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{{Short description|Infraorder of crustaceans}}
{{Short description|Infraorder of crustaceans}}
{{Automatic taxobox
{{Automatic taxobox
| fossil_range = [[Late Triassic]]: [[Norian]] - [[Recent]], {{fossilrange|227|0|ref=<ref name="Jerome2011">{{cite journal |last1=Chablais |first1=Jérôme |last2=Feldmann |first2=Rodney M. |last3=Schweitzer |first3=Carrie E. |date=11 August 2010 |title=A new Triassic decapod, ''Platykotta akaina'', from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura |journal=[[Paläontol Z]] |volume=85 |pages=93–102 |issue=2011 |doi=10.1007/s12542-010-0080-y |s2cid=5612385 |url=https://link.springer.com/article/10.1007/s12542-010-0080-y}}</ref>}}
| fossil_range = {{fossil range|Norian|Recent}}
| image = Dardanus megistos2.jpg
| image = Dardanus megistos2.jpg
| image_caption = The [[hermit crab]] ''[[Dardanus megistos]]''
| image_caption = The [[hermit crab]] ''[[Dardanus megistos]]''
| display_parents = 2
| taxon = Anomura
| taxon = Anomura
| authority = [[William Sharp Macleay|Macleay]], 1838
| authority = [[William Sharp Macleay|Macleay]], 1838
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*[[Galatheoidea]]
*[[Galatheoidea]]
*[[Hippoidea]]
*[[Hippoidea]]
*[[Lithodoidea]]
*[[Lomisoidea]]
*[[Lomisoidea]]
*[[Paguroidea]]
*[[Paguroidea]]
}}
}}


'''Anomura''' (sometimes '''Anomala''') is a group of [[Decapoda|decapod]] [[crustacean]]s, including [[hermit crab]]s and others. Although the names of many anomurans include the word ''crab'', all true [[crab]]s are in the [[sister group]] to the Anomura, the [[Crab|Brachyura]] (the two groups together form the [[clade]] [[Meiura]]).<ref>{{cite journal |author1=Gerhard Scholtz |author2=Stefan Richter |year=1995 |title=Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca) |journal=[[Zoological Journal of the Linnean Society]] |volume=113 |issue=3 |pages=289–328 |url=http://arthroinfo.org/pdfs/826/826.pdf |doi=10.1111/j.1096-3642.1995.tb00936.x}}</ref>
'''Anomura''' (sometimes '''Anomala''') is a group of [[decapod]] [[crustacean]]s, including [[hermit crab]]s and others. Although the names of many anomurans include the word ''crab'', all true [[crab]]s are in the [[sister group]] to the Anomura, the [[Brachyura]] (the two groups together form the [[clade]] [[Meiura]]).<ref>{{cite journal |author1=Gerhard Scholtz |author2=Stefan Richter |year=1995 |title=Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca) |journal=[[Zoological Journal of the Linnean Society]] |volume=113 |issue=3 |pages=289–328 |url=http://arthroinfo.org/pdfs/826/826.pdf |doi=10.1111/j.1096-3642.1995.tb00936.x}}</ref>


==Description==
==Description==
The name Anomura derives from an old classification in which [[Reptantia|reptant]] decapods were divided into Macrura (long-tailed), Brachyura (short-tailed) and Anomura (differently-tailed). The alternative name Anomala reflects the unusual variety of forms in this group; whereas all crabs share some obvious similarities, the various groups of anomurans are quite dissimilar.<ref name="Poore">{{cite book |author=Gary Poore |year=2004 |title=Marine Decapod Crustacea of Southern Australia: a Guide to Identification |publisher=[[CSIRO Publishing]] |isbn=978-0-643-09925-8 |chapter=Anomura – hermit crabs, porcelain crabs and squat lobsters |pages=215–287 |chapter-url=https://books.google.com/books?id=TptuZCY3OU0C&pg=PA215}}</ref>
The name Anomura derives from an old classification in which [[Reptantia|reptant]] decapods were divided into Macrura (long-tailed), Brachyura (short-tailed) and Anomura (differently-tailed). The alternative name Anomala reflects the unusual variety of forms in this group; whereas all crabs share some obvious similarities, the various groups of anomurans are quite dissimilar.<ref name="Poore">{{cite book |author=Gary Poore |year=2004 |title=Marine Decapod Crustacea of Southern Australia: a Guide to Identification |publisher=[[CSIRO Publishing]] |isbn=978-0-643-09925-8 |chapter=Anomura – hermit crabs, porcelain crabs and squat lobsters |pages=215–287 |chapter-url=https://books.google.com/books?id=TptuZCY3OU0C&pg=PA215}}</ref>


The group has been moulded by several instances of [[carcinisation]] – the development of a crab-like body form.<ref name="Ahyong">{{cite book |editor1=Joel W. Martin |editor2=Keith A. Crandall|editor2-link=Keith A. Crandall |editor3=Darryl L. Felder |year=2009 |title=Decapod Crustacean Phylogenetics |volume=18 |series=Crustacean issues |publisher=[[CRC Press]] |author1=Shane T. Ahyong |author2=Kareen E. Schnabel |author3=Elizabeth W. Maas |chapter=Anomuran phylogeny: new insights from molecular data |pages=399–414 |chapter-url=https://books.google.com/books?id=Bxs6SgSW2kQC&pg=PA399 |doi=10.1201/9781420092592-c20 |isbn=978-1-4200-9258-5}}</ref> Thus, the [[king crab]]s (Lithodidae), [[porcelain crab]]s (Porcellanidae) and [[hairy stone crab]] (Lomisidae) are all separate instances of carcinisation.<ref name="Ahyong"/>
The group has been moulded by several instances of [[carcinisation]] – the development of a crab-like body form.<ref name="Ahyong">{{cite book |editor1=Joel W. Martin |editor2=Keith A. Crandall|editor2-link=Keith A. Crandall |editor3=Darryl L. Felder |year=2009 |title=Decapod Crustacean Phylogenetics |volume=18 |series=Crustacean issues |publisher=[[CRC Press]] |author1=Shane T. Ahyong |author2=Kareen E. Schnabel |author3=Elizabeth W. Maas |chapter=Anomuran phylogeny: new insights from molecular data |pages=399–414 |chapter-url=https://books.google.com/books?id=Bxs6SgSW2kQC&pg=PA399 |doi=10.1201/9781420092592 |isbn=978-1-4200-9258-5}}</ref> Thus, the [[king crab]]s (Lithodidae), [[porcelain crab]]s (Porcellanidae) and [[hairy stone crab]] (Lomisidae) are all separate instances of carcinisation.<ref name="Ahyong"/>


As decapods (meaning ''ten-legged''), anomurans have ten [[pereiopod]]s, but the last pair of these is reduced in size, and often hidden inside the gill chamber (under the [[carapace]]) to be used for cleaning the gills.<ref>{{cite journal |author1=Jonas Keiler |author2=Stefan Richter |year=2011 |title=Morphological diversity of setae on the grooming legs in Anomala (Decapoda: Reptantia) revealed by scanning electron microscopy |journal=[[Zoologischer Anzeiger]] |volume=250 |issue=4 |pages=343–366 |doi=10.1016/j.jcz.2011.04.004}}</ref><ref name="Poore"/> Since this arrangement is very rare in [[Crab|true crabs]] (for example, the small family [[Hexapodidae]]),<ref name="S&F">{{cite journal |title=Differentiation of the fossil Hexapodidae Miers, 1886 (Decapoda: Brachyura) from similar forms |author1=Carrie E. Schweitzer |author2=Rodney M. Feldmann |journal=[[Journal of Paleontology]] |volume=75 |issue=2 |year=2001 |pages=330–345 |url=http://decapoda.nhm.org/pdfs/10789/10789.pdf|doi=10.1666/0022-3360(2001)075<0330:DOTFHM>2.0.CO;2}}</ref> a "crab" with only eight visible pereiopods is generally an anomuran.<ref name="Poore"/>
As decapods (meaning ''ten-legged''), anomurans have ten [[pereiopod]]s, but the last pair of these is reduced in size, and often hidden inside the gill chamber (under the [[carapace]]) to be used for cleaning the gills.<ref>{{cite journal |author1=Jonas Keiler |author2=Stefan Richter |year=2011 |title=Morphological diversity of setae on the grooming legs in Anomala (Decapoda: Reptantia) revealed by scanning electron microscopy |journal=[[Zoologischer Anzeiger]] |volume=250 |issue=4 |pages=343–366 |doi=10.1016/j.jcz.2011.04.004}}</ref><ref name="Poore"/> Since this arrangement is very rare in [[Crab|true crabs]] (for example, the small family [[Hexapodidae]]),<ref name="S&F">{{cite journal |title=Differentiation of the fossil Hexapodidae Miers, 1886 (Decapoda: Brachyura) from similar forms |author1=Carrie E. Schweitzer |author2=Rodney M. Feldmann |journal=[[Journal of Paleontology]] |volume=75 |issue=2 |year=2001 |pages=330–345 |url=http://decapoda.nhm.org/pdfs/10789/10789.pdf|doi=10.1666/0022-3360(2001)075<0330:DOTFHM>2.0.CO;2|s2cid=85997166 }}</ref> a "crab" with only eight visible pereiopods is generally an anomuran.<ref name="Poore"/>


==Evolution==
==Evolution==
The [[infraorder]] Anomura belongs to the group [[Reptantia]], which consists of the walking/crawling decapods ([[lobsters]] and crabs). There is wide acceptance from [[morphology (biology)|morphological]] and [[molecular phylogenetics|molecular]] data that Anomura and [[Brachyura]] (true crabs) are [[sister taxa]], together making up the [[clade]] Meiura.<ref name="Ahyong"/> The [[cladogram]] below shows Anomura's placement within the larger [[order (biology)|order]] [[Decapoda]], from analysis by Wolfe ''et al.'', 2019.<ref name="Wolfe2019">{{cite journal |last1=Wolfe |first1=Joanna M. |last2=Breinholt |first2=Jesse W. |last3=Crandall |first3=Keith A. |last4=Lemmon |first4=Alan R. |last5=Lemmon |first5=Emily Moriarty |last6=Timm |first6=Laura E. |last7=Siddall |first7=Mark E. |last8=Bracken-Grissom |first8=Heather D. |title=A phylogenomic framework, evolutionary timeline and genomic resources for comparative studies of decapod crustaceans |journal=Proceedings of the Royal Society B | date=24 April 2019 |volume=286 |issue=1901 |doi=10.1098/rspb.2019.0079 |pmid=31014217 |pmc=6501934 |doi-access=free }}</ref>
The [[infraorder]] Anomura belongs to the group [[Reptantia]], which consists of the walking/crawling decapods ([[lobsters]] and crabs). There is wide acceptance from [[morphology (biology)|morphological]] and [[molecular phylogenetics|molecular]] data that Anomura and [[Brachyura]] ("true" crabs) are [[sister taxa]], together making up the [[clade]] Meiura.<ref name=Ahyong/> Anomura likely diverged from Brachyura in the [[Late Triassic]] period, with the earliest discovered Anomuran fossil ''[[Platykotta akaina]]'' dating from the [[Norian]]–[[Rhaetian]] aged [[Ghalilah Formation]] of the [[United Arab Emirates]].<ref name=Jerome2011/>


The [[cladogram]] below shows Anomura's placement within the larger [[order (biology)|order]] [[Decapoda]], from analysis by Wolfe ''et al.'' (2019).<ref name=Wolfe2019>{{cite journal |last1=Wolfe |first1=Joanna M. |last2=Breinholt |first2=Jesse W. |last3=Crandall |first3=Keith A. |last4=Lemmon |first4=Alan R. |last5=Lemmon |first5=Emily Moriarty |last6=Timm |first6=Laura E. |last7=Siddall |first7=Mark E. |last8=Bracken-Grissom |first8=Heather D. |display-authors=6 |date=24 April 2019 |title=A phylogenomic framework, evolutionary timeline and genomic resources for comparative studies of decapod crustaceans |journal=Proceedings of the Royal Society B |volume=286 |issue=1901 |doi=10.1098/rspb.2019.0079 |doi-access=free |pmid=31014217 |pmc=6501934}}</ref>
{{clade| style=font-size:85%; line-height:85%
|label1='''[[Decapoda]]'''
|1={{clade
|1=[[Dendrobranchiata]] (prawns) [[File:Litopenaeus setiferus.png|50 px]]
|label2=[[Pleocyemata]]
|2={{clade
|1={{clade
|1=[[Stenopodidea]] (boxer shrimp) [[File:Spongicola venustus.png|50 px]]
|2={{clade
|1=[[Procarididea]]
|2=[[Caridea]] (true shrimp) [[File:Macrobrachium sp.jpg|50 px]] }} }}
|label2=[[Reptantia]] (crawling/walking decapods)
|2={{clade
|1={{clade
|1=[[Achelata]] (spiny lobsters, slipper lobsters) [[File:Panulirus argus.png|50 px]]
|2={{clade
|1=[[Polychelida]]
|2=[[Astacidea]] (lobsters, crayfish) [[File:Lobster NSRW rotated2.jpg|50 px]] }} }}
|2={{clade
|1=[[Axiidea]] (mud shrimp, ghost shrimp, or burrowing shrimp)
|2={{clade
|1=[[Gebiidea]]
|label2=[[Meiura]]
|2={{clade
|1='''Anomura''' (hermit crabs and others) [[File:Coenobita variabilis.jpg|50 px]]
|2=[[Brachyura]] (crabs) [[File:Charybdis japonica.jpg|50 px]]
}} }} }} }} }} }} }}


{{Decapoda cladogram}}
⚫
Some of the internal relationships within Anomura can be shown in the cladogram below, which shows Hippidae as [[sister taxon|sister]] to Paguroidea, and resolves Parapaguridae outside of Paguroidea:<ref name="Wolfe2019"/>

{{clade| style=font-size:85%; line-height:85%
⚫
Some of the internal relationships within Anomura are shown in the cladogram below, which shows Hippidae as [[sister taxon|sister]] to Paguroidea, and resolves Parapaguridae outside of Paguroidea:<ref name=Wolfe2019/>
|label1='''Anomura'''
{{Anomura cladogram}}
|1={{clade
|1={{clade
|1=[[Porcellanidae]] (porcelain crabs)
|2=[[Munididae]] (squat lobsters) }}
|2={{clade
|1={{clade
|1=[[Parapaguridae]] (deep water sea anemone hermit crabs)
|2=[[Eumunididae]] (squat lobster-like) }}
|2={{clade
|1=[[Hippidae]] (mole crabs or sand crabs)
|label2=[[Paguroidea]]
|2={{clade
|1={{clade
|1=[[Lithodidae]] (king crabs)
|2=[[Paguridae]] (hermit crabs) }}
|2={{clade
|1=[[Diogenidae]] (left-handed hermit crabs)
|2=[[Coenobitidae]] (terrestrial hermit crabs) }} }} }} }} }} }}


==Classification==
==Classification==
The infraorder Anomura contained seven extant superfamilies:<ref name="Grave">{{cite journal |journal = [[Raffles Bulletin of Zoology]] |year=2009 |volume = Suppl. 21 |pages=1–109 |title = A classification of living and fossil genera of decapod crustaceans |author1=Sammy De Grave |author2=N. Dean Pentcheff |author3=Shane T. Ahyong |url=https://decapoda.nhm.org/pdfs/31051/31051.pdf|display-authors=etal}}</ref><ref name="Chablais">{{cite journal |author1=Jérôme Chablais |author2=Rodney M. Feldmann |author3=Carrie E. Schweitzer |year=2011 |title=A new Triassic decapod, ''Platykotta akaina'', from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura |journal=[[Paläontologische Zeitschrift]] |volume=85 |pages=93–102 |doi=10.1007/s12542-010-0080-y |url=http://decapoda.nhm.org/pdfs/31744/31744.pdf }}</ref><ref name="Schnabel">{{cite journal |author1=K. E. Schnabel |author2=S. T. Ahyong |author3=E. W. Maas |year=2011 |title=Galatheoidea are not monophyletic – molecular and morphological phylogeny of the squat lobsters (Decapoda: Anomura) with recognition of a new superfamily |journal=[[Molecular Phylogenetics and Evolution]] |volume=58 |issue=2 |pages=157–168 |doi=10.1016/j.ympev.2010.11.011 |pmid=21095236}}</ref><ref>{{cite WoRMS |author=WoRMS |year=2018 |title=Anomura |id=106671 |access-date=2018-09-28}}</ref>
The infraorder Anomura contained seven extant superfamilies:<ref name="Grave">{{cite journal |journal = [[Raffles Bulletin of Zoology]] |year=2009 |volume = Suppl. 21 |pages=1–109 |title = A classification of living and fossil genera of decapod crustaceans |author1=Sammy De Grave |author2=N. Dean Pentcheff |author3=Shane T. Ahyong |url=https://decapoda.nhm.org/pdfs/31051/31051.pdf|display-authors=etal}}</ref><ref name="Chablais">{{cite journal |author1=Jérôme Chablais |author2=Rodney M. Feldmann |author3=Carrie E. Schweitzer |year=2011 |title=A new Triassic decapod, ''Platykotta akaina'', from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura |journal=[[Paläontologische Zeitschrift]] |volume=85 |issue=1 |pages=93–102 |doi=10.1007/s12542-010-0080-y |bibcode=2011PalZ...85...93C |s2cid=5612385 |url=http://decapoda.nhm.org/pdfs/31744/31744.pdf }}</ref><ref name="Schnabel">{{cite journal |author1=K. E. Schnabel |author2=S. T. Ahyong |author3=E. W. Maas |year=2011 |title=Galatheoidea are not monophyletic – molecular and morphological phylogeny of the squat lobsters (Decapoda: Anomura) with recognition of a new superfamily |journal=[[Molecular Phylogenetics and Evolution]] |volume=58 |issue=2 |pages=157–168 |doi=10.1016/j.ympev.2010.11.011 |pmid=21095236}}</ref><ref>{{cite WoRMS |author=WoRMS |year=2018 |title=Anomura |id=106671 |access-date=2018-09-28}}</ref>
{| class="wikitable"
{| class="wikitable"
! Superfamily !! Members !! Families !! Photo
! Superfamily !! Members !! Families !! Photo
Line 95: Line 52:
| '''[[Hippoidea]]''' || [[mole crab]]s<br />''or'' sand crabs || [[Albuneidae]]<br />[[Blepharipodidae]]<br />[[Hippidae]] || [[File:Blepharipoda.jpg|200px]]<br />''[[Blepharipoda occidentalis]]''<br />([[Blepharipodidae]])
| '''[[Hippoidea]]''' || [[mole crab]]s<br />''or'' sand crabs || [[Albuneidae]]<br />[[Blepharipodidae]]<br />[[Hippidae]] || [[File:Blepharipoda.jpg|200px]]<br />''[[Blepharipoda occidentalis]]''<br />([[Blepharipodidae]])
|-
|-
| '''[[Lithodoidea]]''' || [[king crab]]s || [[Hapalogastridae]]<br />[[Lithodidae]] || [[File:Dessus.lithodes.santolla.JPG|200px]]<br />''[[Lithodes santolla]]''<br />([[King crab|Lithodidae]])
| '''[[Lomis]]oidea''' || [[hairy stone crab]] || [[Lomis]]idae || [[File:Hairy_Stone_Crab_(Lomis_hirta)_(49782727391).jpg|200px]]<br />''[[Lomis hirta]]''<br />(Lomisidae)
|-
| '''[[Lomis]]oidea''' || [[hairy stone crab]] || [[Lomis]]idae ||
|-
|-
| '''[[Hermit crab|Paguroidea]]''' || [[hermit crab]]s<br />[[coconut crab]] || [[Coenobitidae]]<br />[[Diogenidae]]<br />[[Paguridae]]<br />[[Parapaguridae]]<br />[[Parapylochelidae]]<br />[[Pylochelidae]]<br />[[Pylojacquesidae]] || [[File:Caribbean hermit crab.JPG|200px]]<br />''[[Coenobita clypeatus]]''<br />([[Coenobitidae]])
| '''[[Hermit crab|Paguroidea]]''' || [[hermit crab]]s<br />[[coconut crab]] || [[Coenobitidae]]<br />[[Diogenidae]]<br />[[Paguridae]]<br />[[Parapaguridae]]<br />[[Parapylochelidae]]<br />[[Pylochelidae]]<br />[[Pylojacquesidae]] || [[File:Caribbean hermit crab.JPG|200px]]<br />''[[Coenobita clypeatus]]''<br />([[Coenobitidae]])
Line 112: Line 67:
*{{Wikispecies-inline}}
*{{Wikispecies-inline}}


{{Decapoda}}
{{Taxonbar|from=Q526867}}
{{Taxonbar|from=Q526867}}


Latest revision as of 16:09, 19 November 2024

Anomura
Temporal range: Late Triassic: Norian - Recent, 227–0 Ma[1]
The hermit crab Dardanus megistos
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Malacostraca
Order: Decapoda
Suborder: Pleocyemata
(unranked): Reptantia
Infraorder: Anomura
Macleay, 1838
Superfamilies

Anomura (sometimes Anomala) is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word crab, all true crabs are in the sister group to the Anomura, the Brachyura (the two groups together form the clade Meiura).[2]

Description

[edit]

The name Anomura derives from an old classification in which reptant decapods were divided into Macrura (long-tailed), Brachyura (short-tailed) and Anomura (differently-tailed). The alternative name Anomala reflects the unusual variety of forms in this group; whereas all crabs share some obvious similarities, the various groups of anomurans are quite dissimilar.[3]

The group has been moulded by several instances of carcinisation – the development of a crab-like body form.[4] Thus, the king crabs (Lithodidae), porcelain crabs (Porcellanidae) and hairy stone crab (Lomisidae) are all separate instances of carcinisation.[4]

As decapods (meaning ten-legged), anomurans have ten pereiopods, but the last pair of these is reduced in size, and often hidden inside the gill chamber (under the carapace) to be used for cleaning the gills.[5][3] Since this arrangement is very rare in true crabs (for example, the small family Hexapodidae),[6] a "crab" with only eight visible pereiopods is generally an anomuran.[3]

Evolution

[edit]

The infraorder Anomura belongs to the group Reptantia, which consists of the walking/crawling decapods (lobsters and crabs). There is wide acceptance from morphological and molecular data that Anomura and Brachyura ("true" crabs) are sister taxa, together making up the clade Meiura.[4] Anomura likely diverged from Brachyura in the Late Triassic period, with the earliest discovered Anomuran fossil Platykotta akaina dating from the Norian–Rhaetian aged Ghalilah Formation of the United Arab Emirates.[1]

The cladogram below shows Anomura's placement within the larger order Decapoda, from analysis by Wolfe et al. (2019).[7]


 Decapoda 
     

Dendrobranchiata (prawns)

 Pleocyemata 

Stenopodidea (boxer shrimp)

Procarididea

Caridea ("true" shrimp)

 

 Reptantia 

Achelata (spiny lobsters and slipper lobsters)

Polychelida (benthic crustaceans)

Astacidea (lobsters and crayfish)

Axiidea (mud shrimp, ghost shrimp, and burrowing shrimp)

Gebiidea (mud lobsters and mud shrimp)

Anomura (hermit crabs and allies)

Brachyura ("true" crabs)

(crawling / 
walking 
decapods)
 
 


Some of the internal relationships within Anomura are shown in the cladogram below, which shows Hippidae as sister to Paguroidea, and resolves Parapaguridae outside of Paguroidea:[7]

Brachyura ("true" crabs)

Anomura

Porcellanidae (porcelain crabs)

Munididae (squat lobsters)

Parapaguridae (deep water sea anemone hermit crabs)

Eumunididae (squat lobster-like)

Hippidae (mole crabs or sand crabs)

 Paguroidea 

Lithodidae (king crabs)

Paguridae (hermit crabs)

Diogenidae (left-handed hermit crabs)

Coenobitidae (terrestrial hermit crabs)


Classification

[edit]

The infraorder Anomura contained seven extant superfamilies:[8][9][10][11]

Superfamily Members Families Photo
Aegloidea Aegla Aeglidae
Aegla sp.
Chirostyloidea squat lobsters Chirostylidae
Eumunididae
Kiwaidae
Eumunida picta
† Eocarcinoidea † Eocarcinus
† Platykotta		 † Eocarcinidae
† Platykottidae
Galatheoidea squat lobsters
porcelain crabs		 Galatheidae
Munididae
Munidopsidae
Porcellanidae
† Retrorsichelidae
Munidopsis serricornis
(Munidopsidae)
Hippoidea mole crabs
or sand crabs		 Albuneidae
Blepharipodidae
Hippidae
Blepharipoda occidentalis
(Blepharipodidae)
Lomisoidea hairy stone crab Lomisidae
Lomis hirta
(Lomisidae)
Paguroidea hermit crabs
coconut crab		 Coenobitidae
Diogenidae
Paguridae
Parapaguridae
Parapylochelidae
Pylochelidae
Pylojacquesidae
Coenobita clypeatus
(Coenobitidae)

The oldest fossil attributed to Anomura is Platykotta, from the Norian–Rhaetian (Late Triassic) Period in the United Arab Emirates.[9]

References

[edit]
  1. ^ a b Chablais, Jérôme; Feldmann, Rodney M.; Schweitzer, Carrie E. (11 August 2010). "A new Triassic decapod, Platykotta akaina, from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura". Paläontol Z. 85 (2011): 93–102. doi:10.1007/s12542-010-0080-y. S2CID 5612385.
  2. ^ Gerhard Scholtz; Stefan Richter (1995). "Phylogenetic systematics of the reptantian Decapoda (Crustacea, Malacostraca)" (PDF). Zoological Journal of the Linnean Society. 113 (3): 289–328. doi:10.1111/j.1096-3642.1995.tb00936.x.
  3. ^ a b c Gary Poore (2004). "Anomura – hermit crabs, porcelain crabs and squat lobsters". Marine Decapod Crustacea of Southern Australia: a Guide to Identification. CSIRO Publishing. pp. 215–287. ISBN 978-0-643-09925-8.
  4. ^ a b c Shane T. Ahyong; Kareen E. Schnabel; Elizabeth W. Maas (2009). "Anomuran phylogeny: new insights from molecular data". In Joel W. Martin; Keith A. Crandall; Darryl L. Felder (eds.). Decapod Crustacean Phylogenetics. Crustacean issues. Vol. 18. CRC Press. pp. 399–414. doi:10.1201/9781420092592. ISBN 978-1-4200-9258-5.
  5. ^ Jonas Keiler; Stefan Richter (2011). "Morphological diversity of setae on the grooming legs in Anomala (Decapoda: Reptantia) revealed by scanning electron microscopy". Zoologischer Anzeiger. 250 (4): 343–366. doi:10.1016/j.jcz.2011.04.004.
  6. ^ Carrie E. Schweitzer; Rodney M. Feldmann (2001). "Differentiation of the fossil Hexapodidae Miers, 1886 (Decapoda: Brachyura) from similar forms" (PDF). Journal of Paleontology. 75 (2): 330–345. doi:10.1666/0022-3360(2001)075<0330:DOTFHM>2.0.CO;2. S2CID 85997166.
  7. ^ a b Wolfe, Joanna M.; Breinholt, Jesse W.; Crandall, Keith A.; Lemmon, Alan R.; Lemmon, Emily Moriarty; Timm, Laura E.; et al. (24 April 2019). "A phylogenomic framework, evolutionary timeline and genomic resources for comparative studies of decapod crustaceans". Proceedings of the Royal Society B. 286 (1901). doi:10.1098/rspb.2019.0079. PMC 6501934. PMID 31014217.
  8. ^ Sammy De Grave; N. Dean Pentcheff; Shane T. Ahyong; et al. (2009). "A classification of living and fossil genera of decapod crustaceans" (PDF). Raffles Bulletin of Zoology. Suppl. 21: 1–109.
  9. ^ a b Jérôme Chablais; Rodney M. Feldmann; Carrie E. Schweitzer (2011). "A new Triassic decapod, Platykotta akaina, from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura" (PDF). Paläontologische Zeitschrift. 85 (1): 93–102. Bibcode:2011PalZ...85...93C. doi:10.1007/s12542-010-0080-y. S2CID 5612385.
  10. ^ K. E. Schnabel; S. T. Ahyong; E. W. Maas (2011). "Galatheoidea are not monophyletic – molecular and morphological phylogeny of the squat lobsters (Decapoda: Anomura) with recognition of a new superfamily". Molecular Phylogenetics and Evolution. 58 (2): 157–168. doi:10.1016/j.ympev.2010.11.011. PMID 21095236.
  11. ^ WoRMS (2018). "Anomura". WoRMS. World Register of Marine Species. Retrieved 2018-09-28.
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