Eurasian siskin: Difference between revisions

Eurasian siskin
	Male
	Female; Birds recorded in Surrey, England
Conservation status
	; Least Concern (IUCN 3.1)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Passeriformes
Family:	Fringillidae
Subfamily:	Carduelinae
Genus:	Spinus
Species:	S. spinus
Binomial name
	Spinus spinus; (Linnaeus, 1758)
	Range of the Eurasian siskin; Breeding Resident Non-breeding
Synonyms
	Fringilla spinus Linnaeus, 1758; Carduelis spinus Linnaeus, 1758;

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The Eurasian siskin (Spinus spinus) is a small passerine bird in the finch family Fringillidae. It is also called the European siskin, common siskin or just siskin. Other (archaic) names include black-headed goldfinch,^[2] barley bird and aberdevine.^[3] It is very common throughout Europe and Eurosiberia. It is found in forested areas, both coniferous and mixed woodland where it feeds on seeds of all kinds, especially of alder and conifers.

It can be distinguished from other similar finches by the colour of the plumage. The upper parts are greyish green and the under parts grey-streaked white. Its wings are black with a conspicuous yellow wing bar, and the tail is black with yellow sides. The male has a mainly yellow face and breast, with a neat black cap. Female and young birds have a greyish green head and no cap. It is a trusting, sociable and active bird. The song of this bird is a pleasant mix of twitters and trills. For these reasons it is often raised in captivity.

These birds have an unusual migration pattern as every few years in winter they migrate southwards in large numbers. The reasons for this behaviour are not known but may be related to climatic factors and above all the availability of food. In this way overwintering populations can thrive where food is abundant. This small finch is an acrobatic feeder, often hanging upside-down like a tit. It will visit garden bird feeding stations.

Taxonomy and systematics

The siskin was first described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae as Fringilla spinus, in the 10th edition of Systema Naturae.^[4] In 1760, Brisson described the genus Carduelis, where this species was then placed. Recent taxonomic studies suggest placing it in the genus Spinus.

The scientific name Spinus is from the Ancient Greek spinos, a name for a now-unidentifiable bird.^[5] The English name is from German dialect sisschen or zeischen.^[6]

Despite being found across a wide area it is a monotypic species, that is, there are no distinct subspecies.^[7] This could be explained by a number of factors, such as spatial variability of individuals in breeding areas between years, the large overwintering area which supports a constant genetic interchange, and females having a number of clutches of eggs in one breeding season, each in a different place.^[8]

Description

The siskin is a small, short-tailed bird, 11–12.5 centimetres (4.3–4.9 in) in length^[9]^[10] with a wingspan that ranges from 20 to 23 centimetres (7.9 to 9.1 in).^[11]^[12]^[13] It weighs between 12 and 18 grams (0.42 and 0.63 oz).^[11]^[12]^[14]

The bird's appearance shows sexual dimorphism. The male has a greyish green back; yellow rump; the sides of the tail are yellow and the end is black; the wings are black with a distinctive yellow wing stripe; its breast is yellowish becoming whiter and striped towards the cloaca; it has a black bib (or chin patch) and on its head it has two yellow auriculas and a black cap.^[10] The amount of black on the bib is very variable between males and the size of the bib has been related to dominance within a flock.^[15] The plumage of the female is more olive-coloured than the male. The cap and the auriculas are greenish with a white bib and a rump that is a slightly striped whitish yellow.^[16] The young have a similar colouration to the females, with drab colours and a more subdued plumage.^[10]

The shape of the siskin's beak is determined by its feeding habits. It is strong although it is also slender in order to pick up the seeds on which they feed. The legs and feet are dark brown and the eyes are black.^[16]

It has a rapid and bounding flight pattern that is similar to other finches.^[14]

The siskin is easy to recognize, but in some instances it can be confused with other finches such as the citril finch, the European greenfinch or the European serin.^[16] The Eurasian siskin, in many plumages, is a bright bird. Adult male Eurasian siskins are bright green and yellow with a black cap, and an unstreaked throat and breast. Adult females also usually have green and yellow plumage tones: for example, yellow in the supercilium and on the sides of the breast, green tones in the mantle and yellow in the rump. The ground colour of the underparts of the Eurasian siskin is normally pure white. In females and juveniles, the centre of the belly and lower breast are often largely or entirely unstreaked. The wingbars of the Eurasian siskin are broad and yellow (with the tips white) and the bill is short with a decurved culmen.

male
female

Distribution and habitat

This species can be found across the greater part of Eurosiberia and the north of Africa. Its breeding area is separated into two zones, both on each side of the Palearctic realm: the eastern coast of Asia and the central and northern part of Europe.^[17]

These birds can be found throughout the year in Central Europe and some mountain ranges in the south of the continent. They are present in the north of Scandinavia and in Russia and they over-winter in the Mediterranean basin and the area around the Black Sea. In China they breed in the Khingan Mountains of Inner Mongolia and in Jiangsu province; they spend winter in Tibet, Taiwan, the valleys of the lower Yangtse River and the south east coast.^[18]

The Eurasian siskin is occasionally seen in North America.^[19] There is also a similar and closely related North America counterpart, the pine siskin (Spinus pinus).^[15]

Their seasonal distribution is also marked by the fact that they follow an anomalous migration pattern. Every few years they migrate southwards in larger numbers and the overwintering populations in the Iberian Peninsula are greatly augmented.^[10]^[20] This event has been the object of diverse theories, one theory suggests that it occurs in the years when Norway spruce produces abundant seeds in the centre and north of Europe, causing populations to increase. An alternative theory is that greater migration occurs when the preferred food of alder or birch seed fails. This species will form large flocks outside the breeding season, often mixed with redpolls.

It is a bird that does not remain for long in one area but which varies the areas it uses for breeding and feeding, over-wintering from one year to the next.

Its habitat is forested areas at a particular altitude on a mountain side and they have a certain predilection for humid areas.^[10]^[17] Coniferous woodland, especially spruce, is favoured for breeding. It builds its nest in a tree, laying 2–6 eggs. The British range of this once local breeder has expanded greatly due to an increase in commercial conifer plantations. The siskin also breeds in mixed woodland; while in winter they prefer stubble and crops and areas containing trees with seeds.

Behaviour and ecology

They are very active and restless birds. They are also very social, forming small cohesive flocks especially in autumn and winter.^[21] They are fairly trusting of humans, it being possible to observe them from a short distance. During the breeding season, however, they are much more timid, solitary and difficult to observe. For this reason there is a German legend which says that siskins guard a magic stone in their nests that makes them invisible.^[22]^[23] It is one of the few species which has been described as exhibiting "allofeeding"' behavior, this is where subordinates (of the same sex) regurgitate food for the dominant members of the group,^[24] which creates a strong cohesion in the flocks and implies a hierarchical structure within the group.^[25]

Feeding

The siskin is mainly a granivore, although it varies its diet depending on the season. It feeds in trees, avoiding eating on the ground.^[14]

In autumn and winter, its diet is based on the seeds of deciduous trees such as birch and, above all, alder.^[10]^[20] They also visit cultivated areas and pasture, where they join with other finches in eating the seeds of various Compositae such as thistles, dandelions, Artemisia, knapweeds and other herbaceous plants, such as St. John's wort, meadowsweet and sorrel.^[13]^[20]

In spring, during the breeding season, they are found in coniferous forests. At this time their feeding is based on the seeds of these trees, especially on trees belonging to the genera Abies, Picea and Larix.^[20]^[8] They also feed on elms and poplars. When feeding the young they eat more insects, mainly beetles, as the proteins they contain help the chicks to grow. In summer their feeding is more varied, adding other herbaceous plants to their diet of conifer seeds: goosefoots and other Compositae.^[8]

Reproduction

Pairs are generally formed during the winter period before migration.^[26] The males compete aggressively for the females. As part of the courtship the male plumps up the feathers of the pileus and rump, making itself bigger, extending the tail and singing repeatedly.^[8]^[27] They also make mating flights from tree to tree, although they are not as eye-catching as the flights of the other finches.^[20] They construct a nest that is generally located at the end of a relatively high branch in a conifer, such that the nest is reasonably hidden and difficult to see.^[13]^[20] On the Iberian Peninsula they make their nests in afirs, Scotch pine and Corsican pine.^[28] They form small colonies of up to six pairs with the nests located near to each other.^[25] The nest is small and bowl-shaped. It is made from small twigs, dried grasses, moss and lichen and lined with down.^[20]^[22]^[8]

The first brood is born in mid-April.^[8] The female lays between 2 and 6 eggs.^[20]^[22]^[29] The eggs are white or light grey or light blue, with small brown spots^[20]^[22] and they are approximately 16.5 mm by 12 mm in size.^[13]^[20]^[22] Incubation takes between 10 and 14 days and is carried out entirely by the female.^[12]^[20]^[22] The chicks are altricial and nidicolous. They leave the nest after 15 days in a semi-feathered condition. They then remain close to the nest area for up to a month when, with their plumage now complete, they disperse.^[8] The siskin usually has a second brood, from the middle of June up to the middle of July.^[20]

Song and call

This bird has two calls, both powerful but conflicting, one is descending and the other is ascending, their onomatopoeic sounds can be represented as "tilu" and "tluih".^[10] On occasions they also issue a harsh rattling chirrup.^[10]^[20]

The song is similar to the other finches, a smooth and rapid twitter and trill with a long duration and which is occasionally interrupted by a stronger or shorter syllable. Siskins sing throughout the year and often in groups.^[10]^[20]

Status and conservation

The worldwide population of the siskin is estimated as between 20 and 36 million.^[30] The European population is estimated as between 2.7 and 15 million pairs.^[31]^[32] There does not seem to be a major decline in population numbers and for this reason the IUCN has listed their conservation status as least concern.^[1] The siskin appears in Annex II of the Berne Convention as a protected bird species.^[33]

Relationship with humans

Like many of the finches, the siskin is valued by aviculturalists as a domestic bird for its song and appearance. They do not require specific care and adapt well to captivity, although they do not breed well in captivity.^[29] There are no specific diseases that affect the species, although they can show certain intestinal pathologies associated with a poor diet.^[29] They live for between 11 and 14 years,^[22]^[29] in sharp contrast to the 2 or 3 years it is estimated they live in the wild.^[14]

They form hybrids with some other finches (for example, canaries) giving rise to intermediate birds.^[34]^[35] Hybridisation also occurs in nature without human intervention.^{[failed verification]}^[36] In some areas, individuals that are found are the result of escapes or releases of captive birds.^[37]

Cultural depictions

Poland, Gibraltar, Benin and Belgium have all issued postage stamps bearing the image of the siskin.^[12]^[38]

In Saint Petersburg there is a statue of a siskin, as its colours are the same as the uniform worn by the students at an elite school in the city. These students have come to be known by the sobriquet siskin, Russian: Чиж. This term was popularised in the Russian song "Chizhik-Pyzhik".^[39] There has been a statue of a siskin on the embankment by the First Engineer Bridge since 1994, though it has been stolen and replaced multiple times.

There is a Czech folk song/dance/game "Čížečku, čížečku", in which the siskin is the source of the lore on what happens with the poppy.

Elif Shafak, in the novel Three Daughters of Eve, mentioned a siskin in a pivotal scene in which the heroine, Peri, meets the charismatic and controversial Professor Azur. When Peri entered Professor Azur's office, she found a siskin with yellow-green feathers and a forked tail trapped amid the shelves and stacks of books.

References

^ ^a ^b BirdLife International (2017). "Spinus spinus". IUCN Red List of Threatened Species. 2017: e.T22720354A111126041. doi:10.2305/IUCN.UK.2017-1.RLTS.T22720354A111126041.en. Retrieved 12 November 2021.
^ "Eurasian Siskin (Carduelis spinus) (Linnaeus, 1758)". Avibase. 30 September 2009.
^ Lockwood, W. B. (1993). The Oxford Dictionary of British Bird Names. Oxford University Press. ISBN 0-19-866196-7.
^ "Carduelis spinus". Fauna Europaea. Retrieved 10 October 2008.
^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London, United Kingdom: Christopher Helm. p. 362. ISBN 978-1-4081-2501-4.
^ "Siskin". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
^ Newton, Ian (2003). The Speciation and Biogeography of Birds. London, UK: Academic Press. p. 505. ISBN 0-12-517375-X.
^ ^a ^b ^c ^d ^e ^f ^g "Carduelis spinus" (in Spanish). Terra.es. Archived from the original on 24 January 2008. Retrieved 23 October 2008. This reference is based on theories expounded in Payevsky, V. A. (1994). "Age and sex structure, mortality and spatial winter distribution of siskins (Carduelis spinus) migrating through eastern Baltic area". Vogelwarte. 37: 190–198.
^ Singleton, David G. (11 May 2022). "The alder siskin (Spinus spinus), a bird often found in bands". NATUEND Lifestyle Magazine. Retrieved 30 August 2022.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Mullarney, K.; Svensson, L.; Zetterström, D.; Grant, P. J. (2003). Guía de Campo de las Aves de España y de Europa (in Spanish). Editorial Omega. ISBN 84-282-1218-X.
^ ^a ^b "Eurasian Siskin Carduelis spinus". Madeira Birdwatching. 12 October 2009. Retrieved 30 August 2022.
^ ^a ^b ^c ^d "Eurasian Siskin (Carduelis spinus)". Stamps of Israeli Birds. Archived from the original on 17 July 2011. Retrieved 13 October 2008.
^ ^a ^b ^c ^d Schauenberg, P.; et al. (1979). Fichero Safari Club (Lúgano). Madrid: S.A.P.E. ISBN 84-7461-167-9.
^ ^a ^b ^c ^d Hume, Rob (2002). Guía de Campo de las Aves de España y de Europa (in Spanish). Editorial Omega. ISBN 84-282-1317-8.
^ ^a ^b Senar, J. C.; Camerino, L.; Copete, J. L.; Metcalfe N. B. (1993). "Variation in black bib of the Eurasian siskin (Carduelis spinus) and its role as reliable badge of dominance" (PDF). The Auk. 110 (4): 924–927. doi:10.2307/4088649. JSTOR 4088649.
^ ^a ^b ^c Clement, P (1999). Finches and Sparrows. Princeton University Press. p. 221. ISBN 9780691048789.
^ ^a ^b Senar, J.C.; Borrás, A. "Lúgano en el Atlas de las Aves Reproductoras de España" (PDF) (in Spanish). Retrieved 13 October 2008.
^ China's Species Information Service. "Carduelis spinus". Archived from the original on 15 May 2006. Retrieved 23 October 2008.
^ Borror, A.C. (1963). "Eurasian siskin (Carduelis spinus) in Maine" (PDF). The Auk. 80 (2): 109. doi:10.2307/4082569. JSTOR 4082569. Retrieved 20 October 2008.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ "Lúgano" (in Spanish). Pajaricos.es. Retrieved 12 October 2008.
^ Copete, J.L. (1990). "Observación de un dormidero de Lúganos (Carduelis spinus)". Butlletí del Grup Català d'Anellament (in Spanish). 7.
^ ^a ^b ^c ^d ^e ^f ^g "Tarin des aulnes" (in French). Oiseaux.net. Retrieved 12 October 2008.
^ "Lúgano" (in Spanish). Pajaricos.es. Archived from the original on 12 December 2007. Retrieved 12 October 2008.
^ Senar, J.C.; Borrás, A. (2004). "Sobevivir al invierno: estrategias de las aves invernantes en la Península Ibérica" (PDF). Ardeola. 51 (1): 133–168. Archived from the original (PDF) on 16 September 2006. Retrieved 20 October 2008.
^ ^a ^b Senar, J.C. (April–June 1984). "Allofeeding in Eurasian siskin (Carduelis spinus)" (PDF). Condor. 86 (2). The Cooper Ornithological Society: 213–214. doi:10.2307/1367046. JSTOR 1367046.
^ Senar, J.C.; Copete, J.L. (1990). "Observación de alimentación de cortejo en Lúganos (Carduelis spinus) invernantes". Butlletí del Grup Català d'Anellament. 7. Archived from the original on 1 October 2013. Retrieved 21 July 2010.
^ Newton, I. (1973). Finches. London: Taplinger Publishing. ISBN 0-8008-2720-1.
^ "Eurasian Siskins". Beauty of Birds. 16 September 2021. Retrieved 30 August 2022.
^ ^a ^b ^c ^d "Lúgano-Aves" (in Spanish). Rednaturaleza.com. Archived from the original on 12 February 2007. Retrieved 13 October 2008.
^ BirdLife International. "Species factsheet: Carduelis spinus". Archived from the original on 25 September 2019. Retrieved 12 October 2008.
^ Tucker, G. M.; Heath, M. F. (1994). Birds in Europe: their conservation status. BirdLife Conservation Series 3. Cambridge: BirdLife International. ISBN 0-946888-29-9.
^ Cramp, S.; Perrins, C. M. (1994). Handbook of the Birds of Europe, the Middle East and North Africa. The Birds of the Western Palearctic. Vol. IX. Oxford: Oxford University Press. ISBN 0-19-857506-8.
^ "Boletín Oficial del Estado (España): Ratificación del Convenio de Berna" (PDF) (in Spanish). 1 October 1986. Retrieved 30 August 2022.
^ Global Biodiversity Information Facility. "Carduelis spinus × Serinus canaria". Retrieved 13 October 2008.
^ "Hibridaciones de Carduelis spinus". Archived from the original on 6 November 2007. Retrieved 23 October 2008.
^ McCarthy, Eugene M. (2006). Handbook of Avian Hybrids of the World. Oxford University Press. ISBN 978-0-19-518323-8.
^ Galarza, A. (1989). Urdaibai, avifauna de la ría de Gernika. Diputación Foral de Bizkaia. ISBN 84-404-5084-2.
^ "Eurasian Siskin". Bird Stamps. Archived from the original on 25 October 2000. Retrieved 27 November 2008.{{cite web}}: CS1 maint: unfit URL (link)
^ "Chizhik-Pyzhik". Saint-Petersburg.com. Retrieved 13 October 2008.

External links

Audio recordings from Xeno-canto
Eurasian siskin videos, photos & sounds on the Internet Bird Collection
Ageing and sexing (PDF; 3.1 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
Feathers of Eurasian siskin (Carduelis spinus)
Carduelis spinus in Field Guide: Birds of the World on Flickr
"Carduelis spinus". Avibase.
Chisholm, Hugh, ed. (1911). "Siskin" . Encyclopædia Britannica (11th ed.). Cambridge University Press.

[iucn_status_12_November_2021-1] BirdLife International (2017). "Spinus spinus". IUCN Red List of Threatened Species. 2017: e.T22720354A111126041. doi:10.2305/IUCN.UK.2017-1.RLTS.T22720354A111126041.en. Retrieved 12 November 2021.

[2] "Eurasian Siskin (Carduelis spinus) (Linnaeus, 1758)". Avibase. 30 September 2009.

[Lockwood-3] Lockwood, W. B. (1993). The Oxford Dictionary of British Bird Names. Oxford University Press. ISBN 0-19-866196-7.

[4] "Carduelis spinus". Fauna Europaea. Retrieved 10 October 2008.

[job-5] Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London, United Kingdom: Christopher Helm. p. 362. ISBN 978-1-4081-2501-4.

[OED-6] "Siskin". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)

[Newton-7] Newton, Ian (2003). The Speciation and Biogeography of Birds. London, UK: Academic Press. p. 505. ISBN 0-12-517375-X.

[terra-8] ^ ^a ^b ^c ^d ^e ^f ^g "Carduelis spinus" (in Spanish). Terra.es. Archived from the original on 24 January 2008. Retrieved 23 October 2008. This reference is based on theories expounded in Payevsky, V. A. (1994). "Age and sex structure, mortality and spatial winter distribution of siskins (Carduelis spinus) migrating through eastern Baltic area". Vogelwarte. 37: 190–198.

[9] Singleton, David G. (11 May 2022). "The alder siskin (Spinus spinus), a bird often found in bands". NATUEND Lifestyle Magazine. Retrieved 30 August 2022.

[sven-10] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Mullarney, K.; Svensson, L.; Zetterström, D.; Grant, P. J. (2003). Guía de Campo de las Aves de España y de Europa (in Spanish). Editorial Omega. ISBN 84-282-1218-X.

[madeira-11] "Eurasian Siskin Carduelis spinus". Madeira Birdwatching. 12 October 2009. Retrieved 30 August 2022.

[isr-12] "Eurasian Siskin (Carduelis spinus)". Stamps of Israeli Birds. Archived from the original on 17 July 2011. Retrieved 13 October 2008.

[fichero-13] Schauenberg, P.; et al. (1979). Fichero Safari Club (Lúgano). Madrid: S.A.P.E. ISBN 84-7461-167-9.

[hume-14] Hume, Rob (2002). Guía de Campo de las Aves de España y de Europa (in Spanish). Editorial Omega. ISBN 84-282-1317-8.

[blackrib-15] Senar, J. C.; Camerino, L.; Copete, J. L.; Metcalfe N. B. (1993). "Variation in black bib of the Eurasian siskin (Carduelis spinus) and its role as reliable badge of dominance" (PDF). The Auk. 110 (4): 924–927. doi:10.2307/4088649. JSTOR 4088649.

[Finches-16] Clement, P (1999). Finches and Sparrows. Princeton University Press. p. 221. ISBN 9780691048789.

[mma-17] Senar, J.C.; Borrás, A. "Lúgano en el Atlas de las Aves Reproductoras de España" (PDF) (in Spanish). Retrieved 13 October 2008.

[18] China's Species Information Service. "Carduelis spinus". Archived from the original on 15 May 2006. Retrieved 23 October 2008.

[19] Borror, A.C. (1963). "Eurasian siskin (Carduelis spinus) in Maine" (PDF). The Auk. 80 (2): 109. doi:10.2307/4082569. JSTOR 4082569. Retrieved 20 October 2008.

[paj-20] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ "Lúgano" (in Spanish). Pajaricos.es. Retrieved 12 October 2008.

[21] Copete, J.L. (1990). "Observación de un dormidero de Lúganos (Carduelis spinus)". Butlletí del Grup Català d'Anellament (in Spanish). 7.

[fr-22] ^ ^a ^b ^c ^d ^e ^f ^g "Tarin des aulnes" (in French). Oiseaux.net. Retrieved 12 October 2008.

[23] "Lúgano" (in Spanish). Pajaricos.es. Archived from the original on 12 December 2007. Retrieved 12 October 2008.

[24] Senar, J.C.; Borrás, A. (2004). "Sobevivir al invierno: estrategias de las aves invernantes en la Península Ibérica" (PDF). Ardeola. 51 (1): 133–168. Archived from the original (PDF) on 16 September 2006. Retrieved 20 October 2008.

[allo-25] Senar, J.C. (April–June 1984). "Allofeeding in Eurasian siskin (Carduelis spinus)" (PDF). Condor. 86 (2). The Cooper Ornithological Society: 213–214. doi:10.2307/1367046. JSTOR 1367046.

[26] Senar, J.C.; Copete, J.L. (1990). "Observación de alimentación de cortejo en Lúganos (Carduelis spinus) invernantes". Butlletí del Grup Català d'Anellament. 7. Archived from the original on 1 October 2013. Retrieved 21 July 2010.

[27] Newton, I. (1973). Finches. London: Taplinger Publishing. ISBN 0-8008-2720-1.

[28] "Eurasian Siskins". Beauty of Birds. 16 September 2021. Retrieved 30 August 2022.

[red-29] "Lúgano-Aves" (in Spanish). Rednaturaleza.com. Archived from the original on 12 February 2007. Retrieved 13 October 2008.

[30] BirdLife International. "Species factsheet: Carduelis spinus". Archived from the original on 25 September 2019. Retrieved 12 October 2008.

[31] Tucker, G. M.; Heath, M. F. (1994). Birds in Europe: their conservation status. BirdLife Conservation Series 3. Cambridge: BirdLife International. ISBN 0-946888-29-9.

[32] Cramp, S.; Perrins, C. M. (1994). Handbook of the Birds of Europe, the Middle East and North Africa. The Birds of the Western Palearctic. Vol. IX. Oxford: Oxford University Press. ISBN 0-19-857506-8.

[33] "Boletín Oficial del Estado (España): Ratificación del Convenio de Berna" (PDF) (in Spanish). 1 October 1986. Retrieved 30 August 2022.

[34] Global Biodiversity Information Facility. "Carduelis spinus × Serinus canaria". Retrieved 13 October 2008.

[35] "Hibridaciones de Carduelis spinus". Archived from the original on 6 November 2007. Retrieved 23 October 2008.

[Avi-36] McCarthy, Eugene M. (2006). Handbook of Avian Hybrids of the World. Oxford University Press. ISBN 978-0-19-518323-8.

[37] Galarza, A. (1989). Urdaibai, avifauna de la ría de Gernika. Diputación Foral de Bizkaia. ISBN 84-404-5084-2.

[38] "Eurasian Siskin". Bird Stamps. Archived from the original on 25 October 2000. Retrieved 27 November 2008.{{cite web}}: CS1 maint: unfit URL (link)

[39] "Chizhik-Pyzhik". Saint-Petersburg.com. Retrieved 13 October 2008.

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@@ Line 10: / Line 10: @@
 | status = LC
 | status_system = IUCN3.1
-| status_ref = <ref name=IUCN>{{cite iucn|url=https://www.iucnredlist.org/details/22720354/0 |title=''Carduelis spinus'' |author=BirdLife International |author-link=BirdLife International |year=2012 |access-date=26 November 2013}}</ref>
+| status_ref = <ref name="iucn status 12 November 2021">{{cite iucn |author=BirdLife International |date=2017 |title=''Spinus spinus'' |volume=2017 |page=e.T22720354A111126041 |doi=10.2305/IUCN.UK.2017-1.RLTS.T22720354A111126041.en |access-date=12 November 2021}}</ref>
 | genus = Spinus
 | species = spinus
 | authority = ([[Carl Linnaeus|Linnaeus]], [[10th edition of Systema Naturae|1758]])
-| range_map = Rangemap-tarin.PNG
+| range_map = SpinusSpinusIUCN.svg
-| range_map_caption = Range of the Eurasian siskin<br/>{{legend0|#FFFF00|Summer|border=solid 1px black}} {{legend0|#008000|Resident|border=solid 1px black}} {{legend0|#0080FF|Winter|border=solid 1px black}}
+| range_map_caption = Range of the Eurasian siskin<br/>{{leftlegend|#00FF00|Breeding|outline=gray}} {{leftlegend|#008000|Resident|outline=gray}}  {{leftlegend|#007FFF|Non-breeding|outline=gray}}
 | synonyms =
 *''Fringilla spinus'' <small>Linnaeus, 1758</small>
-*''Carduelis spinus''<small>Linnaeus, 1758</small>
+*''Carduelis spinus'' <small>Linnaeus, 1758</small>
 }}
-The '''Eurasian siskin''' (''Spinus spinus'') is a small [[passerine]] bird in the [[finch]] family Fringillidae. It is also called the '''European siskin''', '''common siskin''' or just '''siskin'''. Other (archaic) names include ''black-headed goldfinch'',<ref>{{cite web| title  = Eurasian Siskin (''Carduelis spinus'') (Linnaeus, 1758)| url = http://avibase.bsc-eoc.org/species.jsp?lang=EN&avibaseid=9820CECA12EC737D| publisher=Avibase| date = 30 September 2009}}</ref> ''barley bird'' and ''aberdevine''.<ref name='Lockwood'>{{cite book|title=The Oxford Dictionary of British Bird Names|last=Lockwood|first=W. B.|year=1993|publisher=[[Oxford University Press]]|isbn=0-19-866196-7}}</ref> It is very common throughout Europe and [[Palearctic|Eurosiberia]]. It is found in forested areas, both [[conifer]]ous and [[mixed woodland]] where it feeds on seeds of all kinds, especially of [[alder]] and conifers.
+The '''Eurasian siskin''' ('''''Spinus spinus''''') is a small [[passerine]] bird in the [[finch]] family Fringillidae. It is also called the '''European siskin''', '''common siskin''' or just '''siskin'''. Other (archaic) names include ''black-headed goldfinch'',<ref>{{cite web| title  = Eurasian Siskin (''Carduelis spinus'') (Linnaeus, 1758)| url = http://avibase.bsc-eoc.org/species.jsp?lang=EN&avibaseid=9820CECA12EC737D | publisher=Avibase| date = 30 September 2009}}</ref> ''barley bird'' and ''aberdevine''.<ref name='Lockwood'>{{cite book|title=The Oxford Dictionary of British Bird Names|last=Lockwood|first=W. B.|year=1993|publisher=[[Oxford University Press]]|isbn=0-19-866196-7}}</ref> It is very common throughout Europe and [[Palearctic|Eurosiberia]]. It is found in forested areas, both [[conifer]]ous and [[mixed woodland]] where it feeds on seeds of all kinds, especially of [[alder]] and conifers.
 It can be distinguished from other similar finches by the colour of the plumage. The upper parts are greyish green and the under parts grey-streaked white. Its wings are black with a conspicuous yellow wing bar, and the tail is black with yellow sides. The male has a mainly yellow face and breast, with a neat black cap. Female and young birds have a greyish green head and no cap. It is a trusting, sociable and active bird. The song of this bird is a pleasant mix of twitters and trills. For these reasons it is often raised in captivity.
@@ Line 28: / Line 28: @@
 ==Taxonomy and systematics==
-The siskin was first was described by [[Carl Linnaeus]] in his landmark 1758 [[10th edition of Systema Naturae|10th edition of ''Systema Naturae'']] as ''Fringilla spinus'', in the [[10th edition of Systema Naturae|10th edition]] of ''[[Systema Naturae]]''.<ref>{{cite web|url = http://www.faunaeur.org/full_results.php?id=97471 |title = ''Carduelis spinus'' |access-date = 10 October 2008|work=Fauna Europaea}}</ref> In 1760, [[Mathurin Jacques Brisson|Brisson]] described the genus ''[[Carduelis]]'', where this species was then placed. Recent taxonomic studies suggest placing it in the genus ''Spinus''.
+The siskin was first described by [[Carl Linnaeus]] in his landmark 1758 [[10th edition of Systema Naturae|10th edition of ''Systema Naturae'']] as ''Fringilla spinus'', in the [[10th edition of Systema Naturae|10th edition]] of ''[[Systema Naturae]]''.<ref>{{cite web|url = https://fauna-eu.org/cdm_dataportal/taxon/7936e3c2-09b8-4618-a8ba-29bc492edc26 |title = ''Carduelis spinus'' |access-date = 10 October 2008|work=Fauna Europaea}}</ref> In 1760, [[Mathurin Jacques Brisson|Brisson]] described the genus ''[[Carduelis]]'', where this species was then placed. Recent taxonomic studies suggest placing it in the genus ''Spinus''.
-The scientific name ''Spinus'' is from the [[Ancient Greek]] ''spinos'', a name for a now-unidentifiable bird.<ref name =job>{{cite book | last= Jobling | first= James A. | year= 2010| title= The Helm Dictionary of Scientific Bird Names | url= https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling | publisher=Christopher Helm | location = London, United Kingdom | isbn = 978-1-4081-2501-4 | page = 362}}</ref> The English name is from German dialect {{lang|de|sisschen}} or {{lang|de|zeischen}},<ref name=OED>{{ OED |Siskin}}</ref>
+The scientific name ''Spinus'' is from the [[Ancient Greek]] ''spinos'', a name for a now-unidentifiable bird.<ref name =job>{{cite book | last= Jobling | first= James A. | year= 2010| title= The Helm Dictionary of Scientific Bird Names | url= https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling | publisher=Christopher Helm | location = London, United Kingdom | isbn = 978-1-4081-2501-4 | page = 362}}</ref> The English name is from German dialect {{lang|de|sisschen}} or {{lang|de|zeischen}}.<ref name=OED>{{ OED |Siskin}}</ref>
-Despite being found across a wide area it is a monotypic species, that is, there are no distinct [[subspecies]].<ref name = "Newton">{{cite book |title = The Speciation and Biogeography of Birds |first = Ian | last = Newton | page = 505 | publisher = Academic Press | location = London, UK | isbn = 0-12-517375-X |year = 2003 |url = https://books.google.com/books?id=dYJ8CMm-zH0C&pg=PA505}}</ref> This could be explained by a number of factors, such as spatial variability of individuals in breeding areas between years, the large overwintering area which supposes a constant genetic interchange,{{Clarify|date=October 2011}} and females having a number of clutches of eggs in one breeding season, each in a different place.<ref name=terra>{{cite web|url = http://www.terra.es/personal/sobradil/lugano/home.htm|title = ''Carduelis spinus''|access-date = 23 October 2008|publisher=Terra.es}} This reference is based on theories expounded in {{cite journal|last=Payevsky|first=V. A.|year=1994|title=Age and sex structure, mortality and spatial winter distribution of siskins (''Carduelis spinus'') migrating through eastern Baltic area|journal=Vogelwarte|volume=37|pages=190–198}}</ref>
+Despite being found across a wide area it is a monotypic species, that is, there are no distinct [[subspecies]].<ref name = "Newton">{{cite book |title = The Speciation and Biogeography of Birds |first = Ian | last = Newton | page = 505 | publisher = Academic Press | location = London, UK | isbn = 0-12-517375-X |year = 2003 |url = https://books.google.com/books?id=dYJ8CMm-zH0C&pg=PA505}}</ref> This could be explained by a number of factors, such as spatial variability of individuals in breeding areas between years, the large overwintering area which supports a constant genetic interchange, and females having a number of clutches of eggs in one breeding season, each in a different place.<ref name=terra>{{cite web|url = http://www.terra.es/personal/sobradil/lugano/home.htm|title = ''Carduelis spinus''|access-date = 23 October 2008|publisher=Terra.es|url-status=dead|archive-date=2008-01-24|archive-url=https://web.archive.org/web/20080124215704/http://www.terra.es/personal/sobradil/lugano/home.htm|language=es}} This reference is based on theories expounded in {{cite journal|last=Payevsky|first=V. A.|year=1994|title=Age and sex structure, mortality and spatial winter distribution of siskins (''Carduelis spinus'') migrating through eastern Baltic area|journal=Vogelwarte|volume=37|pages=190–198}}</ref>
-The [[phylogeny]] has been obtained by Antonio Arnaiz-Villena et al.<ref>{{cite journal|last=Zamora|first=J.|author2=Moscoso, J.|author3=Ruiz-del-Valle, V.|author4=Ernesto, L.|author5=Serrano-Vela, J. I.|author6=Ira-Cachafeiro, J.|author7=Arnaiz-Villena, A.|s2cid=46892185|year=2006|title=Conjoint mitochondrial phylogenetic trees for canaries ''Serinus'' spp. and goldfinches ''Carduelis'' spp. show several specific polytomies|journal=Ardeola|volume=53|issue=1|pages=1–17}}</ref><ref>{{cite journal|last=Arnaiz-Villena|first=A.|author2=Ruiz-del-Valle, V.|author3=Moscoso, J.|author4=Serrano-Vela, J. I.|author5=Zamora, J.|s2cid=46892185|year=2007|title=mtDNA phylogeny of North American ''Carduelis pinus'' group|journal=Ardeola|volume=54|issue=1|pages=1–14}}</ref>
-===Evolution===
-This bird may have reached America either from Asia<ref name="Finches">{{cite book|last=Clement|first=P|title=Finches and Sparrows|publisher=Princeton University Press|year=1999|pages=221|isbn = 9780691048789}}</ref><ref name="Arnaiz">{{cite journal|last=Arnaiz-Villena|first=A|author2=Ruiz-del-Valle V|author3=Reguera R|author4=Gomez-Prieto P|author5=Serrano-Vela JI|title=What Might have been the Ancestor of New World Siskins?|journal=The Open Ornithology Journal|year=2012|volume=1|pages=46–47|url=http://www.benthamscience.com/open/tooenij/articles/V001/46TOOENIJ.pdf|doi=10.2174/1874453200801010046|url-status=dead|archive-url=https://web.archive.org/web/20131001210831/http://www.benthamscience.com/open/tooenij/articles/V001/46TOOENIJ.pdf|archive-date=1 October 2013}}</ref> or from Europe (Greenland/Iceland). It is the extant parental species of one of the ''Spinus''/''Carduelis'' three evolutive North American radiations of ''atriceps'', ''pinus'' and ''dominicensis'' finches. It has been recorded both in the Aleutian Islands<ref name="Finches" /><ref name="Arnaiz" /> and the east: the Labrador Peninsula and St. Lawrence River mouth (Canada).<ref>{{cite journal|last=Arnaiz-Villena |first=A |author2=Areces C |author3=Rey D |author4=Enríquez-de-Salamanca M |author5=Alonso-Rubio J |author6=Ruiz-del-Valle V |title=Three different North American siskin/goldfinch evolutionary radiations (genus ''Carduelis''): pine siskin green morphs and European siskins in America |journal=The Open Ornithology Journal |year=2012 |volume=5 |pages=73–81 |url=http://www.benthamscience.com/open/tooenij/articles/V005/73TOOENIJ.pdf |doi=10.2174/1874453201205010073 |url-status=dead |archive-url=https://web.archive.org/web/20130921054120/http://www.benthamscience.com/open/tooenij/articles/V005/73TOOENIJ.pdf |archive-date=21 September 2013}}</ref> This raises the possibility that this bird entered (or may still try to enter) America through Greenland/Iceland from Western Europe.
 ==Description==
-The siskin is a small, short-tailed bird, {{convert|11|–|12.5|cm|in}} in length<ref name=BWP>{{cite book |title=The Birds of the western Palearctic|edition=abridged<!-- the isbn given is that of the full set, but "abridged" suggests concise ed intended --> |year=1997|publisher=[[Oxford University Press]]|isbn=0-19-854099-X}}</ref><!-- edition/vol needed as given, also page numbers -->{{full citation needed|date=October 2011}}<ref name=sven /> with a wingspan that ranges from {{convert|20|–|23|cm|in}}.<ref name=BWP /><ref name=isr /><ref name=fichero /> It weighs between {{convert|10|–|18|g|oz}}.<ref name=BWP /><ref name=isr /><ref name=hume />
+The siskin is a small, short-tailed bird, {{convert|11|–|12.5|cm|in}} in length<ref>{{cite web|url=https://natuend.com/the-alder-siskin-spinus-spinus-a-bird-often-found-in-bands/|title=The alder siskin (Spinus spinus), a bird often found in bands|last=Singleton|first=David G.|date=2022-05-11|publisher=NATUEND Lifestyle Magazine|access-date=2022-08-30}}</ref><ref name=sven /> with a wingspan that ranges from {{convert|20| to |23|cm|in}}.<ref name=madeira>{{cite web|url=https://www.madeirabirds.com/eurasian-siskin-carduelis-spinus|title=Eurasian Siskin Carduelis spinus|publisher=Madeira Birdwatching|date=2009-10-12|access-date=2022-08-30}}</ref><ref name=isr /><ref name=fichero /> It weighs between {{convert|12| and |18|g|oz}}.<ref name=madeira/><ref name=isr /><ref name=hume/>
-The bird's appearance shows [[sexual dimorphism]]. The male has a greyish green back; yellow rump; the sides of the tail are yellow and the end is black; the wings are black with a distinctive yellow wing stripe; its breast is yellowish becoming whiter and striped towards the [[cloaca]]; it has a black bib (or chin patch) and on its head it has two yellow [[Ear|auricula]]s and a black cap.<ref name=sven /> The amount of black on the bib is very variable between males and the size of the bib has been related to dominance within a flock.<ref name=blackrib /> The [[plumage]] of the female is more olive-coloured than the male. The cap and the auriculas are greenish with a white bib and a rump that is a slightly striped whitish yellow.{{Citation needed|date=October 2011}} The young have a similar colouration to the females, with drab colours and a more subdued plumage.<ref name=sven />
+The bird's appearance shows [[sexual dimorphism]]. The male has a greyish green back; yellow rump; the sides of the tail are yellow and the end is black; the wings are black with a distinctive yellow wing stripe; its breast is yellowish becoming whiter and striped towards the [[cloaca]]; it has a black bib (or chin patch) and on its head it has two yellow [[Ear|auricula]]s and a black cap.<ref name=sven /> The amount of black on the bib is very variable between males and the size of the bib has been related to dominance within a flock.<ref name=blackrib /> The [[plumage]] of the female is more olive-coloured than the male. The cap and the auriculas are greenish with a white bib and a rump that is a slightly striped whitish yellow.<ref name="Finches">{{cite book|last=Clement|first=P|title=Finches and Sparrows|publisher=Princeton University Press|year=1999|pages=221|isbn = 9780691048789}}</ref> The young have a similar colouration to the females, with drab colours and a more subdued plumage.<ref name=sven />
-The shape of the siskin's [[beak]] is determined by its feeding habits. It is strong although it is also slender in order to pick up the seeds on which they feed. The legs and feet are dark brown and the eyes are black.{{Citation needed|date=October 2011}}
+The shape of the siskin's [[beak]] is determined by its feeding habits. It is strong although it is also slender in order to pick up the seeds on which they feed. The legs and feet are dark brown and the eyes are black.<ref name="Finches"/>
 It has a rapid and bounding flight pattern that is similar to other finches.<ref name=hume>{{cite book |last = Hume|first= Rob| publisher=Editorial Omega| title = Guía de Campo de las Aves de España y de Europa| year = 2002| isbn = 84-282-1317-8|language=es}}</ref>
-The siskin is easy to recognize, but in some instances it can be confused with other finches such as the [[citril finch]], the [[European greenfinch]] or the [[European serin]].{{Citation needed|date=October 2011}} The Eurasian siskin, in many plumages, is a bright bird. Adult male Eurasian siskins are bright green and yellow with a black cap, and an unstreaked throat and breast. Adult females also usually have green and yellow plumage tones: for example, yellow in the [[supercilium]] and on the sides of the breast, green tones in the mantle and yellow in the rump. The ground colour of the underparts of the Eurasian siskin is normally pure white. In females and juveniles, the centre of the belly and lower breast are often largely or entirely unstreaked. The wingbars of the Eurasian siskin are broad and yellow (with the tips white) and the bill is short with a decurved culmen.
+The siskin is easy to recognize, but in some instances it can be confused with other finches such as the [[citril finch]], the [[European greenfinch]] or the [[European serin]].<ref name="Finches"/>  The Eurasian siskin, in many plumages, is a bright bird. Adult male Eurasian siskins are bright green and yellow with a black cap, and an unstreaked throat and breast. Adult females also usually have green and yellow plumage tones: for example, yellow in the [[supercilium]] and on the sides of the breast, green tones in the mantle and yellow in the rump. The ground colour of the underparts of the Eurasian siskin is normally pure white. In females and juveniles, the centre of the belly and lower breast are often largely or entirely unstreaked. The wingbars of the Eurasian siskin are broad and yellow (with the tips white) and the bill is short with a decurved culmen.
+<gallery mode = packed heights = 180px>
+Eurasian siskin (Spinus spinus) male Biebrzanski.jpg|male
+Eurasian siskin (Spinus spinus) female Biebrzanski.jpg|female
+</gallery>
 ==Distribution and habitat==
+This species can be found across the greater part of [[Palearctic|Eurosiberia]] and the north of Africa. Its breeding area is separated into two zones, both on each side of the [[Palearctic realm]]: the eastern coast of Asia and the central and northern part of Europe.<ref name=mma>{{cite web|url = https://www.miteco.gob.es/es/biodiversidad/temas/inventarios-nacionales/lugano_tcm30-99965.pdf|title = Lúgano en el Atlas de las Aves Reproductoras de España|last1 = Senar|first1 = J.C.|last2 = Borrás|first2 = A.|access-date = 13 October 2008|language = es}}</ref>
-[[File:Adult male siskin.JPG|thumb|left|An adult male at a bird table]]
-This species can be found across the greater part of [[Palearctic|Eurosiberia]] and the north of Africa. Its breeding area is separated into two zones, both on each side of the [[Palearctic realm]]: the eastern coast of Asia and the central and northern part of Europe.<ref name=mma>{{cite web|url = http://www.mma.es/secciones/biodiversidad/inventarios/inb/atlas_aves_reproductoras/pdf/lugano.pdf|title = Lúgano en el Atlas de las Aves Reproductoras de España|last1 = Senar|first1 = J.C.|last2 = Borrás|first2 = A.|access-date = 13 October 2008|language = es}}{{Dead link|date=August 2019 |bot=InternetArchiveBot |fix-attempted=yes}}</ref>
-These birds can be found throughout the year in Central Europe and some mountain ranges in the south of the continent. They are present in the north of [[Scandinavia]] and in [[Russia]] and they over-winter in the [[Mediterranean]] basin and the area around the [[Black Sea]]. In China they breed in the [[Khingan Mountains]] of [[Inner Mongolia]] and in [[Jiangsu]] province; they spend winter in [[Tibet]], Taiwan, the valleys of the lower [[Yangtse River]] and the south east coast.<ref>{{cite web|url = http://www.chinabiodiversity.com/search/english/detail.shtm?cspcode=022110010|title =''Carduelis spinus''|access-date = 23 October 2008|author=China's Species Information Service|archive-url=https://web.archive.org/web/20060515001752/http://www.chinabiodiversity.com/search/english/detail.shtm?cspcode=022110010|archive-date=15 May 2006}}</ref>
+These birds can be found throughout the year in Central Europe and some mountain ranges in the south of the continent. They are present in the north of [[Scandinavia]] and in [[Russia]] and they over-winter in the [[Mediterranean]] basin and the area around the [[Black Sea]]. In China they breed in the [[Khingan Mountains]] of [[Inner Mongolia]] and in [[Jiangsu]] province; they spend winter in [[Tibet]], Taiwan, the valleys of the lower [[Yangtse River]] and the south east coast.<ref>{{cite web|url = http://www.chinabiodiversity.com/search/english/detail.shtm?cspcode=022110010|title =''Carduelis spinus''|access-date = 23 October 2008|url-status=dead|author=China's Species Information Service|archive-url=https://web.archive.org/web/20060515001752/http://www.chinabiodiversity.com/search/english/detail.shtm?cspcode=022110010|archive-date=15 May 2006}}</ref>
-The Eurasian siskin is occasionally seen in North America.<ref>{{cite journal|last = Borror|first= A.C.|year= 1963|title= Eurasian siskin (''Carduelis spinus'') in Maine|journal=The Auk|volume= 80|issue= 2|page= 109|doi= 10.2307/4082569|jstor= 4082569|url= http://sora.unm.edu/sites/default/files/journals/auk/v080n02/p0201-p0201.pdf|access-date= 20 October 2008}}</ref> There is also a similar and closely related North America counterpart, the [[pine siskin]], ''Spinus pinus''.<ref name=blackrib>{{cite journal|author1=Senar, J. C. |author2=Camerino, L. |author3=Copete, J. L. |author4=Metcalfe N. B. |year= 1993|title= Variation in black bib of the Eurasian siskin (''Carduelis spinus'') and its role as reliable badge of dominance|journal=The Auk|volume= 110|issue= 4|pages= 924–927|url= http://sora.unm.edu/sites/default/files/journals/auk/v110n04/p0924-p0927.pdf|doi=10.2307/4088649|jstor=4088649}}</ref>
+The Eurasian siskin is occasionally seen in North America.<ref>{{cite journal|last = Borror|first= A.C.|year= 1963|title= Eurasian siskin (''Carduelis spinus'') in Maine|journal=The Auk|volume= 80|issue= 2|page= 109|doi= 10.2307/4082569|jstor= 4082569|url= http://sora.unm.edu/sites/default/files/journals/auk/v080n02/p0201-p0201.pdf|access-date= 20 October 2008}}</ref> There is also a similar and closely related North America counterpart, the [[pine siskin]] (''Spinus pinus'').<ref name=blackrib>{{cite journal|author1=Senar, J. C. |author2=Camerino, L. |author3=Copete, J. L. |author4=Metcalfe N. B. |year= 1993|title= Variation in black bib of the Eurasian siskin (''Carduelis spinus'') and its role as reliable badge of dominance|journal=The Auk|volume= 110|issue= 4|pages= 924–927|url= http://sora.unm.edu/sites/default/files/journals/auk/v110n04/p0924-p0927.pdf|doi=10.2307/4088649|jstor=4088649}}</ref>
 Their seasonal distribution is also marked by the fact that they follow an anomalous [[Animal migration|migration]] pattern. Every few years they migrate southwards in larger numbers and the overwintering populations in the [[Iberian Peninsula]] are greatly augmented.<ref name=sven>{{cite book|author1=Mullarney, K. |author2=Svensson, L. |author3=Zetterström, D. |author4=Grant, P. J. | publisher=Editorial Omega| title = Guía de Campo de las Aves de España y de Europa| year = 2003| isbn = 84-282-1218-X|language=es}}</ref><ref name=paj>{{cite web|url = http://www.pajaricos.es/mas/maslugano.htm|title = Lúgano|access-date = 12 October 2008|publisher=Pajaricos.es|language=es}}</ref> This event has been the object of diverse theories, one theory suggests that it occurs in the years when [[Picea abies|Norway spruce]] produces abundant seeds in the centre and north of Europe, causing populations to increase. An alternative theory is that greater migration occurs when the preferred food of [[alder]] or [[birch]] seed fails. This species will form large flocks outside the breeding season, often mixed with [[redpoll]]s.
@@ Line 65: / Line 64: @@
 ==Behaviour and ecology==
-[[File:Carduelis spinus 1 tom (Marek Szczepanek).jpg|thumb|240px|A perched male observing its next destination]]
-They are very active and restless birds. They are also very social, forming small cohesive flocks especially in autumn and winter.<ref>{{cite journal| last = Copete|first=J.L.| title = Observación de un dormidero de Lúganos (''Carduelis spinus'')| year = 1990| journal=Butlletí del Grup Català d'Anellament| volume = 7| url = http://www.raco.cat/index.php/ButlletiAnellament/article/view/70755/89502}}</ref> They are fairly trusting of humans, it being possible to observe them from a short distance. During the breeding season, however, they are much more timid, solitary and difficult to observe. For this reason there is a [[German folklore|German legend]] which says that siskins guard a magic stone in their nests that makes them invisible.<ref name=fr>{{cite web|url = http://www.oiseaux.net/oiseaux/tarin.des.aulnes.html|title = Tarin des aulnes|access-date = 12 October 2008|publisher=Oiseaux.net|language = fr}}</ref><ref>{{cite web|url = http://www.pajaricos.es/l/l1/lugano.htm|title = Lúgano|access-date = 12 October 2008|publisher = Pajaricos.es|language = es|url-status = dead|archive-url = https://web.archive.org/web/20071212211937/http://www.pajaricos.es/l/l1/lugano.htm|archive-date = 12 December 2007}}</ref> It is one of the few species which has been described as exhibiting "allofeeding"' behavior, this is where subordinates (of the same sex) regurgitate food for the dominant members of the group,<ref>{{cite journal|author1= Senar, J.C.|author2= Borrás, A.|year= 2004|title= Sobevivir al invierno: estrategias de las aves invernantes en la Península Ibérica|journal= Ardeola|volume= 51|pages= 133–168|url= http://www.ardeola.org/files/Ardeola_51(1)_133-168.pdf|access-date= 20 October 2008|issue= 1|url-status= dead|archive-url= https://web.archive.org/web/20060916153354/http://www.ardeola.org/files/Ardeola_51(1)_133-168.pdf|archive-date= 16 September 2006}}</ref> which creates a strong cohesion in the flocks and implies a hierarchical structure within the group.<ref name=allo>{{cite journal|author=Senar, J.C.|date=April–June 1984|title= Allofeeding in Eurasian siskin (''Carduelis spinus'')|journal=Condor|volume= 86|publisher=The Cooper Ornithological Society|pages= 213–214|url= http://sora.unm.edu/sites/default/files/journals/condor/v086n02/p0213-p0214.pdf|doi=10.2307/1367046|issue=2|jstor=1367046}}</ref>
+They are very active and restless birds. They are also very social, forming small cohesive flocks especially in autumn and winter.<ref>{{cite journal| last = Copete|first=J.L.| title = Observación de un dormidero de Lúganos (''Carduelis spinus'')| year = 1990| journal=Butlletí del Grup Català d'Anellament| volume = 7| url = http://www.raco.cat/index.php/ButlletiAnellament/article/view/70755/89502|language=es}}</ref> They are fairly trusting of humans, it being possible to observe them from a short distance. During the breeding season, however, they are much more timid, solitary and difficult to observe. For this reason there is a [[German folklore|German legend]] which says that siskins guard a magic stone in their nests that makes them invisible.<ref name=fr>{{cite web|url = http://www.oiseaux.net/oiseaux/tarin.des.aulnes.html|title = Tarin des aulnes|access-date = 12 October 2008|publisher=Oiseaux.net|language = fr}}</ref><ref>{{cite web|url = http://www.pajaricos.es/l/l1/lugano.htm|title = Lúgano|access-date = 12 October 2008|publisher = Pajaricos.es|language = es|url-status = dead|archive-url = https://web.archive.org/web/20071212211937/http://www.pajaricos.es/l/l1/lugano.htm|archive-date = 12 December 2007}}</ref> It is one of the few species which has been described as exhibiting "allofeeding"' behavior, this is where subordinates (of the same sex) regurgitate food for the dominant members of the group,<ref>{{cite journal|author1= Senar, J.C.|author2= Borrás, A.|year= 2004|title= Sobevivir al invierno: estrategias de las aves invernantes en la Península Ibérica|journal= Ardeola|volume= 51|pages= 133–168|url= http://www.ardeola.org/files/Ardeola_51(1)_133-168.pdf|access-date= 20 October 2008|issue= 1|url-status= dead|archive-url= https://web.archive.org/web/20060916153354/http://www.ardeola.org/files/Ardeola_51(1)_133-168.pdf|archive-date= 16 September 2006}}</ref> which creates a strong cohesion in the flocks and implies a hierarchical structure within the group.<ref name=allo>{{cite journal|author=Senar, J.C.|date=April–June 1984|title= Allofeeding in Eurasian siskin (''Carduelis spinus'')|journal=Condor|volume= 86|publisher=The Cooper Ornithological Society|pages= 213–214|url= http://sora.unm.edu/sites/default/files/journals/condor/v086n02/p0213-p0214.pdf|doi=10.2307/1367046|issue=2|jstor=1367046}}</ref>
 ===Feeding===
@@ Line 78: / Line 77: @@
 ===Reproduction===
 [[File:Carduelis spinus MHNT.ZOO.2010.11 Anvers.jpg|thumb|Eggs of ''Spinus spinus'']]
-Pairs are generally formed during the winter period before migration.<ref>{{cite journal| author1 = Senar, J.C.| author2 = Copete, J.L.| title = Observación de alimentación de cortejo en Lúganos (''Carduelis spinus'') invernantes| year = 1990| journal = Butlletí del Grup Català d'Anellament| volume = 7| url = http://www.raco.cat/index.php/ButlletiAnellament/article/viewFile/70753/89500| access-date = 21 July 2010| archive-url = https://web.archive.org/web/20131001210830/http://www.raco.cat/index.php/ButlletiAnellament/article/viewFile/70753/89500| archive-date = 1 October 2013| url-status = dead}}</ref> The males compete aggressively for the females. As part of the courtship the male plumps up the feathers of the pileus and rump, making itself bigger, extending the tail and singing repeatedly.<ref name=terra /><ref>{{cite book|last = Newton|first= I.| publisher=Taplinger Publishing| title = Finches| year = 1973| location = London| isbn = 0-8008-2720-1}}</ref> They also make mating flights from tree to tree, although they are not as eye-catching as the flights of the other finches.<ref name=paj /> They construct a [[bird nest|nest]] that is generally located at the end of a relatively high branch in a conifer, such that the nest is reasonably hidden and difficult to see.<ref name=fichero /><ref name=paj /> On the Iberian Peninsula they make their nests in ''[[Abies alba|afirs]],'' [[Pinus sylvestris|Scotch pine]] and [[Pinus nigra|Corsican pine]].<ref>{{cite web|url = http://www.ausdebalears.org/isnoEnd/isnova/loadBird?lang=es&birdId=445|title = Lúgano (''Carduelis spinus'')|access-date = 19 November 2008|work=Enciclopedia Balear de Ornitología}}</ref> They form small colonies of up to six pairs with the nests located near to each other.<ref name=allo /> The nest is small and bowl-shaped. It is made from small twigs, dried grasses, [[moss]] and [[lichen]] and lined with [[down feather|down]].<ref name=paj /><ref name=fr /><ref name=terra />
+Pairs are generally formed during the winter period before migration.<ref>{{cite journal| author1 = Senar, J.C.| author2 = Copete, J.L.| title = Observación de alimentación de cortejo en Lúganos (''Carduelis spinus'') invernantes| year = 1990| journal = Butlletí del Grup Català d'Anellament| volume = 7| url = http://www.raco.cat/index.php/ButlletiAnellament/article/viewFile/70753/89500| access-date = 21 July 2010| archive-url = https://web.archive.org/web/20131001210830/http://www.raco.cat/index.php/ButlletiAnellament/article/viewFile/70753/89500| archive-date = 1 October 2013| url-status = dead}}</ref> The males compete aggressively for the females. As part of the courtship the male plumps up the feathers of the pileus and rump, making itself bigger, extending the tail and singing repeatedly.<ref name=terra /><ref>{{cite book|last = Newton|first= I.| publisher=Taplinger Publishing| title = Finches| year = 1973| location = London| isbn = 0-8008-2720-1}}</ref> They also make mating flights from tree to tree, although they are not as eye-catching as the flights of the other finches.<ref name=paj /> They construct a [[bird nest|nest]] that is generally located at the end of a relatively high branch in a conifer, such that the nest is reasonably hidden and difficult to see.<ref name=fichero /><ref name=paj /> On the Iberian Peninsula they make their nests in ''[[Abies alba|afirs]],'' [[Pinus sylvestris|Scotch pine]] and [[Pinus nigra|Corsican pine]].<ref>{{cite web|url = https://www.beautyofbirds.com/eurasiansiskins.html|title=Eurasian Siskins|date=2021-09-16|publisher=Beauty of Birds|access-date=2022-08-30}}</ref> They form small colonies of up to six pairs with the nests located near to each other.<ref name=allo /> The nest is small and bowl-shaped. It is made from small twigs, dried grasses, [[moss]] and [[lichen]] and lined with [[down feather|down]].<ref name=paj /><ref name=fr /><ref name=terra />
 The first brood is born in mid-April.<ref name=terra /> The female lays between 2 and 6 eggs.<ref name=paj /><ref name=fr /><ref name=red /> The eggs are white or light grey or light blue, with small brown spots<ref name=paj /><ref name=fr /> and they are approximately 16.5&nbsp;mm by 12&nbsp;mm in size.<ref name=fichero>{{cite book| last1 = Schauenberg|first1=P.| publisher=S.A.P.E.| title = Fichero Safari Club (Lúgano)| year = 1979| location = Madrid| isbn = 84-7461-167-9|display-authors=etal}}</ref><ref name=paj /><ref name=fr /> [[Avian incubation|Incubation]] takes between 10 and 14 days and is carried out entirely by the female.<ref name=isr /><ref name=paj /><ref name=fr /> The chicks are [[altricial]] and [[nidicolous]]. They leave the nest after 15 days in a semi-feathered condition. They then remain close to the nest area for up to a month when, with their plumage now complete, they disperse.<ref name=terra/> The siskin usually has a second brood, from the middle of June up to the middle of July.<ref name=paj />
@@ Line 88: / Line 87: @@
 ==Status and conservation==
-The worldwide population of the siskin is estimated as between 20 and 36 million.<ref>{{cite web|url = http://www.birdlife.org/datazone/species/index.html?action=SpcHTMDetails.asp&sid=8813 |title = Species factsheet: ''Carduelis spinus'' |access-date = 12 October 2008|author=BirdLife International}}</ref> The European population is estimated as between 2.7 and 15 million pairs.<ref>{{cite book|author1=Tucker, G. M. |author2=Heath, M. F. | publisher=BirdLife International| title = Birds in Europe: their conservation status |series=BirdLife Conservation Series 3| year = 1994| location = Cambridge| isbn = 0-946888-29-9}}</ref><ref>{{cite book|author1=Cramp, S. |author2=Perrins, C. M. | publisher=Oxford University Press| title = Handbook of the Birds of Europe, the Middle East and North Africa. The Birds of the Western Palearctic. Vol. IX.| year = 1994| location = Oxford| isbn = 0-19-857506-8}}</ref> There does not seem to be a major decline in population numbers and for this reason the [[IUCN]] has listed their conservation status as [[least concern]].<ref name=IUCN/> The siskin appears in Annex II of the [[Berne Convention (conservation)|Berne Convention]] as a protected bird species.<ref>{{cite web|language=es|url = http://www.club-caza.com/legislacion/docs/europea/4-Convenio_de_Berna.pdf |title = Boletín Oficial del Estado (España): Ratificación del Convenio de Berna |access-date = 12 October 2008|date = 1 October 1986}}</ref>
+The worldwide population of the siskin is estimated as between 20 and 36 million.<ref>{{cite web |url = http://www.birdlife.org/datazone/species/index.html?action=SpcHTMDetails.asp&sid=8813 |title = Species factsheet: ''Carduelis spinus'' |access-date = 12 October 2008 |author = BirdLife International |archive-date = 25 September 2019 |archive-url = https://web.archive.org/web/20190925005302/http://datazone.birdlife.org/home |url-status = dead }}</ref> The European population is estimated as between 2.7 and 15 million pairs.<ref>{{cite book|author1=Tucker, G. M. |author2=Heath, M. F. | publisher=BirdLife International| title = Birds in Europe: their conservation status |series=BirdLife Conservation Series 3| year = 1994| location = Cambridge| isbn = 0-946888-29-9}}</ref><ref>{{cite book|author1=Cramp, S. |author2=Perrins, C. M. | publisher=Oxford University Press| title = Handbook of the Birds of Europe, the Middle East and North Africa. The Birds of the Western Palearctic. Vol. IX.| year = 1994| location = Oxford| isbn = 0-19-857506-8}}</ref> There does not seem to be a major decline in population numbers and for this reason the [[IUCN]] has listed their conservation status as [[least concern]].<ref name="iucn status 12 November 2021" /> The siskin appears in Annex II of the [[Berne Convention (conservation)|Berne Convention]] as a protected bird species.<ref>{{cite web|language=es|url = http://www.club-caza.com/legislacion/docs/europea/4-Convenio_de_Berna.pdf |title = Boletín Oficial del Estado (España): Ratificación del Convenio de Berna |access-date =2022-08-30|date = 1 October 1986}}</ref>
 ==Relationship with humans==
 [[File:Sagra dei Osei Carduelis spinus.JPG|thumb|Structure holding cages used for listening to siskins at the [[Vinkensport|singing contest]] of Sagra dei Osei, Italy]]
-Like many of the finches, the siskin is valued by [[aviculture|aviculturalists]] as a domestic bird for its song and appearance. They do not require specific care and adapt well to captivity, although they do not breed well in captivity.<ref name=red>{{cite web|url = http://www.rednaturaleza.com/aves_doc.asp?p=Lugano|title = Lúgano-Aves|access-date = 13 October 2008|publisher = Rednaturaleza.com|url-status = dead|archive-url = https://web.archive.org/web/20070212123212/http://www.rednaturaleza.com/aves_doc.asp?p=Lugano|archive-date = 12 February 2007}}</ref> There are no specific diseases that affect the species, although they can show certain intestinal pathologies associated with a poor diet.<ref name=red /> They live for between 11 and 14 years,<ref name=fr /><ref name=red /> in sharp contrast to the 2 or 3 years it is estimated they live in the wild.<ref name=hume />
+Like many of the finches, the siskin is valued by [[aviculture|aviculturalists]] as a domestic bird for its song and appearance. They do not require specific care and adapt well to captivity, although they do not breed well in captivity.<ref name=red>{{cite web|url = http://www.rednaturaleza.com/aves_doc.asp?p=Lugano|title = Lúgano-Aves|access-date = 13 October 2008|publisher = Rednaturaleza.com|url-status = dead|archive-url = https://web.archive.org/web/20070212123212/http://www.rednaturaleza.com/aves_doc.asp?p=Lugano|archive-date = 12 February 2007|language=es}}</ref> There are no specific diseases that affect the species, although they can show certain intestinal pathologies associated with a poor diet.<ref name=red /> They live for between 11 and 14 years,<ref name=fr /><ref name=red /> in sharp contrast to the 2 or 3 years it is estimated they live in the wild.<ref name=hume />
 They form hybrids with some other finches (for example, [[Atlantic canary|canaries]]) giving rise to intermediate birds.<ref>{{cite web|url = http://data.gbif.org/species/browse/taxon/20104120|title = ''Carduelis spinus'' × ''Serinus canaria''|access-date = 13 October 2008|author=Global Biodiversity Information Facility}}</ref><ref>{{cite web|url = http://www.geocities.com/mules_hybrids/siskin_crosses.htm|title = Hibridaciones de ''Carduelis spinus''|access-date = 23 October 2008|archive-url=https://web.archive.org/web/20071106172027/http://www.geocities.com/mules_hybrids/siskin_crosses.htm|archive-date=6 November 2007}}</ref> Hybridisation also occurs in nature without human intervention.{{Failed verification|date=October 2011}}<ref name=Avi>{{cite book| last = McCarthy|first=Eugene M.| publisher=Oxford University Press| title = Handbook of Avian Hybrids of the World| year = 2006| url = https://books.google.com/books?id=iZhKTNkpxUIC| isbn = 978-0-19-518323-8}}</ref> In some areas, individuals that are found are the result of escapes or releases of captive birds.<ref>{{cite book| last = Galarza|first= A.| publisher=Diputación Foral de Bizkaia| title = Urdaibai, avifauna de la ría de Gernika| year = 1989| isbn = 84-404-5084-2}}</ref>
 ===Cultural depictions===
-[[Poland]], [[Gibraltar]], Benin and [[Belgium]] have all issued postage stamps bearing the image of the siskin.<ref name=isr>{{cite web|url = http://israelibirdsstamps.yardbirds.org.il//bl8.html|title = Eurasian Siskin (''Carduelis spinus'')|access-date = 13 October 2008|work = Stamps of Israeli Birds|archive-url = https://web.archive.org/web/20110717002158/http://israelibirdsstamps.yardbirds.org.il/bl8.html|archive-date = 17 July 2011|url-status = dead}}</ref><ref>{{cite web|url = http://www.bird-stamps.org/cspecies/19604700.htm|title = Eurasian Siskin|access-date = 27 November 2008|work=Bird Stamps}}</ref>
+[[Poland]], [[Gibraltar]], [[Benin]] and [[Belgium]] have all issued postage stamps bearing the image of the siskin.<ref name=isr>{{cite web|url = http://israelibirdsstamps.yardbirds.org.il//bl8.html|title = Eurasian Siskin (''Carduelis spinus'')|access-date = 13 October 2008|work = Stamps of Israeli Birds|archive-url = https://web.archive.org/web/20110717002158/http://israelibirdsstamps.yardbirds.org.il/bl8.html|archive-date = 17 July 2011|url-status = dead}}</ref><ref>{{cite web|url = http://www.bird-stamps.org/cspecies/19604700.htm|archive-url = https://web.archive.org/web/20001025032810/http://www.bird-stamps.org/cspecies/19604700.htm|url-status = usurped|archive-date = 25 October 2000|title = Eurasian Siskin|access-date = 27 November 2008|work=Bird Stamps}}</ref>
-In [[Saint Petersburg]] there is a statue of a siskin, as its colours are the same as the uniform worn by the students at an elite school in the city. These students have come to be known by the [[sobriquet]] ''siskin'', {{lang-ru|link=no|Чиж}}. This term was popularised in the Russian song "[[Chizhik-Pyzhik]]".<ref>{{cite web|url = http://www.saint-petersburg.com/monuments/chizhik-pyzhik.asp|title = Chizhik-Pyzhik|access-date = 13 October 2008|publisher=Saint-Petersburg.com}}</ref> There has been a statue of siskin on the embankment by the [[First Engineer Bridge]] since 1994, though it has been stolen and replaced multiple times.
+In [[Saint Petersburg]] there is a statue of a siskin, as its colours are the same as the uniform worn by the students at an elite school in the city. These students have come to be known by the [[sobriquet]] ''siskin'', {{langx|ru|link=no|Чиж}}. This term was popularised in the Russian song "[[Chizhik-Pyzhik]]".<ref>{{cite web|url = http://www.saint-petersburg.com/monuments/chizhik-pyzhik.asp|title = Chizhik-Pyzhik|access-date = 13 October 2008|publisher=Saint-Petersburg.com}}</ref> There has been a statue of a siskin on the embankment by the [[First Engineer Bridge]] since 1994, though it has been stolen and replaced multiple times.
-There is a Czech folk song/dance/game "[[Čížečku, čížečku]]", in which the siskin is the source of the lore on what happens with the [[Poppy (flower)|poppy]].
+There is a Czech folk song/dance/game "[[Čížečku, čížečku]]", in which the siskin is the source of the lore on what happens with the [[poppy]].
 [[Elif Shafak]], in the novel ''Three Daughters of Eve'', mentioned a siskin in a pivotal scene in which the heroine, Peri, meets the charismatic and controversial Professor Azur. When Peri entered Professor Azur's office, she found a siskin with yellow-green feathers and a forked tail trapped amid the shelves and stacks of books.
@@ Line 109: / Line 108: @@
 ==External links==
-{{Commons|Carduelis spinus}}
+{{Commons|Spinus spinus}}
 {{Wikispecies|Carduelis spinus}}
 *[http://www.xeno-canto.org/species/Spinus-spinus Audio recordings from Xeno-canto]
 * [https://web.archive.org/web/20131203001948/http://ibc.lynxeds.com/species/eurasian-siskin-carduelis-spinus Eurasian siskin videos, photos & sounds] on the Internet Bird Collection
 *[https://web.archive.org/web/20141202063639/http://aulaenred.ibercaja.es/wp-content/uploads/432_SiskinCspinus.pdf Ageing and sexing (PDF; 3.1 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze]
-*[http://www.ornithos.de/Ornithos/Feather_Collection/Carduelis_spinus/Carduelis_spinus.htm Feathers of Eurasian siskin (''Carduelis spinus'')]
+*[https://web.archive.org/web/20180304043839/http://www.ornithos.de/Ornithos/Feather_Collection/Carduelis_spinus/Carduelis_spinus.htm Feathers of Eurasian siskin (''Carduelis spinus'')]
 *{{field guide birds of the world|Carduelis spinus}}
 *{{Avibase|name=Carduelis spinus}}
@@ Line 123: / Line 122: @@
 [[Category:Spinus (genus)]]
-[[Category:Birds of Eurasia|siskin]]
+[[Category:Birds of Eurasia]]
 [[Category:Birds described in 1758]]
 [[Category:Taxa named by Carl Linnaeus]]

Taxon identifiers
Spinus spinus	Wikidata: Q27075864 Wikispecies: Spinus spinus ABA: eursis Avibase: 9820CECA12EC737D BirdLife: 22720354 BOLD: 745829 BOW: eursis BTO: bob16540 CoL: 4Z65Y eBird: eursis EoL: 45510356 EUNIS: 1274 GBIF: 6092830 iNaturalist: 145303 ITIS: 997858 IUCN: 22720354 NBN: NHMSYS0021004277 NCBI: 160760 Observation.org: 174 Open Tree of Life: 167338 Xeno-canto: Spinus-spinus
Fringilla spinus	Wikidata: Q28530668 GBIF: 4408957 ZooBank: 4707A019-DF32-42F9-ABF4-F99A6F1873B6