[Amphibamus]]''
|[[File:PSM V72 D566 Pelion lyelli quadraped from carboniferous ohio.png|center|237x237px]]
|-
|''[[Stegops|Stegops newberryi]]''
|Linton
|A rare spiny dissorophoid
|[[File:Stegops.jpg|center|150x150px]]
|}

=== Lepospondyls ===
{| class="wikitable" align="center"
! colspan="4" align="center" |'''[[Lepospondyli|Lepospondyls]] of East Kirkton'''
|-
!Species
!Locality
!Notes
!Images
|-
|''[[Brachydectes|Brachydectes newberryi]]''
|Linton
|A [[Lysorophia|lysorophian]], likely encompassing fossils previously referred to ''Pleuroptyx clavatus''<ref>{{Cite journal |last=Mann |first=Arjan |date=2018-10-06 |title=Cranial ornamentation of a large Brachydectes newberryi (Recumbirostra: Lysorophia) from Linton, Ohio. |url=https://journals.library.ualberta.ca/vamp/index.php/VAMP/article/view/29341 |journal=Vertebrate Anatomy Morphology Palaeontology |volume=6 |pages=91–96 |doi=10.18435/vamp29341 |issn=2292-1389|doi-access=free }}</ref> (which are rare) and ''Cocytinus gyrinoides<ref name="WCF91">{{cite journal |last=Wellstead |first=C. F. |year=1991 |title=Taxonomic revision of the Lysorophia, Permo-Carboniferous lepospondyl amphibians |url=http://digitallibrary.amnh.org/dspace/bitstream/2246/904/1/B209a01.pdf |journal=Bulletin of the American Museum of Natural History |volume=209 |pages=1–90}}</ref>'' (which are common)
|[[File:Brachydectes.jpg|center|150x150px]]
|-
|''[[Ctenerpeton|Ctenerpeton remex]]''
|Linton
|A rare [[Urocordylidae|urocordylid]] [[Nectridea|nectridean]]
|
|-
|''[[Diceratosaurus|Diceratosaurus brevirostris]]''
|Linton
|An abundant [[Diplocaulidae|diplocaulid]] nectridean
|[[File:Diceratosaurus.jpg|center|150x150px]]
|-
|''[[Molgophis|Molgophis macrurus]]''
|Linton
|A rare lysorophian
|
|-
|''[[Oestocephalus|Oestocephalus amphiuminus]]''
|Linton, Five Points<ref>{{Cite journal |last1=Pardo |first1=Jason D. |last2=Holmes |first2=Robert |last3=Anderson |first3=Jason S. |date=2018 |title=An enigmatic braincase from Five Points, Ohio (Westphalian D) further supports a stem tetrapod position for aïstopods |url=https://www.researchgate.net/publication/330323000 |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |language=en |volume=109 |issue=1–2 |pages=255–264 |doi=10.1017/S1755691018000567 |bibcode=2018EESTR.109..255P |issn=1755-6910}}</ref>
|An abundant [[Aistopoda|aistopod]], previously considered a species of ''[[Ophiderpeton]]''.
|[[File:Oestocephalus.jpg|center|195x195px]]
|-
|''[[Odonterpeton|Odonterpeton triangulare]]''
|Linton
|A rare small-limbed [[Microsauria|microsaur]] known from a single partial skeleton
|[[File:Odonterpeton.jpg|center|150x150px]]
|-
|''[[Phlegethontia|Phlegethontia linearis]]''
|Linton
|A common aistopod
|[[File:Phlegethontia.jpg|center|150x150px]]
|-
|''[[Ptyonius|Ptyonius marshii]]''
|Linton
|An abundant urocordylid nectridean
|
|-
|''[[Sauropleura|Sauropleura pectinata]]''
|Linton
|An abundant urocordylid nectridean
|[[File:Sauropleura cropped.jpg|center|150x150px]]
|-
|''[[Tuditanus|Tuditanus punctulatus]]''
|Linton
|A rare [[Tuditanidae|tuditanid]] microsaur
|[[File:The Osteology of the Reptiles-234 rty wer rt.png|center|150x150px]]
|}

=== Other amphibians ===
Various indeterminate [[Embolomeri|embolomere]] fossils are known from the fossil sites of the Allegheny Group, including an articulated tail from Five Points,<ref>{{Cite journal |last=Clack |first=Jennifer A. |date=2011 |title=A Carboniferous embolomere tail with supraneural radials |url=http://www.tandfonline.com/doi/abs/10.1080/02724634.2011.595467 |journal=Journal of Vertebrate Paleontology |language=en |volume=31 |issue=5 |pages=1150–1153 |doi=10.1080/02724634.2011.595467 |bibcode=2011JVPal..31.1150C |issn=0272-4634}}</ref> an [[Eogyrinidae|eogyrinid]]-like skull roof from Linton,<ref>{{Cite journal |last1=Witzmann |first1=Florian |last2=Werneburg |first2=Ralf |last3=Milner |first3=Andrew R. |date=2017 |title=A partial skull roof of an embolomere from Linton, Ohio (Middle Pennsylvanian) and its phylogenetic affinities |url=https://www.researchgate.net/publication/318293578 |journal=PalZ |language=en |volume=91 |issue=3 |pages=399–408 |doi=10.1007/s12542-017-0374-4 |bibcode=2017PalZ...91..399W |issn=0031-0220}}</ref> and [[Archeriidae|archeriid]]-like cranial and postcranial fragments from both Linton and Five Points.<ref>{{Cite journal |last1=Holmes |first1=Robert |last2=Baird |first2=Donald |date=2011-05-04 |title=The Smaller Embolomerous Amphibians (Anthracosauria) from the Middle Pennsylvanian (Desmoinesian) Localities at Linton and Five Points Coal Mines, Ohio |url=http://www.bioone.org/doi/abs/10.3099/0006-9698-523.1.1 |journal=Breviora |language=en |volume=523 |pages=1–13 |doi=10.3099/0006-9698-523.1.1 |issn=0006-9698}}</ref>
{| class="wikitable" align="center"
! colspan="4" align="center" |'''Tetrapods of East Kirkton'''
|-
!Species
!Locality
!Notes
!Images
|-
|''[[Baphetes|Baphetes lineolatus]]''
|Linton
|A rare [[Baphetidae|baphetid]]
|
|-
|''[[Colosteus|Colosteus scutellatus]]''
|Linton
|An abundant [[Colosteidae|colosteid]]
|[[File:Colosteus NT.jpg|center|150x150px]]
|-
|''[[Eusauropleura|Eusauropleura digitata]]''
|Linton
|A rare [[Gephyrostegidae|gephyrostegid]]
|
|-
|''[[Leptophractus|Leptophractus obsoletus]]''
|Linton
|A rare embolomere, likely encompassing fossils previously given the name "''Anthracosaurus lancifer''"
|
|-
|''[[Megalocephalus|Megalocephalus lineolatus]]''
|Linton
|A rare baphetid, also known as ''Megalocephalus enchodus''
|[[File:MegalocephalusDB.jpg|center|150x150px]]
|}

=== Invertebrates ===
Marine fossils in the Allegheny Group are concentrated into only a few patchy bands of limestone and shale. Four marine members (Putnam Hill, Vanport, Columbiana, and Washingtonville) have produced a rich fauna of [[cephalopod]] fossils, the best representation of the Desmoinesian stage in the Appalachian region.<ref>{{Cite journal |last1=Sturgeon |first1=M. T. |last2=Windle |first2=D. L. |last3=Mapes |first3=R. H. |last4=Hoare |first4=R. D. |date=1982 |title=New and Revised Taxa of Pennsylvanian Cephalopods in Ohio and West Virginia |url= |journal=Journal of Paleontology |volume=56 |issue=6 |pages=1453–1479 |issn=0022-3360 |jstor=1304678}}</ref><ref>{{Cite journal |last1=Darwin R. II |first1=Boardman |last2=Work |first2=David M. |last3=Mapes |first3=Royal H. |last4=Barrick |first4=James E. |date=1994-03-15 |title=Biostratigraphy of Middle and Late Pennsylvanian (Desmoinesian-Virgilian) ammonoids |url=https://journals.ku.edu/kgsbulletin/article/view/20477 |journal=Bulletin (Kansas Geological Survey) |language=en |volume=232 |issue= |pages=1–122 |issn=0097-4471}}</ref><ref>{{Cite journal |last=Nikolaeva |first=S. V. |date=2022 |title=Carboniferous ammonoid genozones |url=https://www.lyellcollection.org/doi/10.1144/SP512-2020-229 |journal=Geological Society, London, Special Publications |language=en |volume=512 |issue=1 |pages=633–693 |doi=10.1144/SP512-2020-229 |bibcode=2022GSLSP.512..633N |issn=0305-8719}}</ref>

In terrestrial sediments, the Allegheny Group preserves characteristic Desmoinesian [[index fossils]] of [[Conchostracan|conchostracans]] (bivalved [[Crustacean|crustaceans]]). Conchostracan species in the unit belong to an [[Assemblage Zone|assemblage zone]] distinguished by ''[[Anomalonema|Anomalonema reumauxi]]'', ''[[Pseudestheria|Pseudestheria simoni]]'', and potentially ''[[Palaeolimnadiopsis|Palaeolimnadiopsis freysteini]]''.<ref>{{Cite journal |last1=Schneider |first1=Joerg W. |last2=Scholze |first2=Frank |last3=Ross |first3=Andrew J. |last4=Blake |first4=Bascombe M. |last5=Lucas |first5=Spencer G. |date=2022 |title=Improved blattoid insect and conchostracan zonation for the Late Carboniferous, Pennsylvanian, of Euramerica |url=https://www.lyellcollection.org/doi/10.1144/SP512-2021-93 |journal=Geological Society, London, Special Publications |language=en |volume=512 |issue=1 |pages=865–891 |doi=10.1144/SP512-2021-93 |bibcode=2022GSLSP.512..865S |issn=0305-8719}}</ref><ref>{{Cite journal |last1=Schneider |first1=Joerg W. |last2=Scholze |first2=Frank |date=2018 |title=Late Pennsylvanian–Early Triassic conchostracan biostratigraphy: a preliminary approach |url=https://www.lyellcollection.org/doi/10.1144/SP450.6 |journal=Geological Society, London, Special Publications |language=en |volume=450 |issue=1 |pages=365–386 |doi=10.1144/SP450.6 |bibcode=2018GSLSP.450..365S |issn=0305-8719}}</ref> Invertebrate fossils from Linton include '[[Microconchida|spirorbid]]' tubes, [[Ostracod|ostracods]], [[Syncarida|syncarid]] crustaceans, and [[Millipede|millipedes]] (''[[Xyloiulus|Xyloiulus bairdi]]'', ''[[Plagiascetus|Plagiascetus lateralis]]'', and other undescribed species).<ref>{{Cite journal |last=Hoffman |first=Richard L. |date=1963 |title=New Genera and Species of Upper Paleozoic Diplopoda |url= |journal=Journal of Paleontology |volume=37 |issue=1 |pages=167–174 |issn=0022-3360 |jstor=1301419}}</ref><ref name=":0" /> Some groups (insects, [[Arachnid|arachnids]], and freshwater [[Bivalvia|bivalves]]) are curiously absent from Linton.<ref name=":0" />


==Age==
=== Plants ===
The lowermost portion of the Allegheny Group belongs to the ''Laveineopteris rarinervis'' plant [[macrofossil]] zone. This [[biozone]] is distinguished by ''[[Laveineopteris|Laveineopteris rarinervis]]'' and ''[[Neuropteris|Neuropteris ovata]]'', two species of [[Medullosales|medullosalean]] "[[Pteridospermatophyta|seed ferns]]" which first appear in the uppermost Kanawha Formation. In the Kittanning coals and higher stratigraphic sections, the plant fossils transition to the ''Neuropteris flexuosa'' zone, characterized by ''Neuropteris flexuosa'' and the [[Marattiales|marrattialean]] [[fern]] ''[[Cyathocarpus]]''.<ref name=":6" /> The plant macrofossil record is mirrored by [[Microfossil|microfossils]]: "tree fern" spores are the most diverse components of the [[Palynology|palynoflora]], and ''[[Lycospora]]'' (a [[lycopsid]] [[miospore]]) is also abundant.<ref name=":0" /><ref>{{Cite journal |last=Eble |first=Cortland F. |date=2022 |title=Appalachian coal bed palynofloras: changes in composition through time and comparison with other areas |url=https://www.lyellcollection.org/doi/10.1144/SP512-2021-131 |journal=Geological Society, London, Special Publications |language=en |volume=512 |issue=1 |pages=791–812 |doi=10.1144/SP512-2021-131 |bibcode=2022GSLSP.512..791E |issn=0305-8719}}</ref>
Relative age dating of the Allegheny places it in the middle [[Pennsylvanian]] period.


==References==
<references/>


==See also==
== References ==
{{reflist}}
*[[Geology of Pennsylvania]]
*[[Geology of West Virginia]]


{{Chronostratigraphy of Ohio}}
[[Category:Geologic formations]]
[[Category:Geology of Pennsylvania]]
{{Chronostratigraphy of Pennsylvania}}
[[Category:Geology of West Virginia]]
{{Stratigraphic column of West Virginia}}
[[Category:Sandstone]]
[[Category:Siltstone]]
[[Category:Coal]]


[[Category:Pennsylvanian Series]]
{{geologic-formation-stub}}
[[Category:Carboniferous geology of Pennsylvania]]
[[Category:Carboniferous West Virginia]]
[[Category:Sandstone formations of the United States]]
[[Category:Shale formations of the United States]]
[[Category:Coal formations]]
[[Category:Coal in the United States]]
[[Category:Geologic formations of Ohio]]
[[Category:Geologic formations of Pennsylvania]]
[[Category:Geologic formations of West Virginia]]
[[Category:Carboniferous southern paleotropical deposits]]

Latest revision as of 20:15, 26 August 2024

Allegheny Group
Stratigraphic range: (Moscovian)
~311–306.8 Ma
Bituminous coal (Clarion Coal) from the Allegheny Group, Ohio
TypeSedimentary
Sub-unitsFreeport Formation
Upper Freeport Coal
Upper Freeport Limestone Member
Butler Sandstone Member
Lower Freeport Coal
Freeport Sandstone Member

Kittanning Formation

Upper Kittanning Coal
Johnstone Limestone Member
Middle Kittanning Coal
Lower Kittanning Coal
Kittanning Sandstone Member
Vanport Limestone Member
UnderliesConemaugh Group
OverliesKanawha Formation and Pottsville Formation
Lithology
PrimarySandstone, coal
Location
RegionAppalachian Mountains
CountryUnited States
ExtentPennsylvania, Maryland, West Virginia, Ohio
Type section
Named byH. D. Rogers, 1840[1]

The Allegheny Group, often termed the Allegheny Formation,[2] is a Pennsylvanian-age geological unit in the Appalachian Plateau. It is a major coal-bearing unit in the eastern United States, extending through western and central Pennsylvania, western Maryland and West Virginia, and southeastern Ohio. Fossils of fishes such as Bandringa are known from the Kittaning Formation,[3] which is part of the Allegheny Group.

Stratigraphy

[edit]

In Pennsylvania, the Allegheny Group includes rocks from the base of the Brooksville Coal to the top of the Upper Freeport Coal, and was defined to include all economically significant coals in the upper Pennsylvanian sequence.[4] The unit consists of cyclothemic sequences of coal, shale, limestone, sandstone, and clay. It contains six major coal zones, which, in stratigraphic order, are:

  • Upper Freeport Coal
  • Lower Freeport Coal
  • Upper Kittanning Coal
  • Middle Kittanning Coal
  • Lower Kittanning Coal
  • Brookville Coal

Members

[edit]

Glen Richey (PA), Laurel Run (PA), Mineral Springs (PA), Millstone Run (PA), Clearfield Creek (PA); Clarion (OH, MD, PA, WV), Kittanning (PA), Freeport (PA, MD, OH, WV); Putnam Hill (OH, PA); Vanport (PA, MD, OH, WV); Butler (MD, PA), Worthington (MD, PA); Washingtonville (OH, PA, WV), Columbiana (OH)[5]

Age

[edit]

Relative age dating of the Allegheny places it about mid-way through the Pennsylvanian, the second subperiod of the Carboniferous period. The plant and conchostracan fossils in particular can be equated with European strata of the "Asturian" (Westphalian D) regional stage. In North America, the roughly equivalent regional stage is known as the Desmoinesian.[2] The Asturian has an estimated age of 310.7 to 307.5 million years ago (Ma), equivalent to the latter half of the global Moscovian stage.[6]

Notable sites

[edit]

The Allegheny Group has the richest fauna of tetrapod fossils in the entire Appalachian Basin, and practically all of these fossils are concentrated at the Linton site of Ohio.[7][8]

Linton was once a tiny village at the mouth of Yellow Creek (Saline Township) in Jefferson County. The nearby Diamond Coal Mine was active from 1855 to 1892, and again from 1917 to 1921, when the drift mine was permanently closed. The drift mine collapsed soon afterwards and the settlement diminished into obscurity. The main Diamond coal seam is likely equivalent to the Upper Freeport coal. John S. Newberry, from Columbia University and Chief Geologist of the Geological Survey of Ohio, was the first paleontologist to investigate the Linton area.[9] Starting in 1856, he collected thousands of fossils from the mine. Under the stewardship of E.D. Cope, some of Newberry's fossils were transferred to the American Museum of Natural History (AMNH), and others to the Orton Geological Museum at The Ohio State University. Other 19th-century geologists who collected fossils from the site include Frank Howe Bradley (1865, on behalf of Yale), R. N. Fearon (1883, Harvard), and Thomas Stock (1888, British Museum and the National Museum of Natural History). Fossil collection extended into the 20th century, with expeditions led by Jesse Hyde (AMNH), A.S. Romer (Field Museum), Donald Baird (Harvard, Princeton), Richard Lund and David Hamilla (Carnegie). Over 7000 Linton fossils are now in the collections of at least 14 museums across three countries.[7][8]

A second fossiliferous site is Five Points, a similar coal deposit in Beaver Township of Mahoning County. The Five Points cannel coal mine was first publicized as a fossil site by Robert Hook and Donald Baird in 1994, who collected tetrapod fossils comparable to those found at Linton. However, the mine at Five Points was reclaimed around the same time, so the fossiliferous spoil piles are no longer accessible for further collection.[10][11]

Paleobiota

[edit]

Tetrapod records from Hook & Baird (1986/1988)[7][8] unless stated otherwise:

Amniotes

[edit]
Amniotes of East Kirkton
Species Locality Notes Images
Anthracodromeus longipes Linton A "protorothyridid" eureptile
Archaeothyris sp. Linton A possible ophiacodontid synapsid based on rare fragments
Carbonodraco lundi[12] Linton An acleistorhinid parareptile based on fossils previously referred to Cephalerpeton
Malanedaphodon hovaneci[13] Linton A edaphosaurid synapsid

Temnospondyls

[edit]
Temnospondyls of East Kirkton
Species Locality Notes Images
Adamanterpeton ohioensis[14] Linton A rare cochleosaurid edopoid, previously referred to "Gaudrya cf. latistoma"
Erpetosaurus radiatus Linton A common eobrachyopid dvinosaur
Isodectes obtusus[10] Linton, Five Points[10] An abundant eobrachyopid dvinosaur, previously known as Saurerpeton obtusum
Macrerpeton huxleyi Linton A rare temnospondyl, possibly an edopoid or dissorophoid[15]
Palodromeus bairdi[11] Five Points The earliest-braching olsoniform dissorophoid, known from a single skull
Platyrhinops lyelli Linton A common amphibamiform dissorophoid, previously considered a species of Amphibamus
Stegops newberryi Linton A rare spiny dissorophoid

Lepospondyls

[edit]
Lepospondyls of East Kirkton
Species Locality Notes Images
Brachydectes newberryi Linton A lysorophian, likely encompassing fossils previously referred to Pleuroptyx clavatus[16] (which are rare) and Cocytinus gyrinoides[17] (which are common)
Ctenerpeton remex Linton A rare urocordylid nectridean
Diceratosaurus brevirostris Linton An abundant diplocaulid nectridean
Molgophis macrurus Linton A rare lysorophian
Oestocephalus amphiuminus Linton, Five Points[18] An abundant aistopod, previously considered a species of Ophiderpeton.
Odonterpeton triangulare Linton A rare small-limbed microsaur known from a single partial skeleton
Phlegethontia linearis Linton A common aistopod
Ptyonius marshii Linton An abundant urocordylid nectridean
Sauropleura pectinata Linton An abundant urocordylid nectridean
Tuditanus punctulatus Linton A rare tuditanid microsaur

Other amphibians

[edit]

Various indeterminate embolomere fossils are known from the fossil sites of the Allegheny Group, including an articulated tail from Five Points,[19] an eogyrinid-like skull roof from Linton,[20] and archeriid-like cranial and postcranial fragments from both Linton and Five Points.[21]

Tetrapods of East Kirkton
Species Locality Notes Images
Baphetes lineolatus Linton A rare baphetid
Colosteus scutellatus Linton An abundant colosteid
Eusauropleura digitata Linton A rare gephyrostegid
Leptophractus obsoletus Linton A rare embolomere, likely encompassing fossils previously given the name "Anthracosaurus lancifer"
Megalocephalus lineolatus Linton A rare baphetid, also known as Megalocephalus enchodus

Invertebrates

[edit]

Marine fossils in the Allegheny Group are concentrated into only a few patchy bands of limestone and shale. Four marine members (Putnam Hill, Vanport, Columbiana, and Washingtonville) have produced a rich fauna of cephalopod fossils, the best representation of the Desmoinesian stage in the Appalachian region.[22][23][24]

In terrestrial sediments, the Allegheny Group preserves characteristic Desmoinesian index fossils of conchostracans (bivalved crustaceans). Conchostracan species in the unit belong to an assemblage zone distinguished by Anomalonema reumauxi, Pseudestheria simoni, and potentially Palaeolimnadiopsis freysteini.[25][26] Invertebrate fossils from Linton include 'spirorbid' tubes, ostracods, syncarid crustaceans, and millipedes (Xyloiulus bairdi, Plagiascetus lateralis, and other undescribed species).[27][8] Some groups (insects, arachnids, and freshwater bivalves) are curiously absent from Linton.[8]

Plants

[edit]

The lowermost portion of the Allegheny Group belongs to the Laveineopteris rarinervis plant macrofossil zone. This biozone is distinguished by Laveineopteris rarinervis and Neuropteris ovata, two species of medullosalean "seed ferns" which first appear in the uppermost Kanawha Formation. In the Kittanning coals and higher stratigraphic sections, the plant fossils transition to the Neuropteris flexuosa zone, characterized by Neuropteris flexuosa and the marrattialean fern Cyathocarpus.[2] The plant macrofossil record is mirrored by microfossils: "tree fern" spores are the most diverse components of the palynoflora, and Lycospora (a lycopsid miospore) is also abundant.[8][28]


References

[edit]
  1. ^ Rogers, H.D., (1840), Fourth annual report of the Geological Survey of the State of Pennsylvania: Pennsylvania Geological Survey Annual Report, no. 4, 215 p. ([1])
  2. ^ a b c Opluštil, Stanislav; Cleal, Christopher J.; Wang, Jun; Wan, Mingli (2022). "Carboniferous macrofloral biostratigraphy: an overview". Geological Society, London, Special Publications. 512 (1): 813–863. Bibcode:2022GSLSP.512..813O. doi:10.1144/SP512-2020-97. ISSN 0305-8719.
  3. ^ R. Zangerl. (1969). Bandringa rayi: A New Ctenacanthoid Shark form the Pennsylvanian Essex Fauna of Illinois. Fieldiana Geology 12:157-169
  4. ^ The Geology of Pennsylvania, C. H. Shultz, ed., DCNR Special Publication 1, 1999. (Ch. 10) [2]
  5. ^ GEOLEX database, Geologic Unit: Allegheny, retrieved 28 December 2010
  6. ^ Knight, John A.; Cleal, Christopher J.; Álvarez-Vázquez, Carmen (2023-06-14). "The challenge of relating the Kasimovian to west European chronostratigraphy: a critical review of the Cantabrian and Barruelian substages of the Stephanian Stage". Geological Society, London, Special Publications. 535 (1): 31–71. Bibcode:2023GSLSP.535..189K. doi:10.1144/SP535-2022-189. ISSN 0305-8719.
  7. ^ a b c Hook, Robert W.; Baird, Donald (1986-06-19). "The Diamond Coal Mine of Linton, Ohio, and its Pennsylvanian-age vertebrates". Journal of Vertebrate Paleontology. 6 (2): 174–190. Bibcode:1986JVPal...6..174H. doi:10.1080/02724634.1986.10011609. ISSN 0272-4634.
  8. ^ a b c d e f Hook, Robert W.; Baird, Donald (1988). "An Overview of the Upper Carboniferous Fossil Deposit at Linton, Ohio" (PDF). Ohio Journal of Science. 88 (1): 55–60.
  9. ^ Babcock, L. E. (2024). "Some vertebrate types (Chondrichthyes, Actinopterygii, Sarcopterygii, and Tetrapoda) from two Paleozoic Lagerstätten of Ohio, U.S.A." Journal of Vertebrate Paleontology. 44: 1–12. doi:10.1080/02724634.2024.2308621.
  10. ^ a b c Sequeira, Sandra E. K. (1998). "The cranial morphology and taxonomy of the saurerpetontid Isodectes obtusus comb. nov. (Amphibia: Temnospondyli) from the Lower Permian of Texas". Zoological Journal of the Linnean Society. 122 (1–2): 237–259. doi:10.1111/j.1096-3642.1998.tb02531.x.
  11. ^ a b Schoch, Rainer R.; Henrici, Amy C.; Hook, Robert W. (2021). "A new dissorophoid temnospondyl from the Allegheny Group (late Carboniferous) of Five Points, Mahoning County, Ohio (USA)". Journal of Paleontology. 95 (3): 638–651. Bibcode:2021JPal...95..638S. doi:10.1017/jpa.2020.101. ISSN 0022-3360.
  12. ^ Mann, Arjan; McDaniel, Emily J.; McColville, Emily R.; Maddin, Hillary C. (2019). "Carbonodraco lundi gen et sp. nov., the oldest parareptile, from Linton, Ohio, and new insights into the early radiation of reptiles". Royal Society Open Science. 6 (11): 191191. Bibcode:2019RSOS....691191M. doi:10.1098/rsos.191191. PMC 6894558. PMID 31827854.
  13. ^ Mann, A.; Henrici, A. C.; Sues, H.-D.; Pierce, S. E. (2023). "A new Carboniferous edaphosaurid and the origin of herbivory in mammal forerunners". Scientific Reports. 13 (1): 4459. doi:10.1038/s41598-023-30626-8. PMC 10076360. PMID 37019927.
  14. ^ Milner, A.R.; Sequeira, S.E.K. (1998). "A cochleosaurid temnospondyl amphibian from the Middle Pennsylvanian of Linton, Ohio, U.S.A.". Zoological Journal of the Linnean Society. 122 (1): 261–290. doi:10.1111/j.1096-3642.1998.tb02532.x.
  15. ^ Schoch, Rainer R.; Milner, Andrew R. (2021). "Morphology and relationships of the temnospondyl Macrerpeton huxleyi from the Pennsylvanian of Linton, Ohio (USA)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 299 (1): 77–98. doi:10.1127/njgpa/2021/0956. ISSN 0077-7749.
  16. ^ Mann, Arjan (2018-10-06). "Cranial ornamentation of a large Brachydectes newberryi (Recumbirostra: Lysorophia) from Linton, Ohio". Vertebrate Anatomy Morphology Palaeontology. 6: 91–96. doi:10.18435/vamp29341. ISSN 2292-1389.
  17. ^ Wellstead, C. F. (1991). "Taxonomic revision of the Lysorophia, Permo-Carboniferous lepospondyl amphibians" (PDF). Bulletin of the American Museum of Natural History. 209: 1–90.
  18. ^ Pardo, Jason D.; Holmes, Robert; Anderson, Jason S. (2018). "An enigmatic braincase from Five Points, Ohio (Westphalian D) further supports a stem tetrapod position for aïstopods". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 109 (1–2): 255–264. Bibcode:2018EESTR.109..255P. doi:10.1017/S1755691018000567. ISSN 1755-6910.
  19. ^ Clack, Jennifer A. (2011). "A Carboniferous embolomere tail with supraneural radials". Journal of Vertebrate Paleontology. 31 (5): 1150–1153. Bibcode:2011JVPal..31.1150C. doi:10.1080/02724634.2011.595467. ISSN 0272-4634.
  20. ^ Witzmann, Florian; Werneburg, Ralf; Milner, Andrew R. (2017). "A partial skull roof of an embolomere from Linton, Ohio (Middle Pennsylvanian) and its phylogenetic affinities". PalZ. 91 (3): 399–408. Bibcode:2017PalZ...91..399W. doi:10.1007/s12542-017-0374-4. ISSN 0031-0220.
  21. ^ Holmes, Robert; Baird, Donald (2011-05-04). "The Smaller Embolomerous Amphibians (Anthracosauria) from the Middle Pennsylvanian (Desmoinesian) Localities at Linton and Five Points Coal Mines, Ohio". Breviora. 523: 1–13. doi:10.3099/0006-9698-523.1.1. ISSN 0006-9698.
  22. ^ Sturgeon, M. T.; Windle, D. L.; Mapes, R. H.; Hoare, R. D. (1982). "New and Revised Taxa of Pennsylvanian Cephalopods in Ohio and West Virginia". Journal of Paleontology. 56 (6): 1453–1479. ISSN 0022-3360. JSTOR 1304678.
  23. ^ Darwin R. II, Boardman; Work, David M.; Mapes, Royal H.; Barrick, James E. (1994-03-15). "Biostratigraphy of Middle and Late Pennsylvanian (Desmoinesian-Virgilian) ammonoids". Bulletin (Kansas Geological Survey). 232: 1–122. ISSN 0097-4471.
  24. ^ Nikolaeva, S. V. (2022). "Carboniferous ammonoid genozones". Geological Society, London, Special Publications. 512 (1): 633–693. Bibcode:2022GSLSP.512..633N. doi:10.1144/SP512-2020-229. ISSN 0305-8719.
  25. ^ Schneider, Joerg W.; Scholze, Frank; Ross, Andrew J.; Blake, Bascombe M.; Lucas, Spencer G. (2022). "Improved blattoid insect and conchostracan zonation for the Late Carboniferous, Pennsylvanian, of Euramerica". Geological Society, London, Special Publications. 512 (1): 865–891. Bibcode:2022GSLSP.512..865S. doi:10.1144/SP512-2021-93. ISSN 0305-8719.
  26. ^ Schneider, Joerg W.; Scholze, Frank (2018). "Late Pennsylvanian–Early Triassic conchostracan biostratigraphy: a preliminary approach". Geological Society, London, Special Publications. 450 (1): 365–386. Bibcode:2018GSLSP.450..365S. doi:10.1144/SP450.6. ISSN 0305-8719.
  27. ^ Hoffman, Richard L. (1963). "New Genera and Species of Upper Paleozoic Diplopoda". Journal of Paleontology. 37 (1): 167–174. ISSN 0022-3360. JSTOR 1301419.
  28. ^ Eble, Cortland F. (2022). "Appalachian coal bed palynofloras: changes in composition through time and comparison with other areas". Geological Society, London, Special Publications. 512 (1): 791–812. Bibcode:2022GSLSP.512..791E. doi:10.1144/SP512-2021-131. ISSN 0305-8719.