Jump to content

Canada warbler: Difference between revisions

From Wikipedia, the free encyclopedia
Content deleted Content added
Citation bot (talk | contribs)
Add: date, volume. | Use this bot. Report bugs. | Suggested by Abductive | Category:Taxonbars with automatically added original combinations | #UCB_Category 1278/4153
Behavior and ecology: add author-link Judy Gradwohl
 
(16 intermediate revisions by 14 users not shown)
Line 1: Line 1:
{{short description|Species of bird}}
{{Short description|Species of bird}}
{{Speciesbox
{{Speciesbox
| name = Canada warbler
| name = Canada warbler
| status = LC
| status = LC
| status_system = IUCN3.1
| status_system = IUCN3.1
| status_ref = <ref name =IUCN>BirdLife International. 2016. Cardellina canadensis. The IUCN Red List of Threatened Species 2016: e.T22721882A94737489. https://dx.doi.org/10.2305/IUCN.UK.2016-3.RLTS.T22721882A94737489.en. Downloaded on 09 April 2021.</ref>
| status_ref = <ref name =IUCN>{{cite iucn |author=BirdLife International. |year=2016 |title=''Cardellina canadensis'' |volume=2016 |page=e.T22721882A94737489 |doi=10.2305/IUCN.UK.2016-3.RLTS.T22721882A94737489.en |access-date=9 April 2021}}</ref>
| image = 8G7D5475-Canada.jpg
| image = 8G7D5475-Canada.jpg
| image_caption = Male
| image_caption = Male
Line 18: Line 18:
}}
}}


The '''Canada warbler''' (''Cardellina canadensis'') is a small boreal [[songbird]] of the [[New World warbler]] family (Parulidae). It summers in Canada and northeastern United States and winters in northern South America.
The '''Canada warbler''' ('''''Cardellina canadensis''''') is a small boreal [[songbird]] of the [[New World warbler]] family (Parulidae). It summers in Canada and northeastern United States and winters in northern South America.


==Taxonomy==
==Taxonomy==
In 1760 the French zoologist [[Mathurin Jacques Brisson]] included a description of the Canada warbler in his ''Ornithologie'' based on a specimen collected in Canada. He used the French name ''Le gobe-mouche cendré de Canada'' and the Latin name ''Muscicapa Canadensis Cinerea''.<ref name=brisson>{{ cite book | last=Brisson | first=Mathurin Jacques | author-link=Mathurin Jacques Brisson | year=1760 | title=Ornithologie, ou, Méthode contenant la division des oiseaux en ordres, sections, genres, especes & leurs variétés | volume=Volume 2 | language=fr, la | place=Paris | publisher=Jean-Baptiste Bauche | pages=406–408, Plate 39 fig 4 | url=https://biodiversitylibrary.org/page/36011718 }} The two stars (**) at the start of the section indicates that Brisson based his description on the examination of a specimen.</ref> Although Brisson coined Latin names, these do not conform to the [[binomial nomenclature|binomial system]] and are not recognised by the [[International Commission on Zoological Nomenclature]].<ref name=allen>{{cite journal |last=Allen | first=J.A. | author-link=Joel Asaph Allen | year=1910 | title=Collation of Brisson's genera of birds with those of Linnaeus |journal=Bulletin of the American Museum of Natural History |volume=28 | pages=317–335 | hdl=2246/678 }}</ref> When in 1766 the Swedish naturalist [[Carl Linnaeus]] updated his ''[[Systema Naturae]]'' for the [[12th edition of Systema Naturae|twelfth edition]], he added 240 species that had been previously described by Brisson.<ref name=allen/> One of these was the Canada warbler. Linnaeus included a brief description, coined the [[binomial name]] ''Muscicapa canadensis'' and cited Brisson's work.<ref>{{cite book | last=Linnaeus | first=Carl | author-link=Carl Linnaeus | year=1766 | title=Systema naturae : per regna tria natura, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis | edition=12th | volume=Volume 1, Part 1 | publisher=Laurentii Salvii | place=Holmiae (Stockholm) | language=la | page=327 | url=https://biodiversitylibrary.org/page/42946523 }}</ref> The species is now placed in the [[genus]] ''[[Cardellina]]'' that was introduced by the French naturalist [[Charles Lucien Jules Laurent Bonaparte|Charles Lucien Bonaparte]] in 1850.<ref>{{cite book | last=Bonaparte | first=Charles Lucien | author-link=Charles Lucien Bonaparte | year=1850 | title=Conspectus generum avium | volume=1 | language=la | place=Lugduni Batavorum [Leiden] | publisher=E.J. Brill | page=312 | url=https://biodiversitylibrary.org/page/43560473 }}</ref> The species is [[monotypic]].<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | year=2018 | title=New World warblers, mitrospingid tanagers | work=World Bird List Version 8.1 | url=http://www.worldbirdnames.org/bow/warblers/ | publisher=International Ornithologists' Union | access-date=1 May 2018 }}</ref>
In 1760 the French zoologist [[Mathurin Jacques Brisson]] included a description of the Canada warbler in his ''Ornithologie'' based on a specimen collected in Canada. He used the French name ''Le gobe-mouche cendré de Canada'' and the Latin name ''Muscicapa Canadensis Cinerea''.<ref name=brisson>{{ cite book | last=Brisson | first=Mathurin Jacques | author-link=Mathurin Jacques Brisson | year=1760 | title=Ornithologie, ou, Méthode contenant la division des oiseaux en ordres, sections, genres, especes & leurs variétés | volume=2 | language=fr, la | place=Paris | publisher=Jean-Baptiste Bauche | pages=406–408, Plate 39 fig 4 | url=https://biodiversitylibrary.org/page/36011718 }} The two stars (**) at the start of the section indicates that Brisson based his description on the examination of a specimen.</ref> Although Brissn coined Latin names, these do not conform to the [[binomial nomenclature|binomial system]] and are not recognised by the [[International Commission on Zoological Nomenclature]].<ref name=allen>{{cite journal |last=Allen | first=J.A. | author-link=Joel Asaph Allen | year=1910 | title=Collation of Brisson's genera of birds with those of Linnaeus |journal=Bulletin of the American Museum of Natural History |volume=28 | pages=317–335 | hdl=2246/678 }}</ref> When in 1766 the Swedish naturalist [[Carl Linnaeus]] updated his ''[[Systema Naturae]]'' for the [[12th edition of Systema Naturae|twelfth edition]], he added 240 species that had been previously described by Brisson.<ref name=allen/> One of these was the Canada warbler. Linnaeus included a brief description, coined the [[binomial name]] ''Muscicapa canadensis,'' and cited Brisson's work.<ref>{{cite book | last=Linnaeus | first=Carl | author-link=Carl Linnaeus | year=1766 | title=Systema naturae : per regna tria natura, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis | edition=12th | volume=1, Part 1 | publisher=Laurentii Salvii | place=Holmiae (Stockholm) | language=la | page=327 | url=https://biodiversitylibrary.org/page/42946523 }}</ref> The species is now placed in the [[genus]] ''[[Cardellina]]'' that was introduced by the French naturalist [[Charles Lucien Jules Laurent Bonaparte|Charles Lucien Bonaparte]] in 1850.<ref>{{cite book | last=Bonaparte | first=Charles Lucien | author-link=Charles Lucien Bonaparte | year=1850 | title=Conspectus generum avium | volume=1 | language=la | place=Lugduni Batavorum [Leiden] | publisher=E.J. Brill | page=312 | url=https://biodiversitylibrary.org/page/43560473 }}</ref> The species is [[monotypic]].<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | year=2018 | title=New World warblers, mitrospingid tanagers | work=World Bird List Version 8.1 | url=http://www.worldbirdnames.org/bow/warblers/ | publisher=International Ornithologists' Union | access-date=1 May 2018 }}</ref>


==Description==
==Description==
The Canada warbler is sometimes called the "necklaced warbler," because of the band of dark streaks across its chest. The adults have minimal sexual dimorphism, although the male's "necklace" is darker and more conspicuous and also has a longer tail. Adults are {{convert|12–15|cm|in|abbr=on}} long, have a wingspan of {{convert|17–22|cm|in|abbr=on}} and weigh {{convert|9–13|g|oz|abbr=on}}.<ref name=Cornelllh>{{cite web|url=https://www.allaboutbirds.org/guide/Canada_Warbler/lifehistory|title=Canada Warbler—Life History|website=Cornell Lab of Ornithology|access-date=December 15, 2016}}</ref>
The Canada warbler is sometimes called the "necklaced warbler", because of the band of dark streaks across its chest. The adults have minimal sexual dimorphism, although the male's "necklace" is darker and more conspicuous and also has a longer tail. Adults are {{convert|12–15|cm|in|abbr=on}} long, have a wingspan of {{convert|17–22|cm|in|abbr=on}}, and weigh {{convert|9–13|g|oz|abbr=on}}.<ref name=Cornelllh>{{cite web|url=https://www.allaboutbirds.org/guide/Canada_Warbler/lifehistory|title=Canada Warbler—Life History|website=Cornell Lab of Ornithology|access-date=December 15, 2016}}</ref>


The chest, throat and belly of the bird is yellow, and its back is dark grey. It has no wingbars or tail spots, but the underside of the tail is white. It has a yellow line in front of its eye in the direction of the beak, but the most striking facial feature is the white eyerings or "spectacles."<ref name=Cornellid>{{cite web|url=https://www.allaboutbirds.org/guide/Canada_Warbler/id|title=Canada Warbler—Identification|website=Cornell Lab of Ornithology|access-date=December 15, 2016}}</ref> Immature specimens have similar coloration as adults but duller and with less pronounced facial features.<ref name=Cornellid/>
The chest, throat, and belly of the bird are yellow, and its back is dark grey. It has no wingbars or tail spots, but the underside of the tail is white. It has a yellow line in front of its eye in the direction of the beak, but the most striking facial feature is the white eyerings or "spectacles".<ref name=Cornellid>{{cite web|url=https://www.allaboutbirds.org/guide/Canada_Warbler/id|title=Canada Warbler—Identification|website=Cornell Lab of Ornithology|access-date=December 15, 2016}}</ref> Immature specimens have similar coloration as adults but duller and with less pronounced facial features.<ref name=Cornellid/>


===Song===
===Song===
The {{audio|Cardellina_canadensis_-_Canada_Warbler_-_XC78875.ogg|song}} of this bird is loud and highly variable, resembling ''chip chewy sweet dichetty''. Their calls are low ''chup'''s.
The {{audio|Cardellina_canadensis_-_Canada_Warbler_-_XC78875.ogg|song}} of this bird is loud and highly variable, resembling ''chip chewy sweet dichetty''. Their calls are low ''chup'''s.


A 2013 study showed that male Canada warblers have two performance-encoded song types. In Mode I, used mostly during the day, when unpaired either alone or near a female during early nesting, involves stereotyped songs sung slowly and regularly. Mode II, used at dawn, after pairing and when near another male, involves variable songs, sung rapidly with irregular rhythm and chippipng between songs. Most of the phrases used were common to both modes, a feature unique among paulidswhich ordinarily have an individual's repertoire separated into two distinct parts.<ref>{{cite journal|jstor=10.1525/auk.2013.13059|title= Two song categories in the Canada Warbler (''Cardellina canadensis'')|first1=Alana D.|last1=Demko|first2=Leonard R.|last2=Reitsma|first3=Cynthia A.|last3=Staicer|journal=The Auk|volume=130|number=4|date=October 2013|pages=609–16|doi=10.1525/auk.2013.13059|s2cid= 83993091|access-date=28 April 2019|url=https://www.researchgate.net/publication/274239869}}</ref>
A 2013 study showed that male Canada warblers have two performance-encoded song types. Mode I, used mostly during the day, when unpaired either alone or near a female during early nesting, involves stereotyped songs sung slowly and regularly. Mode II, used at dawn, after pairing and when near another male, involves variable songs, sung rapidly with irregular rhythm and chirping between songs. Most of the phrases used were common to both modes, a feature unique among parulids, which ordinarily have an individual's repertoire separated into two distinct parts.<ref>{{cite journal|jstor=10.1525/auk.2013.13059|title= Two song categories in the Canada Warbler (''Cardellina canadensis'')|first1=Alana D.|last1=Demko|first2=Leonard R.|last2=Reitsma|first3=Cynthia A.|last3=Staicer|journal=The Auk|volume=130|number=4|date=October 2013|pages=609–16|doi=10.1525/auk.2013.13059|s2cid= 83993091|access-date=28 April 2019|url=https://www.researchgate.net/publication/274239869|doi-access=free}}</ref>


In 2000, a female Canada warbler (or a post-hatching year old male that failed to moult, something never before observed) in [[Giles County, Virginia|Giles County]], [[Virginia]] was observed singing. Its repertoire consisted of a repeated song of 12 to 13 notes as well as several shorter songs consisting of the first five or six notes of the longer song. The bird did not respond to playback of its own song or a recording of a male. Although female singing among the parulids has long been considered "idiosyncratic," singing by female Canada warblers is supported by observation of female singing in [[Congener (biology)|congener]] [[Wilson's warbler]] and the closely related [[hooded warbler]].<ref>{{cite journal|first=Matthew A.|last=Etterson|title= An Observation of Singing by a Female-Plumaged Canada Warbler|journal=Southeastern Naturalist|volume=2|number=2|year=2003|pages=419–22|jstor=3878011|doi=10.1656/1528-7092(2003)002[0419:aoosba]2.0.co;2}}</ref>
In 2000, a female Canada warbler (or a post-hatching year-old male that failed to moult, something never before observed) in [[Giles County, Virginia|Giles County]], [[Virginia]] was observed singing. Its repertoire consisted of a repeated song of 12 to 13 notes as well as several shorter songs consisting of the first five or six notes of the longer song. The bird did not respond to the playback of its own song or a recording of a male. Although female singing among the parulids has long been considered "idiosyncratic", singing by female Canada warblers is supported by the observation of female singing in [[Congener (biology)|congener]] [[Wilson's warbler]] and the closely related [[hooded warbler]].<ref>{{cite journal|first=Matthew A.|last=Etterson|title= An Observation of Singing by a Female-Plumaged Canada Warbler|journal=Southeastern Naturalist|volume=2|number=2|year=2003|pages=419–22|jstor=3878011|doi=10.1656/1528-7092(2003)002[0419:aoosba]2.0.co;2|s2cid=86052159 }}</ref>


==Distribution and habitat==
==Distribution and habitat==
During the breeding season 82% of the population can be found in Canada and 18% in the United States.<ref name=Cornelllh/> In Canada the summer range extends from southeastern [[Yukon]] to [[Nova Scotia]]. In the United States the range extends from northern [[Minnesota]] to northern [[Pennsylvania]], east to [[Long Island]], [[New York (state)|New York]]. It also nests in the high [[Appalachian Mountains|Appalachians]] as far south as [[Georgia (U.S. state)|Georgia]].<ref name=Audubon>{{cite web|url=http://www.audubon.org/field-guide/bird/canada-warbler|title=Canada Warbler|work=Audubon Guide to North American Birds|date=13 November 2014|access-date=December 15, 2016}}</ref> In winter the Canada warbler's range extends from [[Guyana]] to northwestern [[Bolivia]] around the northern and western side of the Andean crest.<ref name=Boreal>{{cite web|url=http://www.borealbirds.org/bird/canada-warbler|title=Canada warbler|work=Boreal Songbird Initiative|date=25 February 2014|access-date=December 15, 2016}}</ref>
During the breeding season 82% of the population can be found in Canada and 18% in the United States.<ref name=Cornelllh/> In Canada, the summer range extends from southeastern [[Yukon]] to [[Nova Scotia]]. In the United States, the range extends from northern [[Minnesota]] to northern [[Pennsylvania]], east to [[Long Island]], [[New York (state)|New York]]. It also nests in the high [[Appalachian Mountains|Appalachians]] as far south as [[Georgia (U.S. state)|Georgia]].<ref name=Audubon>{{cite web|url=http://www.audubon.org/field-guide/bird/canada-warbler|title=Canada Warbler|work=Audubon Guide to North American Birds|date=13 November 2014|access-date=December 15, 2016}}</ref> In winter, the Canada warbler's range extends from [[Guyana]] to northwestern [[Bolivia]] around the northern and western side of the Andean crest.<ref name=Boreal>{{cite web|url=http://www.borealbirds.org/bird/canada-warbler|title=Canada warbler|work=Boreal Songbird Initiative|date=25 February 2014|access-date=December 15, 2016}}</ref>


In both summer and winter seasons the Canada warbler inhabits moist thickets. During the breeding season the bird "nests in riparian thickets, brushy ravines, forest bogs, etc. at a wide range of elevations and across a variety of forest types. In the northwestern parts of its range it frequents aspen forests; in the center of the range, it is found in forested wetlands and swamps; and in the south it occupies montane rhododendron thickets.<ref name=Boreal/>"
In both summer and winter seasons, the Canada warbler inhabits moist thickets. During the breeding season, the bird "nests in riparian thickets, brushy ravines, forest bogs, etc. at a wide range of elevations and across a variety of forest types. In the northwestern parts of its range, it frequents aspen forests; in the center of the range, it is found in forested wetlands and swamps; and in the south, it occupies montane rhododendron thickets.<ref name=Boreal/>"
In the winter it prefers mid- and upper-elevation habitats.<ref name=Boreal/> In northern [[Minnesota]] a study found that Canada warblers inhabited the shrub-forest edge, rather than marture forests or open fields with shrub.<ref>{{cite journal|url=http://collins.lternet.edu/biblio/habitat-relationships-wood-warblers-parulidae-northern-central-minnesota|last1=Collins|first1=Scott L.|last2=James|first2=Frances C.|last3=Risser|first3=Paul G.|title=Habitat Relationships of Wood Warblers Parulidae in Northern Central Minnesota|journal=Oikos|volume=39|number=1|year=1982|pages=50–58|access-date=December 18, 2016|doi=10.2307/3544530|jstor=3544530}}</ref> In New England the Canada warbler was found to be "disturbance specialists" moving into patches of forests recovering from wind throw or timber removal.<ref>{{cite journal|jstor=27744588|first1=Jameson F.|last1=Chace|first2=Steven D.|last2=Faccio|first3=Abraham|last3=Chacko|title=Canada Warbler Habitat Use of Northern Hardwoods in Vermont|journal=Northeastern Naturalist|volume=16|number=4|year=2009|pages=491–500 |url=https://www.researchgate.net/publication/232683973|doi=10.1656/045.016.n401|s2cid=86430663}}</ref> Because of its preference for low-height foraging in deciduous forests, it may be bounded at higher elevations as suitable habitat disappears and suffer competition from the [[black-throated blue warbler]] which prefers similar habitats.<ref>{{cite journal|last=Sabo|first=Stephen R.|title=Niche and Habitat Relations in Subalpine Bird Communities of the White Mountains of New Hampshire|journal=Ecological Monographs|volume=50|number=2|date=June 1980|doi=10.2307/1942481|pages=241–59|jstor=1942481}}</ref>
In the winter, it prefers mid- and upper-elevation habitats.<ref name=Boreal/> In northern [[Minnesota]], a study found that Canada warblers inhabited the shrub-forest edge, rather than mature forests or open fields with shrub.<ref>{{cite journal|url=http://collins.lternet.edu/biblio/habitat-relationships-wood-warblers-parulidae-northern-central-minnesota|last1=Collins|first1=Scott L.|last2=James|first2=Frances C.|last3=Risser|first3=Paul G.|title=Habitat Relationships of Wood Warblers Parulidae in Northern Central Minnesota|journal=Oikos|volume=39|number=1|year=1982|pages=50–58|access-date=December 18, 2016|doi=10.2307/3544530|jstor=3544530|bibcode=1982Oikos..39...50C }}</ref> In New England, the Canada warbler was found to be "disturbance specialists" moving into patches of forests recovering from wind throw or timber removal.<ref>{{cite journal|jstor=27744588|first1=Jameson F.|last1=Chace|first2=Steven D.|last2=Faccio|first3=Abraham|last3=Chacko|title=Canada Warbler Habitat Use of Northern Hardwoods in Vermont|journal=Northeastern Naturalist|volume=16|number=4|year=2009|pages=491–500 |url=https://www.researchgate.net/publication/232683973|doi=10.1656/045.016.n401|s2cid=86430663}}</ref> Because of its preference for low-height foraging in deciduous forests, it may be bounded at higher elevations as suitable habitat disappears and it suffers competition from the [[black-throated blue warbler]] which prefers similar habitats.<ref>{{cite journal|last=Sabo|first=Stephen R.|title=Niche and Habitat Relations in Subalpine Bird Communities of the White Mountains of New Hampshire|journal=Ecological Monographs|volume=50|number=2|date=June 1980|doi=10.2307/1942481|pages=241–59|jstor=1942481|bibcode=1980EcoM...50..241S }}</ref>


Two accidentals have been observed in Europe. The first a moribund male caught in [[Sandgerði]], [[Iceland]] on September 29, 1973. The second was a first winter, probably female observed for five days in October 2006 in [[County Clare]], [[Ireland]].<ref>{{cite journal|url=http://www.irbc.ie/notes/pdf/CanadaWarblerInCo.Clare(MauriceHanafin).pdf|first=Maurice|last=Hanafin|title=The Canada Warbler in County Clare—The Second for Western Palearctic|journal=Birding World|volume=19|number=10|year=2006|pages=429–34|access-date=December 17, 2016}}</ref>
Two accidentals have been observed in Europe. The first, a moribund male was caught in [[Sandgerði]], [[Iceland]] on September 29, 1973. The second was a first winter, probably female observed for five days in October 2006 in [[County Clare]], [[Ireland]].<ref>{{cite journal|url=http://www.irbc.ie/notes/pdf/CanadaWarblerInCo.Clare(MauriceHanafin).pdf|first=Maurice|last=Hanafin|title=The Canada Warbler in County Clare—The Second for Western Palearctic|journal=Birding World|volume=19|number=10|year=2006|pages=429–34|access-date=December 17, 2016}}</ref>


===Migration===
===Migration===
Line 48: Line 48:
==Behavior and ecology==
==Behavior and ecology==
===Breeding===
===Breeding===
At least 60–65% of the population nests in boreal forests in Canada, the Great Lakes region of the United States, New England and through the Appalachians.<ref name=Boreal/> The birds are at least seasonally monogamous. Sightings of pairs during migration in Panama have led to the conclusion that they are permanently monogamous.<ref>{{cite journal|last1=Greenberg|first1=Russell S.|last2=Gradwohl|first2=Judy A.|title=Observations of Paired Canada Warblers ''Wilsonia canadensis'' during Migration in Panama|journal=Ibis|volume=122|number=4|date=October 1980|pages=509–12|doi=10.1111/j.1474-919X.1980.tb00907.x}}</ref><ref name=Cornelllh/> This conclusion, however, is contradicted by the sexes' wintering at different elevations.<ref name=Boreal/>
At least 60–65% of the population nests in boreal forests in Canada, the Great Lakes region of the United States, New England and through the Appalachians.<ref name=Boreal/> The birds are at least seasonally monogamous. Sightings of pairs during migration in Panama have led to the conclusion that they are permanently monogamous.<ref>{{cite journal|last1=Greenberg|first1=Russell S.|last2=Gradwohl|first2=Judy A. |author-link2=Judy Gradwohl |title=Observations of Paired Canada Warblers ''Wilsonia canadensis'' during Migration in Panama|journal=Ibis|volume=122|number=4|date=October 1980|pages=509–12|doi=10.1111/j.1474-919X.1980.tb00907.x}}</ref><ref name=Cornelllh/> This conclusion, however, is contradicted by the sexes' wintering at different elevations.<ref name=Boreal/>


Males arrive at the breeding grounds in the first two weeks of May.<ref name=Audubon/> Females build the nests on or very close to the ground in dense cover. The nests are made up of root masses, hummocks, stumps, stream banks, mossy logs, and sometimes leaf litter and grass clumps. Moss covering is frequent.<ref name=Boreal/>
Males arrive at the breeding grounds in the first two weeks of May.<ref name=Audubon/> Females build the nests on or very close to the ground in dense cover. The nests are made up of root masses, hummocks, stumps, stream banks, mossy logs, and sometimes leaf litter and grass clumps. Moss covering is frequent.<ref name=Boreal/>
Line 54: Line 54:
The female lays four to five eggs and incubates for about 12 days. The chicks remain in the nest for about 10 days after hatching and are dependent on their parents for two to three weeks after they leave the nest.<ref name=SaR>{{cite web |url=http://www.sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=1008 |title=Species Profile: Canada Warbler |author=<!--Staff writer(s); no by-line.--> |date=May 29, 2014 |website=Species at Risk Public Registry |publisher=Government of Canada |access-date=June 5, 2014 |archive-url=https://web.archive.org/web/20140909231344/http://www.sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=1008 |archive-date=September 9, 2014 |url-status=dead }}</ref>
The female lays four to five eggs and incubates for about 12 days. The chicks remain in the nest for about 10 days after hatching and are dependent on their parents for two to three weeks after they leave the nest.<ref name=SaR>{{cite web |url=http://www.sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=1008 |title=Species Profile: Canada Warbler |author=<!--Staff writer(s); no by-line.--> |date=May 29, 2014 |website=Species at Risk Public Registry |publisher=Government of Canada |access-date=June 5, 2014 |archive-url=https://web.archive.org/web/20140909231344/http://www.sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=1008 |archive-date=September 9, 2014 |url-status=dead }}</ref>


The age at which the young leave the nest is not known. Once independent they spend almost all their time in the understory, on the ground or in bushes.<ref name=Audubon/> The post-juvenile bird undergoes a partial moult involving all body feathres and wing coverlets. This may be completed before the first migration.{{sfn|Hanafin|2006|p=434}}
The age at which the young leave the nest is not known. Once independent they spend almost all their time in the understory, on the ground or in bushes.<ref name=Audubon/> The post-juvenile bird undergoes a partial moult involving all body feathers and wing coverlets. This may be completed before the first migration.{{sfn|Hanafin|2006|p=434}}


The oldest known specimen was a male found in Quebec in 1982 at least 8 years old, having been banded in 1975.<ref>{{cite web|url=http://www.pwrc.usgs.gov/bbl/longevity/longevity_main.cfm|author=USGS Patuxent Wildlife Research Center|title=Longevity Records of North American Birds|year=2016|publisher=U.S. Geological Survey. Department of the Interior|access-date=December 16, 2016}}</ref>
The oldest known specimen was a male found in Quebec in 1982 at least 8 years old, having been banded in 1975.<ref>{{cite web|url=http://www.pwrc.usgs.gov/bbl/longevity/longevity_main.cfm|author=USGS Patuxent Wildlife Research Center|title=Longevity Records of North American Birds|year=2016|publisher=U.S. Geological Survey. Department of the Interior|access-date=December 16, 2016}}</ref>
Line 67: Line 67:
Partners in Flight estimates a global population of 4 million,<ref>{{cite web|url=http://rmbo.org/pifassessment/Database.aspx|title=Species Assessment Database|work=Partners in Flight|access-date=December 15, 2016|archive-url=https://web.archive.org/web/20160821000851/http://rmbo.org/pifassessment/Database.aspx|archive-date=August 21, 2016|url-status=dead}}</ref> while the American Bird Conservancy estimates that 1.5 million individuals exist.<ref name=ABC>{{cite web|url=https://abcbirds.org/bird/canada-warbler/?gclid=CjwKEAiAm8nCBRD7xLj-2aWFyz8SJAAQNalabtPqHXz1dLpot8bhZkv8vLaciiauRiE0cI4QKXUvKBoC_Jrw_wcB|title=Canada Warbler|work=American Bird Conservancy|access-date=December 15, 2016}}</ref>
Partners in Flight estimates a global population of 4 million,<ref>{{cite web|url=http://rmbo.org/pifassessment/Database.aspx|title=Species Assessment Database|work=Partners in Flight|access-date=December 15, 2016|archive-url=https://web.archive.org/web/20160821000851/http://rmbo.org/pifassessment/Database.aspx|archive-date=August 21, 2016|url-status=dead}}</ref> while the American Bird Conservancy estimates that 1.5 million individuals exist.<ref name=ABC>{{cite web|url=https://abcbirds.org/bird/canada-warbler/?gclid=CjwKEAiAm8nCBRD7xLj-2aWFyz8SJAAQNalabtPqHXz1dLpot8bhZkv8vLaciiauRiE0cI4QKXUvKBoC_Jrw_wcB|title=Canada Warbler|work=American Bird Conservancy|access-date=December 15, 2016}}</ref>


Threats to the Canada warbler include forest fragmentation; over-browsing of the understory by deer, acid rain, and the spread of the woolly adelgid (a killer of fir and hemlock trees).<ref name=ABC/> Owing to these factors the Breeding Bird Survey data show a population decline of 3.2 percent per year throughout the Canada warbler's breeding range, with the greatest declines in the Northeast.<ref>{{cite web|url=http://www.mbr-pwrc.usgs.gov/cgi-bin/atlasa15.pl?06860&1&15&csrfmiddlewaretoken=3YKakk7LxT2ki6NSpl4mstudYCqdW02C|author=USGS Paxtuxent Wildlife Research Center|title=Canada Warbler, ''Cardellina canadensis'': North American Breeding Bird Survey Trend Results|publisher=U.S. Geological Survey. Department of Interior|year=2016|access-date=December 16, 2016}}</ref><ref name=ABC/> The species has been assessed as "threatened" by the [[Committee on the Status of Endangered Wildlife in Canada]].<ref name=COSEWIC>{{cite web |url=http://www.cosewic.gc.ca/eng/sct1/searchdetail_e.cfm?id=1008 |title=Warbler, Canada |author=<!--Staff writer(s); no by-line.--> |date=November 11, 2011 |website=Committee on the Status of Endangered Wildlife in Canada (COSEWIC) |publisher=Government of Canada |access-date=June 5, 2014 |archive-url=https://web.archive.org/web/20140606234613/http://www.cosewic.gc.ca/eng/sct1/searchdetail_e.cfm?id=1008# |archive-date=2014-06-06 |url-status=dead }}</ref> The [[IUCN]], however, ranks the Canada warbler as a species of least concern.<ref name =IUCN />
Threats to the Canada warbler include forest fragmentation; over-browsing of the understory by deer, acid rain, and the spread of the woolly adelgid (a killer of fir and hemlock trees).<ref name=ABC/> Owing to these factors the Breeding Bird Survey data show a population decline of 3.2 percent per year throughout the Canada warbler's breeding range, with the greatest declines in the Northeast.<ref>{{cite web|url=http://www.mbr-pwrc.usgs.gov/cgi-bin/atlasa15.pl?06860&1&15&csrfmiddlewaretoken=3YKakk7LxT2ki6NSpl4mstudYCqdW02C|author=USGS Paxtuxent Wildlife Research Center|title=Canada Warbler, ''Cardellina canadensis'': North American Breeding Bird Survey Trend Results|publisher=U.S. Geological Survey. Department of Interior|year=2016|access-date=December 16, 2016}}</ref><ref name=ABC/> The species has been assessed as "threatened" by the [[Committee on the Status of Endangered Wildlife in Canada]].<ref name=COSEWIC>{{cite web |url=http://www.cosewic.gc.ca/eng/sct1/searchdetail_e.cfm?id=1008 |title=Warbler, Canada |author=<!--Staff writer(s); no by-line.--> |date=November 11, 2011 |website=Committee on the Status of Endangered Wildlife in Canada (COSEWIC) |publisher=Government of Canada |access-date=June 5, 2014 |archive-url=https://web.archive.org/web/20140606234613/http://www.cosewic.gc.ca/eng/sct1/searchdetail_e.cfm?id=1008 |archive-date=2014-06-06 |url-status=dead }}</ref> The [[IUCN]], however, ranks the Canada warbler as a species of least concern.<ref name =IUCN />


The Canada warbler is protected at the federal level in both Canada and the United States.<ref name=SaR />
The Canada warbler is protected at the federal level in both Canada and the United States.<ref name=SaR />
Line 73: Line 73:
==In art==
==In art==
<!-- Deleted image removed: [[File:Audubons Canada Warbler - Plate 5 - Birds of America.jpg|thumb|200px|left|Audubon's Bonaparte's flycatcher (Canada warbler) from ''[[The Birds of America]]'']] -->
<!-- Deleted image removed: [[File:Audubons Canada Warbler - Plate 5 - Birds of America.jpg|thumb|200px|left|Audubon's Bonaparte's flycatcher (Canada warbler) from ''[[The Birds of America]]'']] -->
[[John James Audubon]] illustrates the Canada warbler in ''[[Birds of America (book)|Birds of America]]'' (published, London 1827-38) as Plate 73 entitled "Bonaparte's Flycatching-Warbler—''Muscicapa bonapartii''." The single female (now properly identified as a Canada warbler) is shown perched in a great magnolia (''[[Magnolia grandiflora]]'') branch that was painted by [[Joseph Mason (artist)|Joseph Mason]]. The final, combined image was engraved and colored by [[Havell family#Robert Havell Jr.|Robert Havell Jr.]] at the Havell workshops in London. The original painting was purchased by the [[New York Historical Society]].
[[John James Audubon]] illustrates the Canada warbler in ''[[Birds of America (book)|Birds of America]]'' (published, London 1827–38) as Plate 73 entitled "Bonaparte's Flycatching-Warbler—''Muscicapa bonapartii''." The single female (now properly identified as a Canada warbler) is shown perched on a great magnolia (''[[Magnolia grandiflora]]'') branch that was painted by [[Joseph Mason (artist)|Joseph Mason]]. The final, combined image was engraved and colored by [[Havell family#Robert Havell Jr.|Robert Havell Jr.]] at the Havell workshops in London. The original painting was purchased by the [[New York Historical Society]].


==References==
==References==
Line 87: Line 87:
* {{cite journal | last1 = Caroline | first1 = G | last2 = Marcel | first2 = D | last3 = Jean-Pierre | first3 = LS | last4 = Jean | first4 = H | year = 2004 | title = Are temperate mixedwood forests perceived by birds as a distinct forest type? | journal = Canadian Journal of Forest Research | volume = 34 | issue = 9| page = 1895 | doi=10.1139/x04-087}}
* {{cite journal | last1 = Caroline | first1 = G | last2 = Marcel | first2 = D | last3 = Jean-Pierre | first3 = LS | last4 = Jean | first4 = H | year = 2004 | title = Are temperate mixedwood forests perceived by birds as a distinct forest type? | journal = Canadian Journal of Forest Research | volume = 34 | issue = 9| page = 1895 | doi=10.1139/x04-087}}
* {{cite journal | last1 = Christian | first1 = DP | last2 = Hanowski | first2 = JM | last3 = Reuvers-House | first3 = M | last4 = Niemi | first4 = GJ | last5 = Blake | first5 = JG | last6 = Berguson | first6 = WE | year = 1996 | title = Effects of mechanical strip thinning of aspen on small mammals and breeding birds in northern Minnesota, U.S.A | journal = Canadian Journal of Forest Research | volume = 26 | issue = 7| pages = 1284–1294 | doi=10.1139/x26-143}}
* {{cite journal | last1 = Christian | first1 = DP | last2 = Hanowski | first2 = JM | last3 = Reuvers-House | first3 = M | last4 = Niemi | first4 = GJ | last5 = Blake | first5 = JG | last6 = Berguson | first6 = WE | year = 1996 | title = Effects of mechanical strip thinning of aspen on small mammals and breeding birds in northern Minnesota, U.S.A | journal = Canadian Journal of Forest Research | volume = 26 | issue = 7| pages = 1284–1294 | doi=10.1139/x26-143}}
* {{cite journal | last1 = Crawford | first1 = HS | last2 = Jennings | first2 = DT | year = 1989 | title = Predation by Birds on Spruce Budworm Choristoneura-Fumiferana Functional Numerical and Total Responses | journal = Ecology | volume = 70 | issue = 1| pages = 152–163 | doi=10.2307/1938422| jstor = 1938422 }}
* {{cite journal | last1 = Crawford | first1 = HS | last2 = Jennings | first2 = DT | year = 1989 | title = Predation by Birds on Spruce Budworm Choristoneura-Fumiferana Functional Numerical and Total Responses | journal = Ecology | volume = 70 | issue = 1| pages = 152–163 | doi=10.2307/1938422| jstor = 1938422 | bibcode = 1989Ecol...70..152C }}
* {{cite journal | last1 = Dunn | first1 = EH | last2 = Nol | first2 = E | year = 1980 | title = Age Related Migratory Behavior of Warblers | journal = Journal of Field Ornithology | volume = 51 | issue = 3| pages = 254–269 }}
* {{cite journal | last1 = Dunn | first1 = EH | last2 = Nol | first2 = E | year = 1980 | title = Age Related Migratory Behavior of Warblers | journal = Journal of Field Ornithology | volume = 51 | issue = 3| pages = 254–269 }}
* {{cite journal | last1 = Golet | first1 = FC | last2 = Wang | first2 = Y | last3 = Merrow | first3 = JS | last4 = DeRagon | first4 = WR | year = 2001 | title = Relationship between habitat and landscape features and the avian community of red maple swamps in southern Rhode Island | journal = Wilson Bulletin | volume = 113 | issue = 2| pages = 217–227 | doi=10.1676/0043-5643(2001)113[0217:rbhalf]2.0.co;2}}
* {{cite journal | last1 = Golet | first1 = FC | last2 = Wang | first2 = Y | last3 = Merrow | first3 = JS | last4 = DeRagon | first4 = WR | year = 2001 | title = Relationship between habitat and landscape features and the avian community of red maple swamps in southern Rhode Island | journal = Wilson Bulletin | volume = 113 | issue = 2| pages = 217–227 | doi=10.1676/0043-5643(2001)113[0217:rbhalf]2.0.co;2| s2cid = 86169562 }}
* {{cite journal | last1 = Hobson | first1 = KA | last2 = Bayne | first2 = E | year = 2000 | title = The effects of stand age on avian communities in aspen-dominated forests of central Saskatchewan, Canada | journal = Forest Ecology & Management | volume = 136 | issue = 1–3| pages = 121–134 | doi=10.1016/s0378-1127(99)00287-x| citeseerx = 10.1.1.532.1080 }}
* {{cite journal | last1 = Hobson | first1 = KA | last2 = Bayne | first2 = E | year = 2000 | title = The effects of stand age on avian communities in aspen-dominated forests of central Saskatchewan, Canada | journal = Forest Ecology & Management | volume = 136 | issue = 1–3| pages = 121–134 | doi=10.1016/s0378-1127(99)00287-x| bibcode = 2000ForEM.136..121H | citeseerx = 10.1.1.532.1080 }}
* {{cite journal | last1 = Hobson | first1 = KA | last2 = Schieck | first2 = J | s2cid = 28285190 | year = 1999 | title = Changes in bird communities in boreal mixedwood forest: Harvest and wildfire effects over 30 years | journal = Ecological Applications | volume = 9 | issue = 3| pages = 849–863 | doi=10.1890/1051-0761(1999)009[0849:cibcib]2.0.co;2}}
* {{cite journal | last1 = Hobson | first1 = KA | last2 = Schieck | first2 = J | s2cid = 28285190 | year = 1999 | title = Changes in bird communities in boreal mixedwood forest: Harvest and wildfire effects over 30 years | journal = Ecological Applications | volume = 9 | issue = 3| pages = 849–863 | doi=10.1890/1051-0761(1999)009[0849:cibcib]2.0.co;2}}
* {{cite journal | last1 = Jones | first1 = SE | year = 1977 | title = Coexistence in Mixed Species Antwren Flocks | journal = Oikos | volume = 29 | issue = 2| pages = 366–375 | doi=10.2307/3543628| jstor = 3543628 }}
* {{cite journal | last1 = Jones | first1 = SE | year = 1977 | title = Coexistence in Mixed Species Antwren Flocks | journal = Oikos | volume = 29 | issue = 2| pages = 366–375 | doi=10.2307/3543628| jstor = 3543628 | bibcode = 1977Oikos..29..366J }}
* {{cite journal | last1 = Lacki | first1 = MJ | year = 2000 | title = Surveys of bird communities on Little Black and Black mountains: Implications for long-term conservation of Montane birds in Kentucky | journal = Journal of the Kentucky Academy of Science | volume = 61 | issue = 1| pages = 50–59 }}
* {{cite journal | last1 = Lacki | first1 = MJ | year = 2000 | title = Surveys of bird communities on Little Black and Black mountains: Implications for long-term conservation of Montane birds in Kentucky | journal = Journal of the Kentucky Academy of Science | volume = 61 | issue = 1| pages = 50–59 }}
* {{cite journal | last1 = Lebbin | first1 = DJ | year = 2004 | title = Unusual June record of Canada Warbler (Wilsonia canadensis) in Bolivar, Venezuela | journal = Ornitologia Neotropical | volume = 15 | issue = 1| pages = 143–144 }}
* {{cite journal | last1 = Lebbin | first1 = DJ | year = 2004 | title = Unusual June record of Canada Warbler (Wilsonia canadensis) in Bolivar, Venezuela | journal = Ornitologia Neotropical | volume = 15 | issue = 1| pages = 143–144 }}
* {{cite journal | last1 = Merrill | first1 = SB | last2 = Cuthbert | first2 = FJ | last3 = Oehlert | first3 = G | year = 1998 | title = Residual patches and their contribution to forest-bird diversity on northern Minnesota aspen clearcuts | journal = Conservation Biology | volume = 12 | issue = 1| pages = 190–199 | doi=10.1111/j.1523-1739.1998.96067.x}}
* {{cite journal | last1 = Merrill | first1 = SB | last2 = Cuthbert | first2 = FJ | last3 = Oehlert | first3 = G | year = 1998 | title = Residual patches and their contribution to forest-bird diversity on northern Minnesota aspen clearcuts | journal = Conservation Biology | volume = 12 | issue = 1| pages = 190–199 | doi=10.1111/j.1523-1739.1998.96067.x| bibcode = 1998ConBi..12..190M | s2cid = 84727543 }}
* {{cite journal | last1 = Mitchell | first1 = JM | year = 1999 | title = Habitat relationships of five northern bird species breeding in hemlock ravines in Ohio, USA | journal = Natural Areas Journal | volume = 19 | issue = 1| pages = 3–11 }}
* {{cite journal | last1 = Mitchell | first1 = JM | year = 1999 | title = Habitat relationships of five northern bird species breeding in hemlock ravines in Ohio, USA | journal = Natural Areas Journal | volume = 19 | issue = 1| pages = 3–11 }}
* {{cite journal | last1 = Morris | first1 = SR | last2 = Richmond | first2 = ME | last3 = Holmes | first3 = DW | year = 1994 | title = Patterns of stopover by warblers during spring and fall migration on Appledore Island, Maine | journal = Wilson Bulletin | volume = 106 | issue = 4| pages = 703–718 }}
* {{cite journal | last1 = Morris | first1 = SR | last2 = Richmond | first2 = ME | last3 = Holmes | first3 = DW | year = 1994 | title = Patterns of stopover by warblers during spring and fall migration on Appledore Island, Maine | journal = Wilson Bulletin | volume = 106 | issue = 4| pages = 703–718 }}
Line 103: Line 103:
* {{cite journal | last1 = Rappole | first1 = JH | year = 1983 | title = Analysis of Plumage Variation in the Canada Warbler | journal = Journal of Field Ornithology | volume = 54 | issue = 2| pages = 152–159 }}
* {{cite journal | last1 = Rappole | first1 = JH | year = 1983 | title = Analysis of Plumage Variation in the Canada Warbler | journal = Journal of Field Ornithology | volume = 54 | issue = 2| pages = 152–159 }}
* {{cite journal | last1 = Robinson | first1 = SK | last2 = Fitzpatrick | first2 = JW | last3 = Terborgh | first3 = J | year = 1995 | title = Distribution and habitat use of neotropical migrant landbirds in the Amazon basin and Andes | journal = Bird Conservation International | volume = 5 | issue = 2–3| pages = 305–323 | doi=10.1017/s0959270900001064| doi-access = free }}
* {{cite journal | last1 = Robinson | first1 = SK | last2 = Fitzpatrick | first2 = JW | last3 = Terborgh | first3 = J | year = 1995 | title = Distribution and habitat use of neotropical migrant landbirds in the Amazon basin and Andes | journal = Bird Conservation International | volume = 5 | issue = 2–3| pages = 305–323 | doi=10.1017/s0959270900001064| doi-access = free }}
* {{cite journal | last1 = Sabo | first1 = SR | last2 = Whittaker | first2 = RH | year = 1979 | title = Bird Niches in a Subalpine Forest an Indirect Ordination | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 76 | issue = 3| pages = 1338–1342 | doi=10.1073/pnas.76.3.1338| pmid = 16592631 | pmc = 383246 | bibcode = 1979PNAS...76.1338S }}
* {{cite journal | last1 = Sabo | first1 = SR | last2 = Whittaker | first2 = RH | year = 1979 | title = Bird Niches in a Subalpine Forest an Indirect Ordination | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 76 | issue = 3| pages = 1338–1342 | doi=10.1073/pnas.76.3.1338| pmid = 16592631 | pmc = 383246 | bibcode = 1979PNAS...76.1338S | doi-access = free }}
* {{cite journal | last1 = Skinner | first1 = C | year = 2003 | title = A breeding bird survey of the natural areas at Holden Arboretum | journal = Ohio Journal of Science | volume = 103 | issue = 4| pages = 98–110 }}
* {{cite journal | last1 = Skinner | first1 = C | year = 2003 | title = A breeding bird survey of the natural areas at Holden Arboretum | journal = Ohio Journal of Science | volume = 103 | issue = 4| pages = 98–110 }}
* {{cite journal | last1 = Sodhi | first1 = NS | last2 = Paszkowski | first2 = CA | year = 1995 | title = Habitat use and foraging behavior of four parulid warblers in a second-growth forest | journal = Journal of Field Ornithology | volume = 66 | issue = 2| pages = 277–288 }}
* {{cite journal | last1 = Sodhi | first1 = NS | last2 = Paszkowski | first2 = CA | year = 1995 | title = Habitat use and foraging behavior of four parulid warblers in a second-growth forest | journal = Journal of Field Ornithology | volume = 66 | issue = 2| pages = 277–288 }}
* {{cite journal | last1 = Weakland | first1 = CA | last2 = Wood | first2 = PB | last3 = Ford | first3 = WM | year = 2002 | title = Responses of songbirds to diameter-limit cutting in the central Appalachians of West Virginia, USA | journal = Forest Ecology and Management | volume = 155 | issue = 1–3| pages = 115–129 | doi=10.1016/s0378-1127(01)00552-7}}
* {{cite journal | last1 = Weakland | first1 = CA | last2 = Wood | first2 = PB | last3 = Ford | first3 = WM | year = 2002 | title = Responses of songbirds to diameter-limit cutting in the central Appalachians of West Virginia, USA | journal = Forest Ecology and Management | volume = 155 | issue = 1–3| pages = 115–129 | doi=10.1016/s0378-1127(01)00552-7| bibcode = 2002ForEM.155..115W }}


==External links==
==External links==
Line 117: Line 117:


{{Taxonbar |from=Q27076004}}
{{Taxonbar |from=Q27076004}}
{{Authority control}}


{{DEFAULTSORT:warbler, Canada}}
[[Category:Cardellina|Canada warbler]]
[[Category:Cardellina|Canada warbler]]
[[Category:Birds of Canada]]
[[Category:Birds of Canada]]

Latest revision as of 22:18, 6 October 2024

Canada warbler
Male
Female showing characteristic lighter neck markings and grey top plumage
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Family: Parulidae
Genus: Cardellina
Species:
C. canadensis
Binomial name
Cardellina canadensis
(Linnaeus, 1766)
Range of C. canadensis
  Breeding range
  Wintering range
Synonyms
  • Muscicapa canadensis Linnaeus, 1766
  • Wilsonia canadensis (Linnaeus, 1766)

The Canada warbler (Cardellina canadensis) is a small boreal songbird of the New World warbler family (Parulidae). It summers in Canada and northeastern United States and winters in northern South America.

Taxonomy

[edit]

In 1760 the French zoologist Mathurin Jacques Brisson included a description of the Canada warbler in his Ornithologie based on a specimen collected in Canada. He used the French name Le gobe-mouche cendré de Canada and the Latin name Muscicapa Canadensis Cinerea.[2] Although Brissn coined Latin names, these do not conform to the binomial system and are not recognised by the International Commission on Zoological Nomenclature.[3] When in 1766 the Swedish naturalist Carl Linnaeus updated his Systema Naturae for the twelfth edition, he added 240 species that had been previously described by Brisson.[3] One of these was the Canada warbler. Linnaeus included a brief description, coined the binomial name Muscicapa canadensis, and cited Brisson's work.[4] The species is now placed in the genus Cardellina that was introduced by the French naturalist Charles Lucien Bonaparte in 1850.[5] The species is monotypic.[6]

Description

[edit]

The Canada warbler is sometimes called the "necklaced warbler", because of the band of dark streaks across its chest. The adults have minimal sexual dimorphism, although the male's "necklace" is darker and more conspicuous and also has a longer tail. Adults are 12–15 cm (4.7–5.9 in) long, have a wingspan of 17–22 cm (6.7–8.7 in), and weigh 9–13 g (0.32–0.46 oz).[7]

The chest, throat, and belly of the bird are yellow, and its back is dark grey. It has no wingbars or tail spots, but the underside of the tail is white. It has a yellow line in front of its eye in the direction of the beak, but the most striking facial feature is the white eyerings or "spectacles".[8] Immature specimens have similar coloration as adults but duller and with less pronounced facial features.[8]

Song

[edit]

The song of this bird is loud and highly variable, resembling chip chewy sweet dichetty. Their calls are low chup's.

A 2013 study showed that male Canada warblers have two performance-encoded song types. Mode I, used mostly during the day, when unpaired either alone or near a female during early nesting, involves stereotyped songs sung slowly and regularly. Mode II, used at dawn, after pairing and when near another male, involves variable songs, sung rapidly with irregular rhythm and chirping between songs. Most of the phrases used were common to both modes, a feature unique among parulids, which ordinarily have an individual's repertoire separated into two distinct parts.[9]

In 2000, a female Canada warbler (or a post-hatching year-old male that failed to moult, something never before observed) in Giles County, Virginia was observed singing. Its repertoire consisted of a repeated song of 12 to 13 notes as well as several shorter songs consisting of the first five or six notes of the longer song. The bird did not respond to the playback of its own song or a recording of a male. Although female singing among the parulids has long been considered "idiosyncratic", singing by female Canada warblers is supported by the observation of female singing in congener Wilson's warbler and the closely related hooded warbler.[10]

Distribution and habitat

[edit]

During the breeding season 82% of the population can be found in Canada and 18% in the United States.[7] In Canada, the summer range extends from southeastern Yukon to Nova Scotia. In the United States, the range extends from northern Minnesota to northern Pennsylvania, east to Long Island, New York. It also nests in the high Appalachians as far south as Georgia.[11] In winter, the Canada warbler's range extends from Guyana to northwestern Bolivia around the northern and western side of the Andean crest.[12]

In both summer and winter seasons, the Canada warbler inhabits moist thickets. During the breeding season, the bird "nests in riparian thickets, brushy ravines, forest bogs, etc. at a wide range of elevations and across a variety of forest types. In the northwestern parts of its range, it frequents aspen forests; in the center of the range, it is found in forested wetlands and swamps; and in the south, it occupies montane rhododendron thickets.[12]" In the winter, it prefers mid- and upper-elevation habitats.[12] In northern Minnesota, a study found that Canada warblers inhabited the shrub-forest edge, rather than mature forests or open fields with shrub.[13] In New England, the Canada warbler was found to be "disturbance specialists" moving into patches of forests recovering from wind throw or timber removal.[14] Because of its preference for low-height foraging in deciduous forests, it may be bounded at higher elevations as suitable habitat disappears and it suffers competition from the black-throated blue warbler which prefers similar habitats.[15]

Two accidentals have been observed in Europe. The first, a moribund male was caught in Sandgerði, Iceland on September 29, 1973. The second was a first winter, probably female observed for five days in October 2006 in County Clare, Ireland.[16]

Migration

[edit]

The Canada warbler is one of the last birds to arrive at the breeding grounds and one of the first to leave. They may spend only two months there. They fly at night along a route generally south and west to the Texas coast, then to southern Mexico. The arrive at the winter grounds in northwestern South America in late September to early October.[12]

Behavior and ecology

[edit]

Breeding

[edit]

At least 60–65% of the population nests in boreal forests in Canada, the Great Lakes region of the United States, New England and through the Appalachians.[12] The birds are at least seasonally monogamous. Sightings of pairs during migration in Panama have led to the conclusion that they are permanently monogamous.[17][7] This conclusion, however, is contradicted by the sexes' wintering at different elevations.[12]

Males arrive at the breeding grounds in the first two weeks of May.[11] Females build the nests on or very close to the ground in dense cover. The nests are made up of root masses, hummocks, stumps, stream banks, mossy logs, and sometimes leaf litter and grass clumps. Moss covering is frequent.[12]

The female lays four to five eggs and incubates for about 12 days. The chicks remain in the nest for about 10 days after hatching and are dependent on their parents for two to three weeks after they leave the nest.[18]

The age at which the young leave the nest is not known. Once independent they spend almost all their time in the understory, on the ground or in bushes.[11] The post-juvenile bird undergoes a partial moult involving all body feathers and wing coverlets. This may be completed before the first migration.[19]

The oldest known specimen was a male found in Quebec in 1982 at least 8 years old, having been banded in 1975.[20]

Food and feeding

[edit]

The Canada warbler eats insects for the most part, including beetles, mosquitoes, flies, moths, and smooth caterpillars such as cankerworms, supplemented by spiders, snails, worms, and, at least seasonally, fruit.[12][11] It employs several foraging tactics, such as flushing insects from foliage and catching them on the wing (which it does more frequently than other warblers),[12] and searching upon the ground among fallen leaves.[11] When they occasionally hover glean, males tend to fly higher than females on breeding grounds.[12] In the tropics of South America, it forages in mixed flocks with other birds, usually 3–30 feet above ground in denser foliage.[11]

Diseases and parasites

[edit]

In the summer of 1947 a single specimen of Canada warbler from Virginia (and one specimen of another warbler from Georgia) were found to be hosts of a new species of acanthocephalan worm, which was named Apororhynchus amphistomi, the third species of that genus and the first in North America.[21] In the southern part of the breeding range, nest parasitism by cowbirds is frequent.[12]

Status

[edit]

Partners in Flight estimates a global population of 4 million,[22] while the American Bird Conservancy estimates that 1.5 million individuals exist.[23]

Threats to the Canada warbler include forest fragmentation; over-browsing of the understory by deer, acid rain, and the spread of the woolly adelgid (a killer of fir and hemlock trees).[23] Owing to these factors the Breeding Bird Survey data show a population decline of 3.2 percent per year throughout the Canada warbler's breeding range, with the greatest declines in the Northeast.[24][23] The species has been assessed as "threatened" by the Committee on the Status of Endangered Wildlife in Canada.[25] The IUCN, however, ranks the Canada warbler as a species of least concern.[1]

The Canada warbler is protected at the federal level in both Canada and the United States.[18]

In art

[edit]

John James Audubon illustrates the Canada warbler in Birds of America (published, London 1827–38) as Plate 73 entitled "Bonaparte's Flycatching-Warbler—Muscicapa bonapartii." The single female (now properly identified as a Canada warbler) is shown perched on a great magnolia (Magnolia grandiflora) branch that was painted by Joseph Mason. The final, combined image was engraved and colored by Robert Havell Jr. at the Havell workshops in London. The original painting was purchased by the New York Historical Society.

References

[edit]
  1. ^ a b BirdLife International. (2016). "Cardellina canadensis". IUCN Red List of Threatened Species. 2016: e.T22721882A94737489. doi:10.2305/IUCN.UK.2016-3.RLTS.T22721882A94737489.en. Retrieved 9 April 2021.
  2. ^ Brisson, Mathurin Jacques (1760). Ornithologie, ou, Méthode contenant la division des oiseaux en ordres, sections, genres, especes & leurs variétés (in French and Latin). Vol. 2. Paris: Jean-Baptiste Bauche. pp. 406–408, Plate 39 fig 4. The two stars (**) at the start of the section indicates that Brisson based his description on the examination of a specimen.
  3. ^ a b Allen, J.A. (1910). "Collation of Brisson's genera of birds with those of Linnaeus". Bulletin of the American Museum of Natural History. 28: 317–335. hdl:2246/678.
  4. ^ Linnaeus, Carl (1766). Systema naturae : per regna tria natura, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1, Part 1 (12th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 327.
  5. ^ Bonaparte, Charles Lucien (1850). Conspectus generum avium (in Latin). Vol. 1. Lugduni Batavorum [Leiden]: E.J. Brill. p. 312.
  6. ^ Gill, Frank; Donsker, David, eds. (2018). "New World warblers, mitrospingid tanagers". World Bird List Version 8.1. International Ornithologists' Union. Retrieved 1 May 2018.
  7. ^ a b c "Canada Warbler—Life History". Cornell Lab of Ornithology. Retrieved December 15, 2016.
  8. ^ a b "Canada Warbler—Identification". Cornell Lab of Ornithology. Retrieved December 15, 2016.
  9. ^ Demko, Alana D.; Reitsma, Leonard R.; Staicer, Cynthia A. (October 2013). "Two song categories in the Canada Warbler (Cardellina canadensis)". The Auk. 130 (4): 609–16. doi:10.1525/auk.2013.13059. JSTOR 10.1525/auk.2013.13059. S2CID 83993091. Retrieved 28 April 2019.
  10. ^ Etterson, Matthew A. (2003). "An Observation of Singing by a Female-Plumaged Canada Warbler". Southeastern Naturalist. 2 (2): 419–22. doi:10.1656/1528-7092(2003)002[0419:aoosba]2.0.co;2. JSTOR 3878011. S2CID 86052159.
  11. ^ a b c d e f "Canada Warbler". Audubon Guide to North American Birds. 13 November 2014. Retrieved December 15, 2016.
  12. ^ a b c d e f g h i j k "Canada warbler". Boreal Songbird Initiative. 25 February 2014. Retrieved December 15, 2016.
  13. ^ Collins, Scott L.; James, Frances C.; Risser, Paul G. (1982). "Habitat Relationships of Wood Warblers Parulidae in Northern Central Minnesota". Oikos. 39 (1): 50–58. Bibcode:1982Oikos..39...50C. doi:10.2307/3544530. JSTOR 3544530. Retrieved December 18, 2016.
  14. ^ Chace, Jameson F.; Faccio, Steven D.; Chacko, Abraham (2009). "Canada Warbler Habitat Use of Northern Hardwoods in Vermont". Northeastern Naturalist. 16 (4): 491–500. doi:10.1656/045.016.n401. JSTOR 27744588. S2CID 86430663.
  15. ^ Sabo, Stephen R. (June 1980). "Niche and Habitat Relations in Subalpine Bird Communities of the White Mountains of New Hampshire". Ecological Monographs. 50 (2): 241–59. Bibcode:1980EcoM...50..241S. doi:10.2307/1942481. JSTOR 1942481.
  16. ^ Hanafin, Maurice (2006). "The Canada Warbler in County Clare—The Second for Western Palearctic" (PDF). Birding World. 19 (10): 429–34. Retrieved December 17, 2016.
  17. ^ Greenberg, Russell S.; Gradwohl, Judy A. (October 1980). "Observations of Paired Canada Warblers Wilsonia canadensis during Migration in Panama". Ibis. 122 (4): 509–12. doi:10.1111/j.1474-919X.1980.tb00907.x.
  18. ^ a b "Species Profile: Canada Warbler". Species at Risk Public Registry. Government of Canada. May 29, 2014. Archived from the original on September 9, 2014. Retrieved June 5, 2014.
  19. ^ Hanafin 2006, p. 434.
  20. ^ USGS Patuxent Wildlife Research Center (2016). "Longevity Records of North American Birds". U.S. Geological Survey. Department of the Interior. Retrieved December 16, 2016.
  21. ^ Byrd, Elon E.; Denton, Fred (August 1949). "The Helminth Parasites of Birds. II. A New Species of Acanthocephala from North American Birds". The Journal of Parasitology. 35 (4): 391–410. doi:10.2307/3273430. JSTOR 3273430. PMID 18133320.
  22. ^ "Species Assessment Database". Partners in Flight. Archived from the original on August 21, 2016. Retrieved December 15, 2016.
  23. ^ a b c "Canada Warbler". American Bird Conservancy. Retrieved December 15, 2016.
  24. ^ USGS Paxtuxent Wildlife Research Center (2016). "Canada Warbler, Cardellina canadensis: North American Breeding Bird Survey Trend Results". U.S. Geological Survey. Department of Interior. Retrieved December 16, 2016.
  25. ^ "Warbler, Canada". Committee on the Status of Endangered Wildlife in Canada (COSEWIC). Government of Canada. November 11, 2011. Archived from the original on 2014-06-06. Retrieved June 5, 2014.

General sources

[edit]

Further reading

[edit]
  • Barrowclough, GF; Corbin, KW (1978). "Genetic Variation and Differentiation in the Parulidae". Auk. 95 (4): 691–702.
  • Caroline, G; Marcel, D; Jean-Pierre, LS; Jean, H (2004). "Are temperate mixedwood forests perceived by birds as a distinct forest type?". Canadian Journal of Forest Research. 34 (9): 1895. doi:10.1139/x04-087.
  • Christian, DP; Hanowski, JM; Reuvers-House, M; Niemi, GJ; Blake, JG; Berguson, WE (1996). "Effects of mechanical strip thinning of aspen on small mammals and breeding birds in northern Minnesota, U.S.A". Canadian Journal of Forest Research. 26 (7): 1284–1294. doi:10.1139/x26-143.
  • Crawford, HS; Jennings, DT (1989). "Predation by Birds on Spruce Budworm Choristoneura-Fumiferana Functional Numerical and Total Responses". Ecology. 70 (1): 152–163. Bibcode:1989Ecol...70..152C. doi:10.2307/1938422. JSTOR 1938422.
  • Dunn, EH; Nol, E (1980). "Age Related Migratory Behavior of Warblers". Journal of Field Ornithology. 51 (3): 254–269.
  • Golet, FC; Wang, Y; Merrow, JS; DeRagon, WR (2001). "Relationship between habitat and landscape features and the avian community of red maple swamps in southern Rhode Island". Wilson Bulletin. 113 (2): 217–227. doi:10.1676/0043-5643(2001)113[0217:rbhalf]2.0.co;2. S2CID 86169562.
  • Hobson, KA; Bayne, E (2000). "The effects of stand age on avian communities in aspen-dominated forests of central Saskatchewan, Canada". Forest Ecology & Management. 136 (1–3): 121–134. Bibcode:2000ForEM.136..121H. CiteSeerX 10.1.1.532.1080. doi:10.1016/s0378-1127(99)00287-x.
  • Hobson, KA; Schieck, J (1999). "Changes in bird communities in boreal mixedwood forest: Harvest and wildfire effects over 30 years". Ecological Applications. 9 (3): 849–863. doi:10.1890/1051-0761(1999)009[0849:cibcib]2.0.co;2. S2CID 28285190.
  • Jones, SE (1977). "Coexistence in Mixed Species Antwren Flocks". Oikos. 29 (2): 366–375. Bibcode:1977Oikos..29..366J. doi:10.2307/3543628. JSTOR 3543628.
  • Lacki, MJ (2000). "Surveys of bird communities on Little Black and Black mountains: Implications for long-term conservation of Montane birds in Kentucky". Journal of the Kentucky Academy of Science. 61 (1): 50–59.
  • Lebbin, DJ (2004). "Unusual June record of Canada Warbler (Wilsonia canadensis) in Bolivar, Venezuela". Ornitologia Neotropical. 15 (1): 143–144.
  • Merrill, SB; Cuthbert, FJ; Oehlert, G (1998). "Residual patches and their contribution to forest-bird diversity on northern Minnesota aspen clearcuts". Conservation Biology. 12 (1): 190–199. Bibcode:1998ConBi..12..190M. doi:10.1111/j.1523-1739.1998.96067.x. S2CID 84727543.
  • Mitchell, JM (1999). "Habitat relationships of five northern bird species breeding in hemlock ravines in Ohio, USA". Natural Areas Journal. 19 (1): 3–11.
  • Morris, SR; Richmond, ME; Holmes, DW (1994). "Patterns of stopover by warblers during spring and fall migration on Appledore Island, Maine". Wilson Bulletin. 106 (4): 703–718.
  • Morse, DH (1977). "The Occupation of Small Islands by Passerine Birds". Condor. 79 (4): 399–412. doi:10.2307/1367719. JSTOR 1367719.
  • Patten, MA; Burger, JC (1998). "Spruce budworm outbreaks and the incidence of vagrancy in eastern North American wood-warblers". Canadian Journal of Zoology. 76 (3): 433–439. doi:10.1139/z97-213.
  • Prins, TG; Debrot, AO (1996). "First record of the Canada Warbler for Bonaire, Netherlands Antilles". Caribbean Journal of Science. 32 (2): 248–249.
  • Rappole, JH (1983). "Analysis of Plumage Variation in the Canada Warbler". Journal of Field Ornithology. 54 (2): 152–159.
  • Robinson, SK; Fitzpatrick, JW; Terborgh, J (1995). "Distribution and habitat use of neotropical migrant landbirds in the Amazon basin and Andes". Bird Conservation International. 5 (2–3): 305–323. doi:10.1017/s0959270900001064.
  • Sabo, SR; Whittaker, RH (1979). "Bird Niches in a Subalpine Forest an Indirect Ordination". Proceedings of the National Academy of Sciences of the United States of America. 76 (3): 1338–1342. Bibcode:1979PNAS...76.1338S. doi:10.1073/pnas.76.3.1338. PMC 383246. PMID 16592631.
  • Skinner, C (2003). "A breeding bird survey of the natural areas at Holden Arboretum". Ohio Journal of Science. 103 (4): 98–110.
  • Sodhi, NS; Paszkowski, CA (1995). "Habitat use and foraging behavior of four parulid warblers in a second-growth forest". Journal of Field Ornithology. 66 (2): 277–288.
  • Weakland, CA; Wood, PB; Ford, WM (2002). "Responses of songbirds to diameter-limit cutting in the central Appalachians of West Virginia, USA". Forest Ecology and Management. 155 (1–3): 115–129. Bibcode:2002ForEM.155..115W. doi:10.1016/s0378-1127(01)00552-7.
[edit]