Bewick's wren: Difference between revisions

Bewick's wren
	Song Call
Conservation status
	; Least Concern (IUCN 3.1)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Passeriformes
Family:	Troglodytidae
Genus:	Thryomanes; P.L. Sclater, 1862
Species:	T. bewickii
Binomial name
	Thryomanes bewickii; (Audubon, 1827)
Subspecies
	1–2 dozen living, 2 recently extinct; see article text
	Breeding Year-round Nonbreeding
Synonyms
	Thryomanes leucophrys; Thryothorus bewickii; Thryothorus brevicauda; Thryothorus brevicaudus; Thryothorus bairdi;

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The Bewick's wren (Thryomanes bewickii) is a wren native to North America. It is the only species placed in the genus Thryomanes. At about 14 cm (5.5 in) long, it is grey-brown above, white below, with a long white eyebrow. While similar in appearance to the Carolina wren, it has a long tail that is tipped in white. The song is loud and melodious, much like the song of other wrens. It lives in thickets, brush piles and hedgerows, open woodlands and scrubby areas, often near streams. It eats insects and spiders, which it gleans from vegetation or finds on the ground.^[2]

Its historic range was from southern British Columbia, Nebraska, southern Ontario, and southwestern Pennsylvania, Maryland, south to Mexico, Arkansas and the northern Gulf States. However, it is now extremely rare east of the Mississippi River.^[3]

Taxonomy

In 1827, the American ornithologist John James Audubon included an illustration of Bewick's wren under the binomial name Troglodytes bewickii in his The Birds of America.^[4] In the companion Ornithological Biography, published four years later, Audubon explained that he had shot the specimen near St. Francisville, Louisiana in 1821 and had chosen the specific epithet bewickii in honour of his friend the engraver Thomas Bewick.^[5] Bewick's wren is now the only species placed in the genus Thryomanes that was introduced by the English zoologist Philip Sclater in 1862.^[6]^[7]

The Socorro wren was formerly also placed in Thryomanes, but is now known to be a close relative of the house wren complex, as indicated by biogeography and mtDNA NADH dehydrogenase subunit 2 sequence analysis, whereas Thryomanes seems not too distant from the Carolina wren.^[8]

Subspecies

Fifteen subspecies are recognised of which two are now extinct.^[7] Subspecies can be distinguished by the small differences in the color of the dorsal feathers but this can be difficult for museum specimens as the colors tend to change after a few years of storage.^[9]

T. b. calophonus Oberholser, 1898 – southwest Canada and northwest USA
T. b. drymoecus Oberholser, 1898 – west Canada. Includes T. b. atrestus.
T. b. marinensis Grinnell, 1910 – coastal northwest California
T. b. spilurus (Vigors, 1839) – coastal central California
† T. b. leucophrys (Anthony, 1895) – Extinct, formerly San Clemente Island, California
T. b. charienturus Oberholser, 1898 – southern California and northwest Baja California
T. b. cerroensis (Anthony, 1897) – west central Baja California
T. b. magdalenensis Huey, 1942 – southwest Baja California
† T. b. brevicauda Ridgway, 1876 – Extinct, formerly Guadalupe Island, Mexico
T. b. eremophilus Oberholser, 1898 – interior southwest USA to central Mexico
T. b. cryptus Oberholser, 1898 – west Kansas, west Oklahoma and central, east Texas and northeast Mexico
T. b. pulichi (Phillips, AR, 1986) – east Kansas and Oklahoma
T. b. sadai (Phillips, AR, 1986) – south Texas (southern USA) to central Tamaulipas (northeast Mexico)
T. b. mexicanus (Deppe, 1830) – central and south Mexico. Includes T. b. murinus.
T. b. bewickii (Audubon, 1827) – Nominate subspecies, central and east central USA. Includes T. b. altus.

Description

The Bewick's wren has an average length of 5.1 inches (13 cm) an average weight of 0.3 to 0.4 ounces (8–12 g), and a wingspan of 18 cm.^[10] Its plumage is brown on top and light grey underneath, with a white stripe above each eye. Its beak is long, slender, and slightly curved.^[2] Its most distinctive feature is its long tail with black bars and white corners. It moves its tail around frequently, making this feature even more obvious for observers.^[11]

Juveniles look similar to adults, with only a few key differences. Their beaks are usually shorter and stockier. In addition, their underbelly might feature some faint speckling.^[2] Males and females are very similar in appearance.^[2]

Vocalizations

Bewick's wrens, like many wrens, are very vocal. Both females and males make short calls while foraging and both use a harsh scolding call when agitated.^[2] Males also sing in order to attract mates and protect their territory.^[2] The song is broken into two or three individual parts; one individual male may exhibit up to twenty-two different variations on the song pattern, and may even throw in a little ventriloquism to vary it even further.^[12] A male wren learns its song from neighboring males, so its song will be different from its father's.^[2]

Geographic variation

Geographic differences have been observed in the appearance of the Bewick's wren. Eastern populations, prior to their decline, were described as being more colorful, such as having a reddish tint to its brown feathers. Pacific populations are described as being darker in appearance, while populations in the Southwest are described as having a grayer plumage.^[11]

Geographic differences have also been noted in the song of Bewick's wrens. Each regional population of Bewick's wrens have distinctive vocalizations, in particular their call notes. Pacific populations sing notably more complicated songs than Southwestern populations. Eastern populations were also noted to be excellent singers.^[11]

Distribution and habitat

The Bewick's wren once had a range that extended throughout much of the United States and Mexico and parts of Canada. It used to be fairly common in the Midwest and in the Appalachian Mountains, but it is now extremely rare east of the Mississippi River. It is still found along the Pacific Coast from Baja California to British Columbia, in Mexico, and in a significant portion of the Southwest, including Texas, Arizona, New Mexico, and Oklahoma.^[3] Western populations do not tend to migrate. Eastern populations, prior to their decline, used to migrate from its northern range to the Gulf Coast.^[2]

The preferred habitat of the Bewick's wren is that of arid open woodlands and brush-filled areas such as hillsides and uplands, but will reside in humid areas locally (Subtropical and Temperate zones).^[13] They are more common than house wrens in drier habitats, such as those found in the Southwest.^[3] In California, Bewick's wrens inhabit a shrubland area called chaparral.^[14]

Behavior

Bewick's Wren feeding young and cleaning the nestbox

Feeding

Bewick's wrens are insect eaters. They glean insects and insect eggs from vegetation, including the trunks of trees. They typically do not feed on vegetation higher than 3 meters, but they will forage on the ground.^[3] Bewick's wrens are capable of hanging upside down in order to acquire food, such as catching an insect on the underside of a branch. When it catches an insect, it kills the insect prior to swallowing it whole. Bewick's wrens will repeatedly wipe their beaks on its perch after a meal.

Bewick's wrens will visit backyard feeders. They will eat suet, peanut hearts, hulled sunflower seeds, and mealworms.^[15] Like many insect-eating birds, the Bewick's wren widens its diet to include seeds in the winter.^[16]

Breeding

Courtship begins with the male singing from its perch. It will occasionally pause its song in order to chase its competitors. Bewick's wrens form monogamous pairs that will then forage together.^[2] The male wren begins building the nest in a cavity or birdhouse, with the female joining in later. The nest is constructed from twigs and other plant materials and is often lined with feathers. The nest is cup-shaped and located in a nook or cavity of some kind. It lays 5–7 eggs, which are white with brown spots. The Bewick's wren produces two broods in a season. Pairs are more or less monogamous when it comes to breeding, but go solitary throughout the winter.^[17]

Status and conservation

In 2016, the Bewick's wren was listed as least concern on the IUCN Red List of threatened species due to the size of its range and estimates of its population size.^[1] However, ornithologists have noted a severe decline in its eastern range and parts of its western range.^[3] In particular, it has virtually disappeared from east of the Mississippi. In 1984, the state of Maryland classified the Bewick's wren as endangered under its Maryland Endangered Species Act of 1971. Despite this classification, no breeding pairs of Bewick's wrens are known to remain in Maryland.^[18] In 2014, the North American Bird Conservation Initiative placed the eastern Bewick's wren on its watch list.^[19]

Several theories have been proposed to explain its decline in its eastern range, including pesticide use and competition from other bird species.^[3] The most likely reason seems to be competition from house wrens. House wrens compete with Bewick's wrens for similar nesting sites. House wrens will destroy both the nests and eggs of Bewick's wrens.^[2] The reforestation of once open land has also negatively impacted the eastern Bewick's wrens.^[2]

In California, habitat loss due to development has impacted the Bewick's wren. In San Diego, the development of canyons has led to the gradual decline of native bird species, including the Bewick's wren.^[14]

In Washington, development has actually benefited the Bewick's wren, leading to an increase in its population. However, this has coincided with the decline of the Pacific wren thanks to increased competition between the two species.^[20]

References

^ ^a ^b BirdLife International (2018). "Thryomanes bewickii". IUCN Red List of Threatened Species. 2018: e.T22711377A132096463. doi:10.2305/IUCN.UK.2018-2.RLTS.T22711377A132096463.en. Retrieved 12 November 2021.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k "Bewick's Wren". www.allaboutbirds.org. Retrieved 2017-05-02.
^ ^a ^b ^c ^d ^e ^f "Bewick's Wren - Introduction | Birds of North America Online". birdsna.org. Retrieved 2017-05-02.
^ Audubon, John James (1827). "Bewick's wren". The Birds of America; from original drawing. Vol. 1. London: Published by the author. Plate 18.
^ Audubon, John James (1831). Ornithological Biography, or an account of the habits of the birds of the United States of America ; accompanied by descriptions of the objects represented in the work entitled The Birds of America, and interspersed with delineations of American scenery and manners. Vol. 1. Edinburgh: Adam Black. pp. 96–97.
^ Sclater, P.L. (1862). Catalogue of a Collection of American Birds. London: N. Trubner and Co. p. 22.
^ ^a ^b Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2020). "Dapple-throats, sugarbirds, fairy-bluebirds, kinglets, hyliotas, wrens & gnatcatchers". IOC World Bird List Version 10.2. International Ornithologists' Union. Retrieved 10 October 2020.
^ Martínez Gómez; Juan E.; Barber, Bruian R. & Peterson, A. Townsend (2005). "Phylogenetic position and generic placement of the Socorro Wren (Thryomanes sissonii)" (PDF). Auk. 122 (1): 50–56. doi:10.1642/0004-8038(2005)122[0050:PPAGPO]2.0.CO;2. hdl:1808/16612. S2CID 20879561. Archived from the original (PDF) on 2008-12-17.
^ Kennedy, E.D.; White, D.W. (2020). Poole, A.F. (ed.). "Bewick's Wren (Thryomanes bewickii), version 1.0". Birds of the World. Ithaca, NY, USA: Cornell Lab of Ornithology. doi:10.2173/bow.bewwre.01. Retrieved 11 October 2020.
^ Oiseaux.net. "Troglodyte de Bewick - Thryomanes bewickii - Bewick's Wren". www.oiseaux.net. Retrieved 2020-09-28.
^ ^a ^b ^c Kaufman, Kenn (2006). "Bewick's Wren". Birder's World. 20: 60–61.
^ Beedy, Edward C.; Pandolfino, Edward R. (2013-06-17). Birds of the Sierra Nevada: Their Natural History, Status, and Distribution. University of California Press. ISBN 9780520274938.
^ The Committee on Classification and Nomenclature of the American Ornithologists' Union (1983). Check-List of North American Birds (sixth ed.). American Ornithologists' Union. p. 530. ISBN 0-943610-32-X.
^ ^a ^b Diamond, Jared (1988). "Urban extinction of birds". Nature. 333 (6172): 393–394. Bibcode:1988Natur.333..393D. doi:10.1038/333393a0. S2CID 4340734.
^ "Common Feeder Birds - FeederWatch". feederwatch.org. Retrieved 2017-05-02.
^ "Winter - Wild Birds Unlimited". Wild Birds Unlimited. Retrieved 2017-05-02.
^ Kennedy, E.D.; White, D.W. (1997). Poole, A.; Gill, F. (eds.). "Bewick's Wren (Thryomanes bewickii)". The Birds of North America (315). The Academy of Natural Sciences, Philadelphia, PA & The American Ornithologists' Union, Washington, D.C. doi:10.2173/bna.315.
^ "Wrens of Maryland - Maryland's Wild Acres". dnr2.maryland.gov. Retrieved 2017-03-26.
^ "2014 Report — The State of the Birds Report 2014". www.stateofthebirds.org. Retrieved 2017-03-26.
^ Farwell, Laura; Marzluff, John (2013). "A new bully on the block: Does urbanization promote Bewick's wren (Thryomanes bewickii) aggressive exclusion of Pacific wrens (Troglodytes pacificus)?". Biological Conservation. 161: 128–141. Bibcode:2013BCons.161..128F. doi:10.1016/j.biocon.2013.03.017.

External links

Bewick's wren - Thryomanes bewickii - USGS Patuxent Bird Identification InfoCenter
Bewick's wren Species Account – Cornell Lab of Ornithology
"Bewick's wren media". Internet Bird Collection.
Bewick's wren photo gallery at VIREO (Drexel University)
Interactive range map of Thryomanes bewickii at IUCN Red List maps
Audio recordings of Bewick's wren on Xeno-canto.

[iucn_status_12_November_2021-1] BirdLife International (2018). "Thryomanes bewickii". IUCN Red List of Threatened Species. 2018: e.T22711377A132096463. doi:10.2305/IUCN.UK.2018-2.RLTS.T22711377A132096463.en. Retrieved 12 November 2021.

[:0-2] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k "Bewick's Wren". www.allaboutbirds.org. Retrieved 2017-05-02.

[:2-3] ^ ^a ^b ^c ^d ^e ^f "Bewick's Wren - Introduction | Birds of North America Online". birdsna.org. Retrieved 2017-05-02.

[4] Audubon, John James (1827). "Bewick's wren". The Birds of America; from original drawing. Vol. 1. London: Published by the author. Plate 18.

[5] Audubon, John James (1831). Ornithological Biography, or an account of the habits of the birds of the United States of America ; accompanied by descriptions of the objects represented in the work entitled The Birds of America, and interspersed with delineations of American scenery and manners. Vol. 1. Edinburgh: Adam Black. pp. 96–97.

[6] Sclater, P.L. (1862). Catalogue of a Collection of American Birds. London: N. Trubner and Co. p. 22.

[ioc-7] Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (2020). "Dapple-throats, sugarbirds, fairy-bluebirds, kinglets, hyliotas, wrens & gnatcatchers". IOC World Bird List Version 10.2. International Ornithologists' Union. Retrieved 10 October 2020.

[Martinez-8] Martínez Gómez; Juan E.; Barber, Bruian R. & Peterson, A. Townsend (2005). "Phylogenetic position and generic placement of the Socorro Wren (Thryomanes sissonii)" (PDF). Auk. 122 (1): 50–56. doi:10.1642/0004-8038(2005)122[0050:PPAGPO]2.0.CO;2. hdl:1808/16612. S2CID 20879561. Archived from the original (PDF) on 2008-12-17.

[bow-9] Kennedy, E.D.; White, D.W. (2020). Poole, A.F. (ed.). "Bewick's Wren (Thryomanes bewickii), version 1.0". Birds of the World. Ithaca, NY, USA: Cornell Lab of Ornithology. doi:10.2173/bow.bewwre.01. Retrieved 11 October 2020.

[10] Oiseaux.net. "Troglodyte de Bewick - Thryomanes bewickii - Bewick's Wren". www.oiseaux.net. Retrieved 2020-09-28.

[:1-11] Kaufman, Kenn (2006). "Bewick's Wren". Birder's World. 20: 60–61.

[12] Beedy, Edward C.; Pandolfino, Edward R. (2013-06-17). Birds of the Sierra Nevada: Their Natural History, Status, and Distribution. University of California Press. ISBN 9780520274938.

[13] The Committee on Classification and Nomenclature of the American Ornithologists' Union (1983). Check-List of North American Birds (sixth ed.). American Ornithologists' Union. p. 530. ISBN 0-943610-32-X.

[:3-14] Diamond, Jared (1988). "Urban extinction of birds". Nature. 333 (6172): 393–394. Bibcode:1988Natur.333..393D. doi:10.1038/333393a0. S2CID 4340734.

[15] "Common Feeder Birds - FeederWatch". feederwatch.org. Retrieved 2017-05-02.

[16] "Winter - Wild Birds Unlimited". Wild Birds Unlimited. Retrieved 2017-05-02.

[kennedy&white1997-17] Kennedy, E.D.; White, D.W. (1997). Poole, A.; Gill, F. (eds.). "Bewick's Wren (Thryomanes bewickii)". The Birds of North America (315). The Academy of Natural Sciences, Philadelphia, PA & The American Ornithologists' Union, Washington, D.C. doi:10.2173/bna.315.

[18] "Wrens of Maryland - Maryland's Wild Acres". dnr2.maryland.gov. Retrieved 2017-03-26.

[19] "2014 Report — The State of the Birds Report 2014". www.stateofthebirds.org. Retrieved 2017-03-26.

[20] Farwell, Laura; Marzluff, John (2013). "A new bully on the block: Does urbanization promote Bewick's wren (Thryomanes bewickii) aggressive exclusion of Pacific wrens (Troglodytes pacificus)?". Biological Conservation. 161: 128–141. Bibcode:2013BCons.161..128F. doi:10.1016/j.biocon.2013.03.017.

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@@ Line 1: / Line 1: @@
+{{Short description|Species of bird}}
 {{Speciesbox
 | name = Bewick's wren
@@ Line 6: / Line 7: @@
 | status = LC
 | status_system = IUCN3.1
-| status_ref = <ref>{{IUCN|id=22711377 |title=''Thryomanes bewickii'' |assessor=BirdLife International |assessor-link=BirdLife International |version=2013.2 |year=2012 |accessdate=26 November 2013}}</ref>
+| status_ref = <ref name="iucn status 12 November 2021">{{cite iucn |author=BirdLife International |date=2018 |title=''Thryomanes bewickii'' |volume=2018 |page=e.T22711377A132096463 |doi=10.2305/IUCN.UK.2018-2.RLTS.T22711377A132096463.en |access-date=12 November 2021}}</ref>
 | genus = Thryomanes
 | parent_authority = [[Philip Lutley Sclater|P.L. Sclater]], 1862
 | species = bewickii
-| authority = ([[John James Audubon|Audubon]], 1829)
+| authority = ([[John James Audubon|Audubon]], 1827)
 | subdivision_ranks = Subspecies
 | subdivision =
@@ Line 20: / Line 21: @@
 *''Thryothorus bairdi''
 | range_map = Thryomanes bewickii map.svg
+| range_map_caption = {{leftlegend|#FF7F2A|Breeding}}{{leftlegend|#7137C8|Year-round}}{{leftlegend|#5F8DD3|Nonbreeding}}
 }}
-[[File:ThryothorusBairdiKeulemans.jpg|thumb|right|Subspecies ''T. b. bairdi''; illustration by [[John Gerrard Keulemans|Keulemans]], 1881]]
+[[File:18 Bewick's Wren.jpg|thumb|right|Illustration from [[John James Audubon|Audubon]]'s ''[[The Birds of America]]'']]
-The '''Bewick's wren''' (''Thryomanes bewickii'') is a [[wren]] native to North America. At about {{Convert|14|cm|in|abbr = on}} long, it is grey-brown above, white below, with a long white eyebrow. While similar in appearance to the [[Carolina wren]], it has a long tail that is tipped in white. The song is loud and melodious, much like the song of other wrens. It lives in thickets, brush piles and hedgerows, open woodlands and scrubby areas, often near streams. It eats insects and spiders, which it [[Gleaning (birds)|gleans]] from vegetation or finds on the ground.<ref name=":0" />
+The '''Bewick's wren''' ('''''Thryomanes bewickii''''') is a [[wren]] native to North America. It is the only species placed in the [[genus]] '''''Thryomanes'''''. At about {{Convert|14|cm|in|abbr = on}} long, it is grey-brown above, white below, with a long white eyebrow. While similar in appearance to the [[Carolina wren]], it has a long tail that is tipped in white. The song is loud and melodious, much like the song of other wrens. It lives in thickets, brush piles and hedgerows, open woodlands and scrubby areas, often near streams. It eats insects and spiders, which it [[Gleaning (birds)|gleans]] from vegetation or finds on the ground.<ref name=":0" />
 Its historic range was from southern [[British Columbia]], [[Nebraska]], southern [[Ontario]], and southwestern [[Pennsylvania]], [[Maryland]], south to [[Mexico]], [[Arkansas]] and the northern [[States of the Gulf Coast of the United States|Gulf States]]. However, it is now extremely rare east of the Mississippi River.<ref name=":2" />
 == Taxonomy ==
+In 1827, the American ornithologist [[John James Audubon]] included an illustration of Bewick's wren under the [[binomial name]] ''Troglodytes bewickii'' in his ''[[The Birds of America]]''.<ref>{{ cite book | last=Audubon | first=John James | author-link=John James Audubon | year=1827 | chapter=Bewick's wren | title=The Birds of America; from original drawing  | volume=1 | location=London | publisher=Published by the author | at=Plate 18 | chapter-url=https://digital.library.pitt.edu/islandora/object/pitt%3Aaud0018 }}</ref> In the companion ''Ornithological Biography'', published four years later, Audubon explained that he had shot the specimen near [[St. Francisville, Louisiana]] in 1821 and had chosen the specific epithet ''bewickii'' in honour of his friend the engraver [[Thomas Bewick]].<ref>{{ cite book | last=Audubon | first=John James | author-link=John James Audubon | year=1831 | title=Ornithological Biography, or an account of the habits of the birds of the United States of America ; accompanied by descriptions of the objects represented in the work entitled The Birds of America, and interspersed with delineations of American scenery and manners | volume=1 | location=Edinburgh | publisher=Adam Black | pages=96–97 | url=https://www.biodiversitylibrary.org/page/33238890 }}</ref> Bewick's wren is now the only species placed in the [[genus]] ''Thryomanes'' that was introduced by the English zoologist [[Philip Sclater]] in 1862.<ref>{{ cite book | last=Sclater | first=P.L. | author-link=Philip Sclater  | year=1862 | title=Catalogue of a Collection of American Birds | location=London | publisher=N. Trubner and Co. | page=22 | url=https://www.biodiversitylibrary.org/page/13302204}}</ref><ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela Rasmussen | year=2020 | title=Dapple-throats, sugarbirds, fairy-bluebirds, kinglets, hyliotas, wrens & gnatcatchers | work=IOC World Bird List Version 10.2 | url=http://www.worldbirdnames.org/bow/sugarbirds/ | publisher=International Ornithologists' Union | access-date=10 October 2020 }}</ref>
-[[John James Audubon]] is credited with having first described a specimen of the Bewick's wren, collecting the first known specimen in 1821. The Bewick's Wren is named after Audubon's friend [[Thomas Bewick]], an English engraver and natural historian.<ref name=":2">{{Cite web|url=https://birdsna.org/Species-Account/bna/species/315/articles/introduction?__hstc=75100365.e28ce7d114be476872047861c646d666.1487121674173.1490454734250.1490535196317.6&__hssc=75100365.1.1490535196317&__hsfp=514135907#_ga=1.251930624.800997852.1487121665|title=Bewick's Wren - Introduction {{!}} Birds of North America Online|website=birdsna.org|language=en|access-date=2017-05-02}}</ref>
-This is currently the [[Monotypic taxon|only species]] of its [[genus]], '''''Thryomanes'''''. The [[Socorro wren]], formerly placed here too, is actually a close relative of the [[house wren]] complex, as indicated by [[biogeography]] and [[mtDNA]] [[NADH dehydrogenase]] [[Protein subunit|subunit]] 2 [[DNA sequence|sequence]] analysis, whereas ''Thryomanes'' seems not too distant from the [[Carolina wren]].<ref name="Martinez" />
+The [[Socorro wren]] was formerly also placed in ''Thryomanes'', but is now known to be a close relative of the [[Northern house wren|house wren]] complex, as indicated by [[biogeography]] and [[mtDNA]] [[NADH dehydrogenase]] [[Protein subunit|subunit]] 2 [[DNA sequence|sequence]] analysis, whereas ''Thryomanes'' seems not too distant from the [[Carolina wren]].<ref name="Martinez" />
 ===Subspecies===
+Fifteen [[subspecies]] are recognised of which two are now extinct.<ref name=ioc/> Subspecies can be distinguished by the small differences in the color of the dorsal feathers but this can be difficult for museum specimens as the colors tend to change after a few years of storage.<ref name=bow>{{cite journal | last1=Kennedy | first1=E.D. | last2=White | first2=D.W. | year=2020 | title=Bewick's Wren (''Thryomanes bewickii''), version 1.0 | editor-last=Poole | editor-first=A.F. | journal=Birds of the World | location=Ithaca, NY, USA | publisher=Cornell Lab of Ornithology | doi=10.2173/bow.bewwre.01 | url=https://doi.org/10.2173/bow.bewwre.01 | access-date=11 October 2020 | url-access=subscription }}</ref>
-A list of commonly recognized subspecies follows. Two have gone [[extinct]] during the 20th century, mainly due to [[habitat destruction]] and [[Cat predation on wildlife|cat predation]].<ref name=kennedy&white1997/>
+* ''T. b. calophonus'' Oberholser, 1898 – southwest Canada and northwest USA
-* ''T. b. bewickii'' – <small>(Audubon, 1827)</small>: [[Nominate subspecies#Nominotypical subspecies and subspecies autonyms|nominate]], [[Midwest]]ern USA from NE [[Kansas]] to [[Missouri]] and E [[Texas]]. Includes ''T. b. pulichi'' as a [[junior synonym]].
+* ''T. b. drymoecus'' Oberholser, 1898 – west Canada. Includes ''T. b. atrestus''.
-* ''T. b. altus'' – <small>Aldrich, 1944</small>: Formerly in [[Appalachian Mountains|Appalachian]] region; S Ontario to [[South Carolina]], now quite rare. Possibly an [[Endangered species|endangered subspecies]], but possibly not distinct from ''bewickii''.
+* ''T. b. marinensis'' Grinnell, 1910 – coastal northwest California
-* ''T. b. cryptus'' – <small>Oberholser, 1898</small>: Central Kansas to N [[Tamaulipas]] in Mexico. Includes ''T. b. niceae''. Southeastern birds are sometimes separated as ''T. b. sadai''.
+* ''T. b. spilurus'' (Vigors, 1839) – coastal central California
-* ''T. b. eremophilus'' – <small>Oberholser, 1898</small>: E [[California]] inland, south to [[Zacatecas]] in Mexico.
+*† ''[[Thryomanes bewickii leucophrys|T. b. leucophrys]]'' (Anthony, 1895) – Extinct, formerly [[San Clemente Island]], California
-* ''T. b. calophonus'' – <small>Oberholser, 1898</small>: SW British Columbia, [[Canada]], to W [[Oregon]]. Includes ''T. b. ariborius'' and ''T. b. hurleyi''. The former name refers to the population found in the area of [[Seattle]] and [[Vancouver]]; these birds are sometimes called '''Seattle wren'''.
-* ''T. b. marinensis'' – <small>Grinnell, 1910</small>: Coastal NW California to [[Marin County]].
+* ''T. b. charienturus'' Oberholser, 1898 – southern California and northwest [[Baja California]]
+* ''T. b. cerroensis'' (Anthony, 1897) – west central Baja California
-* ''T. b. spilurus'' – <small>(Vigors, 1839)</small>: Coastal California from [[San Francisco Bay]] to [[Santa Cruz County, California|Santa Cruz County]].
-* ''T. b. drymoecus'' – <small>Oberholser, 1898</small>: SW Oregon to [[California Central Valley]].
+* ''T. b. magdalenensis'' Huey, 1942 – southwest Baja California
+*† ''[[Guadalupe wren|T. b. brevicauda]]'' Ridgway, 1876 – Extinct, formerly [[Guadalupe Island]], Mexico
-* ''T. b. atrestus'' – <small>Oberholser, 1932</small>: S Oregon to W [[Nevada]]. Probably not valid.
+* ''T. b. eremophilus'' Oberholser, 1898 – interior southwest USA to central Mexico
-* ''T. b. correctus'' <small>Grinnell</small>. SW coastal California to Mexican border; possibly synonym of ''charienturus''.
-* ''T. b. charienturus'' – <small>Oberholser, 1898</small>: N [[Baja California Peninsula]] to about 30°N.
+* ''T. b. cryptus'' Oberholser, 1898 – west Kansas, west Oklahoma and central, east Texas and northeast Mexico
+* ''T. b. pulichi'' (Phillips, AR, 1986) – east Kansas and Oklahoma
-* ''T. b. magdalenensis'' – <small>Huey, 1942</small>: SW Baja California Peninsula from 26 to 24°N.
+* ''T. b. sadai'' (Phillips, AR, 1986) – south Texas (southern USA) to central [[Tamaulipas]] (northeast Mexico)
-* ''T. b. nesophilus'' <small>Oberholser</small>. [[Santa Rosa Island, California|Santa Rosa]], [[Santa Cruz Island|Santa Cruz]], and [[Anacapa Island]]s, California; probably also [[Santa Barbara Island|Santa Barbara]] and [[San Nicolas Island|San Nicolas]]; found on the mainland in winter. Possibly synonym of ''charienturus''.
+* ''T. b. mexicanus'' (Deppe, 1830) – central and south Mexico. Includes ''T. b. murinus''.
-* ''T. b. catalinae'' – <small>Grinnell</small>: [[Santa Catalina Island, California]]; found on the mainland in winter. Possibly synonym of ''charienturus''.
-* ''T. b. cerroensis'' – <small>(Anthony, 1897)</small>: [[Cedros Island]] (Mexico) and W central Baja California. Includes ''T. b. atricauda''.
+* ''T. b. bewickii'' (Audubon, 1827) – [[Nominate subspecies]], central and east central USA. Includes ''T. b. altus''.
-* ''[[Thryomanes bewickii leucophrys|T. b. leucophrys]]'' [[extinction|†]] – <small>(Anthony, 1895)</small>: '''San Clemente Bewick's wren'''. Formerly [[San Clemente Island]], California.
-::[[Extinct]] since the 1940s due to [[habitat destruction]] by feral [[goat]]s and [[sheep]]. Also called ''T. b. anthonyi''. Observations of ''leucophrys'' in 1897<ref name=kaeding1905/> refer to ''cerroensis''; at that time, the San Clemente wren was considered a good species which included the Cedros population.
-* ''T. b. brevicauda'' [[extinction|†]] – <small>Ridgway, 1876</small>: '''Guadalupe Bewick's wren'''. Formerly [[Guadalupe Island]], Mexico.
-::This subspecies is [[extinct]] since (probably) the late 1890s due to habitat destruction by feral goats and predation by feral [[cat]]s. Overcollecting by scientists might have hastened its demise.<ref name = anthony1901/> It was last collected (3 specimens) by Anthony and Streator in May 1892<ref name=anthony1901/> and seen but found to be "nearly extinct" on March 22, 1897.<ref name = kaeding1905 /> It was not found by Anthony in several searches between 1892 and 1901 and considered certainly extinct by 1901;<ref name=anthony1901/> a thorough search in 1906 confirmed the subspecies' extinction.<ref name=Thayer/><ref>The often-reported extinction date of 1903 seems to be the first record of its absence rather than the last record of its presence{{Citation needed|date=October 2007}}<!-- secondary source is AOU checklist of 1957. Primary source is? Probably given in Grinnell 1928 -->. Actually, there appears to be no post-1897 record. The schedule of Anthony's visits after 1892 is not known; if he visited the island before 1897 he must have overlooked the last remnant of the population and thus his extinction date of 1901 may be called into question. By the balance of evidence, it is likely however that the subspecies became extinct between 1897 and 1901.</ref>
-* ''T. b. murinus'' – <small>(Hartlaub, 1852)</small>: Eastern and central Mexico.
-* ''T. b. bairdi'' – <small>(Salvin and Goodman)</small>: SE Mexico to S [[Puebla]].
-* ''T. b. percnus'' – <small>(Oberholser)</small>: [[Jalisco]] to [[Guerrero]], Mexico.
-The last three are sometimes united as ''T. b. mexicanus''. The validity of subspecies needs to be verified using freshly caught birds and/or molecular data, as specimens are prone to [[foxing]] quickly.<ref name=kennedy&white1997/>
 == Description ==
+The Bewick's wren has an average length of 5.1 inches (13&nbsp;cm) an average weight of 0.3 to 0.4 ounces (8–12 g), and a wingspan of 18&nbsp;cm.<ref>{{Cite web|last=Oiseaux.net|title=Troglodyte de Bewick - Thryomanes bewickii - Bewick's Wren|url=https://www.oiseaux.net/birds/bewick.s.wren.html|access-date=2020-09-28|website=www.oiseaux.net|language=en}}</ref> Its plumage is brown on top and light grey underneath, with a white stripe above each eye. Its beak is long, slender, and slightly curved.<ref name=":0">{{Cite web|url=https://www.allaboutbirds.org/guide/Bewicks_Wren/lifehistory|title=Bewick's Wren|website=www.allaboutbirds.org|access-date=2017-05-02}}</ref> Its most distinctive feature is its long tail with black bars and white corners. It moves its tail around frequently, making this feature even more obvious for observers.<ref name=":1">{{Cite journal|last=Kaufman|first=Kenn|date=2006|title=Bewick's Wren|journal=Birder's World|volume=20|pages=60–61}}</ref>
-[[File:BewicksWren-13JAN2017.jpg|alt=Bewick's Wren|left|thumb|326x326px|Bewick's Wren in [[Sacramento, California]].]]
-The Bewick's wren has an average length of 5.1 inches (13&nbsp;cm) and an average weight of 0.3 to 0.4 ounces (8 -12 g). Its plumage is brown on top and light grey underneath, with a white stripe above each eye. Its beak is long, slender, and slightly curved.<ref name=":0">{{Cite web|url=https://www.allaboutbirds.org/guide/Bewicks_Wren/lifehistory|title=Bewick's Wren|website=www.allaboutbirds.org|access-date=2017-05-02}}</ref> Its most distinctive feature is its long tail with black bars and white corners. It moves its tail around frequently, making this feature even more obvious for observers.<ref name=":1">{{Cite journal|last=Kaufman|first=Kenn|date=2006|title=Bewick's Wren|url=|journal=Birder's World|volume=20|pages=60–61}}</ref>
 Juveniles look similar to adults, with only a few key differences. Their beaks are usually shorter and stockier. In addition, their underbelly might feature some faint speckling.<ref name=":0" />  Males and females are very similar in appearance.<ref name=":0" />
 === Vocalizations ===
-Bewick's wrens, like many wrens, are very vocal. Both females and males make short calls while foraging and both use a harsh scolding call when agitated.<ref name=":0" />  Males also sing in order to attract mates and protect their territory.<ref name=":0" />  The song is broken into two or three individual parts; one individual male may exhibit up to twenty-two different variations on the song pattern, and may even throw in a little ventriloquism to vary it even further.<ref>https://books.google.com/books?id=oBYzqOAZHpcC&pg=PA260</ref> A male wren learns its song from neighboring males, so its song will be different from its father's.<ref name=":0" />
+Bewick's wrens, like many wrens, are very vocal. Both females and males make short calls while foraging and both use a harsh scolding call when agitated.<ref name=":0" />  Males also sing in order to attract mates and protect their territory.<ref name=":0" />  The song is broken into two or three individual parts; one individual male may exhibit up to twenty-two different variations on the song pattern, and may even throw in a little ventriloquism to vary it even further.<ref>{{Cite book | url=https://books.google.com/books?id=oBYzqOAZHpcC&pg=PA260 |title = Birds of the Sierra Nevada: Their Natural History, Status, and Distribution|isbn = 9780520274938|last1 = Beedy|first1 = Edward C.|last2 = Pandolfino|first2 = Edward R.|date = 2013-06-17| publisher=University of California Press }}</ref> A male wren learns its song from neighboring males, so its song will be different from its father's.<ref name=":0" />
 === Geographic variation ===
+[[File:BewicksWren-13JAN2017.jpg|alt=Bewick's Wren|left|thumb|Bewick's Wren in [[Sacramento, California]]]]
 Geographic differences have been observed in the appearance of the Bewick's wren. Eastern populations, prior to their decline, were described as being more colorful, such as having a reddish tint to its brown feathers. Pacific populations are described as being darker in appearance, while populations in the Southwest are described as having a grayer plumage.<ref name=":1" />
@@ Line 75: / Line 68: @@
 == Distribution and habitat ==
-The Bewick's wren once had a range that extended throughout much of the United States and Mexico and parts of Canada. It used to be fairly common in the Midwest and in the Appalachian Mountains, but it is now extremely rare east of the Mississippi River. It is still found along the Pacific Coast from [[Baja California Peninsula|Baja California]] to [[British Columbia]], in [[Mexico]], and in a significant portion of the [[Southwestern United States|Southwest]], including Texas, Arizona, New Mexico, and Oklahoma.<ref name=":2" /> Western populations do not tend to migrate. Eastern populations, prior to their decline, used to migrate from its northern range to the Gulf Coast.<ref name=":0" />
+The Bewick's wren once had a range that extended throughout much of the United States and Mexico and parts of Canada. It used to be fairly common in the Midwest and in the Appalachian Mountains, but it is now extremely rare east of the Mississippi River. It is still found along the Pacific Coast from [[Baja California Peninsula|Baja California]] to [[British Columbia]], in [[Mexico]], and in a significant portion of the [[Southwestern United States|Southwest]], including Texas, Arizona, New Mexico, and Oklahoma.<ref name=":2">{{Cite web|url=https://birdsna.org/Species-Account/bna/species/315/articles/introduction?__hstc=75100365.e28ce7d114be476872047861c646d666.1487121674173.1490454734250.1490535196317.6&__hssc=75100365.1.1490535196317&__hsfp=514135907#_ga=1.251930624.800997852.1487121665|title=Bewick's Wren - Introduction {{!}} Birds of North America Online|website=birdsna.org|language=en|access-date=2017-05-02}}</ref> Western populations do not tend to migrate. Eastern populations, prior to their decline, used to migrate from its northern range to the Gulf Coast.<ref name=":0" />
-The preferred habitat of the Bewick's wren is that of open woodlands and brush-filled areas such as hillsides and uplands. They are more common than house wrens in drier habitats, such as those found in the Southwest.<ref name=":2" />  In California, Bewick's wrens inhabit a [[shrubland]] area called [[chaparral]].<ref name=":3">{{Cite journal|last=Diamond|first=Jared|date=1988|title=Urban extinction of birds|url=|journal=Nature|volume=333|pages=393–394|doi=10.1038/333393a0}}</ref>
+The preferred habitat of the Bewick's wren is that of arid open woodlands and brush-filled areas such as hillsides and uplands, but will reside in humid areas locally (Subtropical and Temperate zones).<ref>{{cite book |author1=The Committee on Classification and Nomenclature of the American Ornithologists' Union |title=Check-List of North American Birds |date=1983 |publisher=American Ornithologists' Union |isbn=0-943610-32-X |page=530 |edition=sixth |language=en}}</ref> They are more common than house wrens in drier habitats, such as those found in the Southwest.<ref name=":2" /> In California, Bewick's wrens inhabit a [[shrubland]] area called [[chaparral]].<ref name=":3">{{Cite journal|last=Diamond|first=Jared|date=1988|title=Urban extinction of birds|journal=Nature|volume=333|issue=6172|pages=393–394|doi=10.1038/333393a0|bibcode=1988Natur.333..393D|s2cid=4340734|doi-access=free}}</ref>
 == Behavior ==
-[[File:Bewick's Wren nestbox feeding cleaning.webm|thumbtime=1.4|thumb|200px|alt=Video of a Bewick's Wren parent in a nestbox in Oakland, California, feeding her young an insect and cleaning out a fecal sac.|Bewick's Wren feeding young and cleaning the nestbox.]]
+[[File:Bewick's Wren nestbox feeding cleaning.webm|thumbtime=1.4|thumb|200px|alt=Video of a Bewick's Wren parent in a nestbox in Oakland, California, feeding her young an insect and cleaning out a fecal sac.|Bewick's Wren feeding young and cleaning the nestbox]]
 === Feeding ===
@@ Line 88: / Line 81: @@
 === Breeding ===
-Courtship begins with the male singing from its perch. It will occasionally pause its song in order to chase its competitors. Bewick's wrens form monogamous pairs that will then forage together.<ref name=":0" />  The male wren begins building the nest in a cavity or birdhouse, with the female joining in later. The nest is constructed from twigs and other plant materials and is often lined with feathers. The [[Bird nest|nest]] is cup-shaped and located in a nook or cavity of some kind. It lays 5–7 eggs that are white with brown spots. The Bewick's wren produces two broods in a season. Pairs are more or less monogamous when it comes to breeding, but go solitary throughout the winter.<ref name="kennedy&white1997" />
+Courtship begins with the male singing from its perch. It will occasionally pause its song in order to chase its competitors. Bewick's wrens form monogamous pairs that will then forage together.<ref name=":0" />  The male wren begins building the nest in a cavity or birdhouse, with the female joining in later. The nest is constructed from twigs and other plant materials and is often lined with feathers. The [[Bird nest|nest]] is cup-shaped and located in a nook or cavity of some kind. It lays 5–7 eggs, which are white with brown spots. The Bewick's wren produces two broods in a season. Pairs are more or less monogamous when it comes to breeding, but go solitary throughout the winter.<ref name="kennedy&white1997" />
 == Status and conservation ==
-In 2016, the Bewick's wren was listed as [[Least-concern species|least concern]] on the [[IUCN Red List]] of [[threatened species]] due to the size of its range and estimates of its population size.<ref>{{Cite web|url=http://www.iucnredlist.org/details/22711377/0|title=Thryomanes bewickii (Bewick's Wren)|website=www.iucnredlist.org|access-date=2017-03-26}}</ref> However, ornithologists have noted a severe decline in its eastern range and parts of its western range.<ref name=":2" /> In particular, it has virtually disappeared from east of the Mississippi. In 1984, the state of [[Maryland]] classified the Bewick's wren as [[Endangered species|endangered]] under its Maryland Endangered Species Act of 1971. Despite this classification, no breeding pairs of Bewick's wrens are known to remain in Maryland.<ref>{{Cite web|url=http://dnr2.maryland.gov/wildlife/Pages/habitat/wawren.aspx|title=Wrens of Maryland - Maryland's Wild Acres|website=dnr2.maryland.gov|language=en-us|access-date=2017-03-26}}</ref> In 2014, the North American Bird Conservation Initiative placed the eastern Bewick's wren on its watch list.<ref>{{Cite web|url=http://www.stateofthebirds.org/2014/?__hstc=75100365.e28ce7d114be476872047861c646d666.1487121674173.1490537359243.1490548559189.8&__hssc=75100365.2.1490548559189&__hsfp=514135907#_ga=1.248709518.800997852.1487121665|title=2014 Report         —         The State of the Birds Report 2014|website=www.stateofthebirds.org|language=en|access-date=2017-03-26}}</ref>
+In 2016, the Bewick's wren was listed as [[Least-concern species|least concern]] on the [[IUCN Red List]] of [[threatened species]] due to the size of its range and estimates of its population size.<ref name="iucn status 12 November 2021" /> However, ornithologists have noted a severe decline in its eastern range and parts of its western range.<ref name=":2" /> In particular, it has virtually disappeared from east of the Mississippi. In 1984, the state of [[Maryland]] classified the Bewick's wren as [[Endangered species|endangered]] under its Maryland Endangered Species Act of 1971. Despite this classification, no breeding pairs of Bewick's wrens are known to remain in Maryland.<ref>{{Cite web|url=http://dnr2.maryland.gov/wildlife/Pages/habitat/wawren.aspx|title=Wrens of Maryland - Maryland's Wild Acres|website=dnr2.maryland.gov|language=en-us|access-date=2017-03-26}}</ref> In 2014, the North American Bird Conservation Initiative placed the eastern Bewick's wren on its watch list.<ref>{{Cite web|url=http://www.stateofthebirds.org/2014/?__hstc=75100365.e28ce7d114be476872047861c646d666.1487121674173.1490537359243.1490548559189.8&__hssc=75100365.2.1490548559189&__hsfp=514135907#_ga=1.248709518.800997852.1487121665|title=2014 Report — The State of the Birds Report 2014|website=www.stateofthebirds.org|language=en|access-date=2017-03-26}}</ref>
-Several theories have been proposed to explain its decline in its eastern range, including pesticide use and competition from other bird species.<ref name=":2" /> The most likely reason seems to be competition from [[house wren]]s. House wrens compete with Bewick's wrens for similar nesting sites. House wrens will destroy both the nests and eggs of Bewick's wrens.<ref name=":0" /> The reforestation of once open land has also negatively impacted the eastern Bewick's wrens.<ref name=":0" />
+Several theories have been proposed to explain its decline in its eastern range, including pesticide use and competition from other bird species.<ref name=":2" /> The most likely reason seems to be competition from [[Northern house wren|house wren]]s. House wrens compete with Bewick's wrens for similar nesting sites. House wrens will destroy both the nests and eggs of Bewick's wrens.<ref name=":0" /> The reforestation of once open land has also negatively impacted the eastern Bewick's wrens.<ref name=":0" />
 In California, habitat loss due to development has impacted the Bewick's wren. In [[San Diego]], the development of canyons has led to the gradual decline of native bird species, including the Bewick's wren.<ref name=":3" />
-In Washington, development has actually benefited the Bewick's wren, leading to an increase in its population. However, this has coincided with the decline of the [[Pacific wren]] thanks to increased competition between the two species.<ref>{{Cite journal|last=Farwell|first=Laura|last2=Marzluff|first2=John|year=2013|title=A new bully on the block: Does urbanization promote Bewick's wren (Thryomanes bewickii) aggressive exclusion of Pacific wrens (Troglodytes pacificus)?|url=|journal=Biological Conservation|volume=161|pages=128–141|via=EBSCOhost|doi=10.1016/j.biocon.2013.03.017}}</ref>
+In Washington, development has actually benefited the Bewick's wren, leading to an increase in its population. However, this has coincided with the decline of the [[Pacific wren]] thanks to increased competition between the two species.<ref>{{Cite journal|last1=Farwell|first1=Laura|last2=Marzluff|first2=John|year=2013|title=A new bully on the block: Does urbanization promote Bewick's wren (Thryomanes bewickii) aggressive exclusion of Pacific wrens (Troglodytes pacificus)?|journal=Biological Conservation|volume=161|pages=128–141|doi=10.1016/j.biocon.2013.03.017|bibcode=2013BCons.161..128F }}</ref>
 == References ==
 {{Reflist|refs=
+<ref name=kennedy&white1997>{{cite journal |last1=Kennedy |first1=E.D. |last2=White |first2=D.W. |year=1997 |title=Bewick's Wren (''Thryomanes bewickii'') |editor-last1=Poole |editor-first1=A.  |editor-last2=Gill |editor-first2=F. |journal=The Birds of North America |number=315 |publisher=The Academy of Natural Sciences, Philadelphia, PA & The American Ornithologists' Union, Washington, D.C.|doi=10.2173/bna.315 }}</ref>
-<ref name=anthony1901>{{cite journal |last=Anthony |first=A.W. |year=1901 |title=The Guadalupe Wren |journal=[[Condor (journal)|Condor]] |volume=3 |issue=3 |pages=73 |url=http://sora.unm.edu/sites/default/files/journals/condor/v003n03/p0073-p0073.pdf |doi=10.2307/1361475}}</ref>
+<ref name=Martinez>{{cite journal|author1=Martínez Gómez|author2=Juan E.|author3=Barber, Bruian R.|author4=Peterson, A. Townsend|name-list-style=amp|year=2005|title=Phylogenetic position and generic placement of the Socorro Wren (''Thryomanes sissonii'')|journal=[[Auk (journal)|Auk]]|volume=122|issue=1|pages=50–56|doi=10.1642/0004-8038(2005)122[0050:PPAGPO]2.0.CO;2|url=http://www.specifysoftware.org/Informatics/bios/biostownpeterson/MBP_A_2005.pdf|url-status=dead|archive-url=https://web.archive.org/web/20081217114932/http://www.specifysoftware.org/Informatics/bios/biostownpeterson/MBP_A_2005.pdf|archive-date=2008-12-17|hdl=1808/16612|s2cid=20879561 |hdl-access=free}}</ref>
-<ref name=kaeding1905>{{cite journal |author=Kaeding, Henry B. |year=1905| title= Birds from the West Coast of Lower California and Adjacent Islands (Part II) |journal=[[Condor (journal)|Condor]] |volume=7 |issue=4 |pages=134–138 |url=http://sora.unm.edu/sites/default/files/journals/condor/v007n05/p0134-p0138.pdf |doi=10.2307/1361667}}</ref>
+}}
-<ref name=kennedy&white1997>{{cite journal |last1=Kennedy |first1=E.D. |last2=White |first2=D.W. |year=1997 |title=Bewick's Wren (''Thryomanes bewickii'') |editor-last1=Poole |editor-first1=A.  |editor-last2=Gill |editor-first2=F. |work=The Birds of North America |number=315 |publisher=The Academy of Natural Sciences, Philadelphia, PA & The American Ornithologists' Union, Washington, D.C.}}</ref>
+<!--
+<ref name=Thayer>{{cite journal |last1=Thayer |first1=John E. |last2=Bangs |first2=Outram |year=1908 |title=The Present State of the Ornis of Guadaloupe Island |journal=[[Condor (journal)|Condor]] |volume=10 |issue=3 |pages=101–106 |doi=10.2307/1360977 |url=http://sora.unm.edu/sites/default/files/journals/condor/v010n03/p0101-p0106.pdf|jstor=1360977 |hdl=2027/hvd.32044072250186 }}</ref>
-<ref name=Martinez>{{cite journal|author1=Martínez Gómez|author2=Juan E.|author3=Barber, Bruian R.|author4=Peterson, A. Townsend|lastauthoramp=yes|year=2005|title=Phylogenetic position and generic placement of the Socorro Wren (''Thryomanes sissonii'')|journal=[[Auk (journal)|Auk]]|volume=122|issue=1|pages=50–56|doi=10.1642/0004-8038(2005)122[0050:PPAGPO]2.0.CO;2|url=http://www.specifysoftware.org/Informatics/bios/biostownpeterson/MBP_A_2005.pdf|deadurl=yes|archiveurl=https://web.archive.org/web/20081217114932/http://www.specifysoftware.org/Informatics/bios/biostownpeterson/MBP_A_2005.pdf|archivedate=2008-12-17|df=}}</ref>
+<ref name=anthony1901>{{cite journal |last=Anthony |first=A.W. |year=1901 |title=The Guadalupe Wren |journal=[[Condor (journal)|Condor]] |volume=3 |issue=3 |pages=73 |url=http://sora.unm.edu/sites/default/files/journals/condor/v003n03/p0073-p0073.pdf |doi=10.2307/1361475|jstor=1361475 }}</ref>
-<ref name=Thayer>{{cite journal |last1=Thayer |first1=John E. |last2=Bangs |first2=Outram |year=1908 |title=The Present State of the Ornis of Guadaloupe Island |journal=[[Condor (journal)|Condor]] |volume=10 |issue=3 |pages=101–106 |doi=10.2307/1360977 |url=http://sora.unm.edu/sites/default/files/journals/condor/v010n03/p0101-p0106.pdf}}</ref>
+<ref name=kaeding1905>{{cite journal |author=Kaeding, Henry B. |year=1905| title= Birds from the West Coast of Lower California and Adjacent Islands (Part II) |journal=[[Condor (journal)|Condor]] |volume=7 |issue=4 |pages=134–138 |url=http://sora.unm.edu/sites/default/files/journals/condor/v007n05/p0134-p0138.pdf |doi=10.2307/1361667|jstor=1361667}}</ref>
+-->
-}}
 == External links ==
-{{Commons category|Thryomanes bewickii}}
-{{Wikispecies|Thryomanes bewickii}}
 * [http://www.mbr-pwrc.usgs.gov/id/framlst/i7190id.html Bewick's wren - ''Thryomanes bewickii''] - USGS Patuxent Bird Identification InfoCenter
-* [http://www.birds.cornell.edu/AllAboutBirds/BirdGuide/Bewicks_Wren.html Bewick's wren Species Account] – Cornell Lab of Ornithology
+* [https://www.allaboutbirds.org/guide/Bewicks_Wren/ Bewick's wren Species Account] – Cornell Lab of Ornithology
 * {{InternetBirdCollection|bewicks-wren-thryomanes-bewickii|Bewick's wren}}
 * {{VIREO|Bewick's+wren}}
@@ Line 122: / Line 115: @@
 * {{Xeno-canto species|Thryomanes|bewickii|Bewick's wren}}
+{{Passerida|M.|state=collapsed}}
+{{commons category|Thryomanes bewickii}}
 {{Taxonbar|from=Q997457}}
-{{DEFAULTSORT:wren, Bewick's}}
 [[Category:Troglodytidae|Bewick's wren]]
 [[Category:Birds of Mexico]]