Tyrannasorus rex: Difference between revisions

Tyrannasorus rex; Temporal range: Miocene 20.4–13.8 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Life restoration
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Hybosoridae
Genus:	†Tyrannasorus; Ratcliffe & Ocampo, 2001
Species:	†T. rex
Binomial name
	†Tyrannasorus rex; Ratcliffe & Ocampo, 2001

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Tyrannasorus rex is an extinct species of hybosorid beetle known to exist in the Miocene epoch and the sole member of the monotypic genus Tyrannasorus. A fossilized example scarabaeoid was found embedded in the amber resin of Hymenaea protera in the Dominican Republic. The species was described by Brett C. Ratcliffe and Federico Carlos Ocampo in 2001.^[1]

Holotype

The observed specimen is assumed to be female based on similarities to the female specimens of the genus Apalonychus. She was trapped in the resin produced by Hymenaea protera, now also extinct. The amber was previously dated to Oligocene or Eocene, but these datings are since considered incorrect, and it is accepted that the amber was formed between late Early Miocene and Middle Miocene (15–20 million years ago). The amber came from Dominican Republic, probably from the mountain range north of Santiago de los Caballeros. It is too dark for ventral characteristics of the insect's body to be observed.^[1]

Description

Unlike other hybosorids from the West Indies, whose antennae are 10-segmented, Tyrannasorus rex had nine-segmented antennae. The species is most similar to the genera Coilodes and Apalonychus; their shared characteristics include convex and not quite spherical body and reddish-brown colour. The most prominent difference between Coilodes and Tyrannasorus is in the shape of the antennal club, which is slightly concave in Tyrannasorus but cup-shaped in Coilodes. Furthermore, the former's labrum is wider than the latter's and, unlike the former's, the anterior margin of the latter's pronotum is sinuate. Species of the genus Apalonychus have a much more elongated club of the antenna compared to Tyrannasorus and, unlike Tyrannasorus, eyes of the specimen are subglose and easily visible in dorsal view. Their labrum is also wider than that of Tyrannasorus and the anterior margin of their pronotum is not sinuate. As with other insects, the beetle's six legs consisted of a pair of prolegs, a pair of middle legs and a pair of posterior legs. The insect's elytra have a smooth surface.^[1]

Etymology

It was named after Tyrannosaurus rex, the carnivorous dinosaur, although its discoverers provide an etymology for the name based on the word Hybosorus, the type genus of the family Hybosoridae, to which the species belongs. The stem of the generic name is derived from the Latin tyrannus, meaning "master" or "tyrannical", while the suffix -sorus means "pile" or "hump" in Latin. The name, "tyrannical hump", ultimately refers to the mound of sap in which the observed specimen was fatally trapped.^[1]^[2]

References

^ ^a ^b ^c ^d Ratcliffe, Brett C.; Ocampo, Federico Carlos (2001). "Tyrannasorus rex Ratcliffe and Ocampo, a New Genus and Species of Miocene Hybosorid in Amber from the Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae)". The Coleopterists Bulletin. 55 (3): 351–355. doi:10.1649/0010-065X(2001)055[0351:TRRAOA]2.0.CO;2. JSTOR 4009640. S2CID 10948175.
^ Acorn, John (2007). Deep Alberta: Fossil Facts and Dinosaur Digs. University of Alberta. pp. ^{&#91, page needed&#93,}. ISBN 978-0-88864-481-7.

[RatOca01-1] Ratcliffe, Brett C.; Ocampo, Federico Carlos (2001). "Tyrannasorus rex Ratcliffe and Ocampo, a New Genus and Species of Miocene Hybosorid in Amber from the Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae)". The Coleopterists Bulletin. 55 (3): 351–355. doi:10.1649/0010-065X(2001)055[0351:TRRAOA]2.0.CO;2. JSTOR 4009640. S2CID 10948175.

[Alberta-2] Acorn, John (2007). Deep Alberta: Fossil Facts and Dinosaur Digs. University of Alberta. pp. ^{&#91, page needed&#93,}. ISBN 978-0-88864-481-7.

[1]

[2]

@@ Line 1: / Line 1: @@
+{{Short description|Species of beetle}}
-{{about|the beetle|the dinosaur|Tyrannosaurus}}
+{{About|the beetle|the dinosaur|Tyrannosaurus|the fungus|Tyrannosorus pinicola}}
-{{Taxobox
+{{Speciesbox
-| name =
+| fossil_range = [[Miocene]] {{Fossil range|20.4|13.8}}
-| status =
-| image =
+| image = Tyrannasorus.png
+| image_caption = [[Paleoart|Life restoration]]
-| image_width = 250px
+| parent_authority = Ratcliffe & Ocampo, 2001
-| regnum = [[Animal]]ia
+| taxon = Tyrannasorus rex
-| phylum = [[Arthropod]]a
+| authority = [[Brett C. Ratcliffe|Ratcliffe]] & [[Federico Carlos Ocampo|Ocampo]], 2001
-| classis = [[Insect]]a
-| ordo = [[Beetle|Coleoptera]]
-| familia = [[Scarabaeidae]]
-| subfamilia = [[Hybosoridae]]
-| genus = '''''Tyrannasorus'''''
-| species = '''''T. rex'''''
-| binomial = ''Tyrannasorus rex''
-| binomial_authority = [[Brett C. Ratcliffe|Ratcliffe]] & [[Federico Carlos Ocampo|Ocampo]], 2001
 }}
-'''''Tyrannasorus rex''''' was a species of [[beetle]] known to exist in the [[Miocene]] epoch. A [[fossil]]ized example scarabaeoid was found embedded in the [[amber]] [[resin]] of ''[[Hymenaea protera]]'' in the [[Dominican Republic]]. The species was described by [[Brett C. Ratcliffe]] and [[Federico Carlos Ocampo]] in 2001.<ref>[http://digitalcommons.unl.edu/entomologypapers/46/ ''Tyrannasorus rex'' Raltliffe and Ocampo] (abstract)</ref>
+'''''Tyrannasorus rex''''' is an extinct species of hybosorid [[beetle]] known to exist in the [[Miocene]] epoch and the sole member of the [[monotypic]] genus '''''Tyrannasorus'''''. A [[fossil]]ized example scarabaeoid was found embedded in the [[amber]] [[resin]] of ''[[Hymenaea protera]]'' in the [[Dominican Republic]]. The species was described by [[Brett C. Ratcliffe]] and [[Federico Carlos Ocampo]] in 2001.<ref name="RatOca01"/>
 == Holotype ==
+The observed specimen is assumed to be female based on similarities to the female specimens of the genus ''Apalonychus''. She was trapped in the resin produced by ''[[Hymenaea protera]]'', now also extinct. The [[amber]] was previously dated to [[Oligocene]] or [[Eocene]], but these datings are since considered incorrect, and it is accepted that the amber was formed between late [[Early Miocene]] and [[Middle Miocene]] (15–20 million years ago). The amber came from [[Dominican Republic]], probably from the mountain range north of [[Santiago de los Caballeros]]. It is too dark for ventral characteristics of the insect's body to be observed.<ref name="RatOca01"/>
-The observed specimen is assumed to be female based on similarities to the female specimens of the genus ''Apalonychus''. She was trapped in the resin produced by ''[[Hymenaea protera]]'', now also extinct. The [[amber]] was previously dated to [[Oligocene]] or [[Eocene]], but these datings are now considered wrong, and it is accepted that the amber was formed between late [[Early Miocene]] and [[Middle Miocene]] (15–20 million years ago). The amber came from [[Dominican Republic]], probably from the mountain range north of [[Santiago de los Caballeros]]. It is too dark for ventral characteristics of the insect's body to be observed.<ref name="Nebraska"/>
 == Description ==
+Unlike other hybosorids from the [[West Indies]], whose antennae are 10-segmented, ''Tyrannasorus rex'' had nine-segmented antennae. The species is most similar to the genera ''[[Coilodes]]'' and ''[[Apalonychus]]''; their shared characteristics include convex and not quite spherical body and reddish-brown colour. The most prominent difference between ''Coilodes'' and ''Tyrannasorus'' is in the shape of the antennal club, which is slightly concave in ''Tyrannasorus'' but cup-shaped in ''Coilodes''. Furthermore, the former's [[Labrum (arthropod mouthpart)|labrum]] is wider than the latter's and, unlike the former's, the anterior margin of the latter's [[pronotum]] is sinuate. Species of the genus ''Apalonychus'' have a much more elongated club of the antenna compared to ''Tyrannasorus'' and, unlike ''Tyrannasorus'', eyes of the specimen are subglose and easily visible in dorsal view. Their labrum is also wider than that of ''Tyrannasorus'' and the anterior margin of their pronotum is not sinuate. As with other insects, the beetle's six legs consisted of a pair of prolegs, a pair of middle legs and a pair of posterior legs. The insect's [[elytra]] have a smooth surface.<ref name="RatOca01">{{cite journal|url=http://digitalcommons.unl.edu/entomologypapers/46/|title=''Tyrannasorus rex'' Ratcliffe and Ocampo, a New Genus and Species of Miocene Hybosorid in Amber from the Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae)|date=2001|journal=The Coleopterists Bulletin|volume=55|issue=3|pages=351–355|last1=Ratcliffe|first1=Brett C.|last2=Ocampo|first2=Federico Carlos|doi=10.1649/0010-065X(2001)055[0351:TRRAOA]2.0.CO;2|jstor=4009640|s2cid=10948175 }}</ref>
-Unlike other hybosorids from the [[West Indies]], whose antennae are 10-segmented, ''Tyrannasorus rex'' had a 9-segmented antennae. The species is most similar to the genera ''[[Coilodes]]'' and ''[[Apalonychus]]''; their shared characteristics include convex and not quite spherical body and reddish brown colour. The most prominent difference between ''Coilodes'' and ''Tyrannasorus'' is in the shape of antennal club, which is slightly concave in ''Tyrannosorus'' but cup-shaped in ''Coilodes''. Furthermore, the former's [[Labrum (arthropod mouthpart)|labrum]] is wider than the latter's and, unlike the former's, the anterior margin of the latter's [[pronotum]] is sinuate. Species of the genus ''Apalonychus'' have a much more elongated club of the antenna compared to ''Tyrannasorus'' and, unlike ''Tyrannasorus'', eyes of the specimen are subglose and easily visible in dorsal view. Their labrum is also wider than that of ''Tyrannasorus'' and the anterior margin of their pronotum is not sinuate. As with other insects, the beetle's six legs consisted of a pair of prolegs, a pair of middle legs and a pair of posterior legs. The insect's [[elytra]] have a smooth surface.<ref name="Nebraska">{{cite news|url=http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1045&context=entomologypapers&sei-redir=1&referer=http%3A%2F%2Fwww.google.ba%2Furl%3Fsa%3Dt%26rct%3Dj%26q%3D%2522Tyrannasorus%2Brex%2522%2B%26source%3Dweb%26cd%3D3%26cad%3Drja%26ved%3D0CDUQFjAC%26url%3Dhttp%253A%252F%252Fdigitalcommons.unl.edu%252Fcgi%252Fviewcontent.cgi%253Farticle%253D1045%2526context%253Dentomologypapers%26ei%3DAtACUeGOKbDc4QTZgIHYDw%26usg%3DAFQjCNEUzZE80EHdbf09GLp3gx22ynnWMA%26bvm%3Dbv.41524429%2Cd.Yms#search=%22Tyrannasorus%20rex%22|title=TYRANNASORUS REX RATCLIFFE AND OCAMPO, A NEW GENUS AND SPECIES OF MIOCENE HYBOSORID IN AMBER FROM THE DOMINICAN REPUBLIC (COLEOPTERA: SCARABAEOIDEA: HYBOSORIDAE)|date=3 September 2001|publisher=[[University of Nebraska]]|accessdate=25 January 2013|last1=Ratcliffe|first1=Brett C.|last2=Ocampo|first2=Federico Carlos}}</ref>
 == Etymology ==
+It was named after ''[[Tyrannosaurus|Tyrannosaurus rex]]'', the carnivorous dinosaur, although its discoverers provide an [[etymology]] for the name based on the word ''[[Hybosorus]]'', the [[type genus]] of the [[family (biology)|family]]  [[Hybosoridae]], to which the species belongs. The stem of the generic name is derived from the [[Latin]] ''tyrannus'', meaning "master" or "tyrannical", while the suffix ''-sorus'' means "pile" or "hump" in Latin. The name, "tyrannical hump", ultimately refers to the mound of sap in which the observed specimen was fatally trapped.<ref name="RatOca01"/><ref name="Alberta">{{cite book|title=Deep Alberta: Fossil Facts and Dinosaur Digs|date=2007|isbn=978-0-88864-481-7|publisher=[[University of Alberta]]|last=Acorn|first=John|pages={{Page needed|date=November 2017}}|url-access=registration|url=https://archive.org/details/deepalbertafossi00acor}}</ref>
-It was named after ''[[Tyrannosaurus|Tyrannosaurus rex]]'', the dinosaur, although its discoverers provide an [[etymology]] for the name based on the word ''[[Hybosorus]]'', the genus that formed the basis for the [[subfamily]], known as [[Hybosoridae]], to which the species belongs. The stem of the generic name is derived from the [[Latin]] ''tyrannus'', meaning "master" or "tyrannical", while the suffix ''-sorus'' means "pile" or "hump" in Latin. The name, "tyrannical hump", ultimately refers to the mound of sap in which the observed specimen was fatally trapped.<ref name="Nebraska"/><ref name="Alberta">{{cite books|title=Deep Alberta: Fossil Facts and Dinosaur Digs|date=2007|isbn=0888644817|publisher=[[University of Alberta]]|accessdate=25 January 2013|last=Harrison Acorn|first=John}}</ref>
 == References ==
+{{Wikispecies|Tyrannasorus rex}}
 {{Reflist}}
+{{Taxonbar|from=Q2338326}}
-{{Wikispecies|Tyrannasorus rex}}
 {{DEFAULTSORT:Tyrannasorus Rex}}
-[[Category:Scarabaeidae]]
+[[Category:Scarabaeiformia|†]]
-[[Category:Extinct insects]]
+[[Category:Prehistoric beetles]]
-[[Category:Animals described in 2001]]
+[[Category:Prehistoric insects of the Caribbean]]
+[[Category:Miocene insects]]
+[[Category:Neogene Dominican Republic]]
+[[Category:Dominican amber]]
+[[Category:Fossils of the Dominican Republic]]
+[[Category:Fossil taxa described in 2001]]
+<!-- [[Category:Monotypic animal genera]] moved to genus redirect -->