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| type_species_authority = Perry, 1811
| type_species_authority = Perry, 1811
| subdivision_ranks = Species
| subdivision_ranks = Species
| subdivision = 3-4, ''See text.''
| subdivision = 3–4, ''See text.''
}}
}}
The '''Babirusas''', whose name directly translates to '''Pig Deer''' in Malaysian Malay and English ({{lang-id|babi rusa}}<ref>{{cite book|last=Skeat|first=Walter W.|title=A Concise Etymological Dictionary of the English Language|publisher=Clarendon Press|url=https://archive.org/details/dli.bengal.10689.15458|access-date=17 July 2012|year=1901|page=[https://archive.org/details/dli.bengal.10689.15458/page/n53 35]}}</ref>), are a [[genus]], '''''Babyrousa''''', in the [[Suidae|swine family]] found in the [[Indonesia]]n islands of [[Sulawesi]], [[Togian Islands|Togian]], [[Sula Islands|Sula]] and [[Buru]].<ref name= MSW3>{{MSW3 Artiodactyla | id = 14200008 | page = 637}}</ref> All members of this genus were considered part of a single species until 2002, the babirusa, ''B. babyrussa'', but following that was split into several species. This scientific name is restricted to the [[Buru babirusa]] from Buru and Sula, whereas the best-known species, the [[north Sulawesi babirusa]], is named ''B. celebensis''.<ref name= taxonomy>Meijaard, E., & C. Groves. (2002). ''Proposal for taxonomic changes within the genus Babyrousa.'' Asian Wild Pig News 2 (1), 9-10.</ref> The remarkable "prehistoric" appearance of these mammals is largely due to the prominent upwards incurving canine tusks of the males, which actually pierce the flesh in the snout.<ref name="ADW">{{cite web|url=http://animaldiversity.org/accounts/Babyrousa_babyrussa/|title=''Babyrousa babyrussa''|access-date=2015-12-02|publisher=Animal Diversity Web}}</ref>
The '''babirusas''', also called '''deer-pigs''' ({{langx|id|babi rusa}}<ref>{{cite book|last=Skeat|first=Walter W.|title=A Concise Etymological Dictionary of the English Language|publisher=Clarendon Press|url=https://archive.org/details/dli.bengal.10689.15458|access-date=17 July 2012|year=1901|page=[https://archive.org/details/dli.bengal.10689.15458/page/n53 35]}}</ref>), are a [[genus]], '''''Babyrousa''''', in the [[Suidae|swine family]] found in the [[Indonesia]]n islands of [[Sulawesi]], [[Togian Islands|Togian]], [[Sula Islands|Sula]] and [[Buru]].<ref name= MSW3>{{MSW3 Artiodactyla | id = 14200008 | page = 637}}</ref> All members of this genus were considered part of a single species until 2002, the babirusa, ''B. babyrussa'', but following that was split into several species. This scientific name is restricted to the [[Buru babirusa]] from Buru and Sula, whereas the best-known species, the [[North Sulawesi babirusa]], is named ''B. celebensis''.<ref name= taxonomy>Meijaard, E., & C. Groves. (2002). ''Proposal for taxonomic changes within the genus Babyrousa.'' Asian Wild Pig News 2 (1), 9–10.</ref> The males have prominent upwards incurving canine tusks, which pierce the flesh in the snout.<ref name="ADW">{{cite web|url=http://animaldiversity.org/accounts/Babyrousa_babyrussa/|title=''Babyrousa babyrussa''|access-date=2015-12-02|publisher=Animal Diversity Web}}</ref>

==Etymology==
The first part of the name '''"Babi"''' partially originates of the phrase "Barba Rossa" (Red Bearded) and has been adopted into Malaysian Malay and Malaysian English. "Barba's" phrasing and origin could certainly trace back to the extensive presence of interactions with the Spanish and Portuguese, who similarly named '''Barbados.'''<ref>{{cite web |first1=a |first2=b |first3=c |first4=d |first5=e |title='Barbados' |url=https://web.archive.org/web/20210123091149/https://www.cia.gov/the-world-factbook/countries/barbados/ |website=World Factbook}}</ref>
The second part of the word "Rusa" meaning deer is Malay, is phonetically similar to "Rosa," however it originates from the Proto-Austronesian languages itself.<ref>https://dictionary.cambridge.org/dictionary/english-malaysian/deer</ref>


All species of babirusa are listed as threatened by the [[International Union for Conservation of Nature]] (IUCN).<ref name=RedList2010 />
All species of babirusa are listed as threatened by the [[International Union for Conservation of Nature]] (IUCN).<ref name=RedList2010 />
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The genus is [[monotypic]] within the [[subfamily]] Babyrousinae, or alternatively considered to form a [[Tribe (biology)|tribe]], Babyrousini, of the subfamily [[Suinae]]. To date, only one fossil skull has been found to suggest a larger ancestor.<ref>Ian Metcalfe, ''Faunal and floral migrations and evolution in SE Asia-Australasia''CRC Press, 2001. {{ISBN|978-90-5809-349-3}}. 416 pages: pp338.</ref>
The genus is [[monotypic]] within the [[subfamily]] Babyrousinae, or alternatively considered to form a [[Tribe (biology)|tribe]], Babyrousini, of the subfamily [[Suinae]]. To date, only one fossil skull has been found to suggest a larger ancestor.<ref>Ian Metcalfe, ''Faunal and floral migrations and evolution in SE Asia-Australasia''CRC Press, 2001. {{ISBN|978-90-5809-349-3}}. 416 pages: pp338.</ref>


All members of the genus were considered part of a single species, the babirusa or pig-deer, ''B. babyrussa''. They were split into several species, this scientific name is restricted to the Buru babirusa from Buru and the Sula Islands, whereas the best-known species, the north Sulawesi babirusa, is named ''B. celebensis''.<ref name= taxonomy /> The split, which uses the [[phylogenetic species concept]], is based on differences in size, amount of hair on body and tail-tuft, and measurements of the skull and teeth.<ref name= taxonomy />
All members of the genus were considered part of a single species, the babirusa or pig-deer, ''B. babyrussa''. After they were split into several species, this scientific name is restricted to the Buru babirusa from Buru and the Sula Islands, whereas the best-known species, the north Sulawesi babirusa, is named ''B. celebensis''.<ref name= taxonomy /> The split, which uses the [[phylogenetic species concept]], is based on differences in size, amount of hair on the body and tail-tuft, and measurements of the skull and teeth.<ref name= taxonomy />


=== Species ===
=== Species ===
{| class="wikitable"
* [[Buru babirusa]] (''B. babyrussa''), also known as the hairy or golden babirusa
|-
* [[Bola Batu babirusa]] (''B. bolabatuensis''), proposed species
! Image !! Name !! Distribution
* [[North Sulawesi babirusa]] (''B. celebensis''), also known as the Sulawesi babirusa
|-
* [[Togian babirusa]] (''B. togeanensis'')
| || [[Buru babirusa]] (''B. babyrussa''), also known as the hairy or golden babirusa || Sula Islands of Mangole and Taliabu.
|-
| || [[Bola Batu babirusa]] (''B. bolabatuensis'') || Sulawesi.
|-
|[[File:Babirusa (29411754088).jpg|120px]] ||[[North Sulawesi babirusa]] (''B. celebensis''), also known as the Sulawesi babirusa ||Sulawesi
|-
|[[File:Babyrousa togeanensis.jpg|120px]] || [[Togian babirusa]] (''B. togeanensis'') ||Togian Islands
|-
|}


''B. babyrussa beruensis'' was described as an extinct, [[Pleistocene]] subspecies from southwestern Sulawesi before babirusas were split into multiple species.
''B. babyrussa beruensis'' was described as an extinct, [[Pleistocene]] subspecies from southwestern Sulawesi before babirusas were split into multiple species.


== Description ==
== Description ==
[[File:Babyrousa celebensis - Crane.jpg|thumb|left|Skull of a male North Sulawesi babirusa (37&nbsp;cm long).]]
[[File:Babyrousa celebensis - Crane.jpg|thumb|left|Skull of a male North Sulawesi babirusa ({{convert|37|cm|in|abbr=in|disp=or}} long).]]
Babirusa are notable for the long upper canines in the males. The upper canines of male babirusa emerge vertically from the [[alveolar process]], penetrating through the skin and curving backward over the front of the face and towards the forehead.<ref name=MacDonald1993>MacDonald, A. A. 1993. The Babirusa (Babyrousa babyrussa). In: W. L. R. Oliver (ed.), Pigs, Peccaries, and Hippos: Status Survey and Conservation Action Plan, IUCN, Gland, Switzerland.</ref><ref>{{Cite journal|last=Nasoori|first=A|year=2020|title=Tusks, the extra-oral teeth |journal=Archives of Oral Biology|volume=117|page=104835|doi=10.1016/j.archoralbio.2020.104835|pmid=32668361|s2cid=220585014}}</ref> The lower canines also grow upwards. The canines of the female are either reduced or absent.<ref name=MacDonald1993 /> The structure of the male's canines vary by species. In the golden babirusa, the upper canines are short and slender with the alveolar rotated forward to allow the lower canines to cross the lateral view.<ref name=MacDonald1993 /> The Togian babirusa also has the same characteristics and the upper canines always converge. The North Sulawesi babirusa has long and thick upper canines with a vertically implanted alveolar. This caused the upper canines to emerge vertically and not cross with the lower canines.<ref>Groves, C. 1980. Notes on the systematics of Babyrousa (Artiodactyla, Suidae). Zoologische Mededelingen 55:29–46.</ref> Babirusa also vary by species in other characteristics. The golden babirusa has a long, thick pelage that is white, creamy gold, black or gold overall and black at the rump.<ref name=MacDonald1993 /> The pelage of the Togian babirusa is also long but not as that of the golden babirusa. The Togian babirusa has a tawny, brown or black pelage that is darker on the upper parts than in the lower parts.<ref name=MacDonald1993 /> The North Sulawesi babirusa has very short hair and appears bald. The female babirusa has only one pair of [[teat]]s.<ref name=BBC>[[BBC]] (2010). ''[http://www.bbc.co.uk/nature/wildfacts/factfiles/597.shtml Babirusa.]'' Downloaded 18 April 2010.</ref>
Babirusas are notable for the long upper canines in the males. The upper canines of males emerge vertically from the [[alveolar process]], penetrating through the skin and curving backward over the front of the face and towards the forehead.<ref name=MacDonald1993>MacDonald, A. A. 1993. The Babirusa (Babyrousa babyrussa). In: W. L. R. Oliver (ed.), Pigs, Peccaries, and Hippos: Status Survey and Conservation Action Plan, IUCN, Gland, Switzerland.</ref><ref>{{Cite journal|last=Nasoori|first=A|year=2020|title=Tusks, the extra-oral teeth |journal=Archives of Oral Biology|volume=117|page=104835|doi=10.1016/j.archoralbio.2020.104835|pmid=32668361|s2cid=220585014}}</ref> The lower canines also grow upwards. The canines of females are either reduced or absent.<ref name=MacDonald1993 /> The structure of the male's canines varies by species. In the golden babirusa, the upper canines are short and slender with the alveolar rotated forward to allow the lower canines to cross the lateral view.<ref name=MacDonald1993 /> The Togian babirusa also has the same characteristics and the upper canines always converge. The North Sulawesi babirusa has long and thick upper canines with a vertically implanted alveolar. This causes the upper canines to emerge vertically and not cross with the lower canines.<ref>Groves, C. 1980. Notes on the systematics of Babyrousa (Artiodactyla, Suidae). Zoologische Mededelingen 55:29–46.</ref>
Babirusas also vary by species in other characteristics. The golden babirusa has a long, thick pelage that is white, creamy gold, black or gold overall, and black at the rump.<ref name=MacDonald1993 /> The pelage of the Togian babirusa is also long but not as that of the golden babirusa. The Togian babirusa has a tawny, brown, or black pelage that is darker on the upper parts than in the lower parts.<ref name=MacDonald1993 /> The North Sulawesi babirusa has very short hair and appears bald. The female babirusa has only one pair of [[teat]]s.<ref name=BBC>[[BBC]] (2010). ''[http://www.bbc.co.uk/nature/wildfacts/factfiles/597.shtml Babirusa.]'' Downloaded 18 April 2010.</ref>


== Biology and ecology ==
== Biology and ecology ==
[[File:Babirusa berkelahi - edited.jpg|thumb|Two babirusas fighting in Indonesia.]]
[[File:Babirusa berkelahi - edited.jpg|thumb|Two babirusas fighting in Indonesia.]]
[[File:Babirusa Skeleton.jpg|thumb|[[North Sulawesi babirusa]] skeleton ([[Museum of Osteology]])]]
[[File:Babirusa Skeleton.jpg|thumb|[[North Sulawesi babirusa]] skeleton ([[Museum of Osteology]])]]
Babirusa are native to [[Sulawesi]], some of the [[Togian Islands]], the [[Sula Islands Regency|Sula Islands]] and [[Buru]].<ref name=MacDonald1993 /> In Sulawesi, they range from the [[Minahasa Peninsula|northern peninsula]] to the [[South Sulawesi|south]] and [[Southeast Sulawesi|south east]] provinces. Although babirusas are present on both Sulawesi and Sula, they are not found on the large islands between the two, the [[Banggai Regency|Banggai Archipelago]]. It has been hypothesized that the unusual distribution is due to their being transported by humans as gifts bestowed by native royalty.<ref name=Umberto>Umberto Albarella, [[Keith Dobney]], Anton Ervynck: 2007. ''Pigs and humans: 10,000 years of interaction'' Oxford University Press 2007. {{ISBN|978-0-19-920704-6}}. 454 pages: pp20.</ref> The preferred habitat of babirusa are tropical rainforests along river banks.<ref name=MacDonald1993 /> It appears that they have been confined to the higher grounds in the interior despite occurring in lowland areas near coasts in the past.<ref name=MacDonald1993 /> They are also active during the daytime. Like all pig species, babirusa have an omnivorous diet with an intestinal tract similar to that of the domestic pig.<ref>Langer, P. (1988): The Mammalian Herbivore Stomach – Comparative Anatomy, Function and Evolution. Gustav Fischer, Stuttgart and New York: 136-161.</ref> The stomach [[diverticulum]] of a babirusa is enlarged which may indicate that it is a [[ruminant]] but evidence shows otherwise.<ref name=MacDonald1993 /> Because it does not have a rostral bone in the nose, a babirusa does not dig with its snout like other pigs do except in mud and swampy grounds. The diet of the babirusa includes leaves, roots, fruits and animal material. Apparently, the strong jaws of a babirusa are capable of easily cracking hard nuts.<ref name=MacDonald1993 />
Babirusas are native to [[Sulawesi]], some of the [[Togian Islands]], the [[Sula Islands]], and [[Buru]].<ref name=MacDonald1993 /> In Sulawesi, they range from the [[Minahasa Peninsula]] to the provinces of [[South Sulawesi]] and [[Southeast Sulawesi]]. Although they are present on both Sulawesi and Sula, they are not found on the large islands between the two, the [[Banggai Islands Regency|Banggai Archipelago]]. It has been hypothesized that the unusual distribution may be due to their being transported by humans as gifts bestowed by native royalty.<ref name=Umberto>Umberto Albarella, [[Keith Dobney]], Anton Ervynck: 2007. ''Pigs and humans: 10,000 years of interaction'' Oxford University Press 2007. {{ISBN|978-0-19-920704-6}}. 454 pages: pp20.</ref> The preferred habitat of babirusa is tropical rainforest along river banks.<ref name=MacDonald1993 /> It appears that they have been confined to the higher grounds in the interior despite occurring in lowland areas near coasts in the past.<ref name=MacDonald1993 /> They are active during the daytime. Like all pig species, babirusa has an omnivorous diet with an intestinal tract similar to that of the domestic pig.<ref>Langer, P. (1988): The Mammalian Herbivore Stomach – Comparative Anatomy, Function and Evolution. Gustav Fischer, Stuttgart and New York: 136–161.</ref> The stomach [[diverticulum]] of a babirusa is enlarged which may indicate that it is a [[ruminant]] but evidence shows otherwise.<ref name=MacDonald1993 /> Because it does not have a rostral bone in the nose, a babirusa does not dig with its snout like other pigs do except in mud and swampy grounds. The diet of the babirusa includes leaves, roots, fruits and animal material. The strong jaws of a babirusa are capable of easily cracking hard nuts.<ref name=MacDonald1993 />


Male babirusa tend to live solitarily while adult females can be found in groups with young.<ref name=Patry>Patry, Maurice; Leus, Kristin; Macdonald, Alastair A (1995) "Group Structure and Behaviour of Babirusa (''Babyrousa babyrussa'') in Northern Sulawesi", Australian Journal of ''Zoology'', 43:643-655.</ref> Groups of female and young babirusa may number up to 84 individuals, most of which contain no adult males. Males rarely travel in pairs or trios.<ref name=Patry /> There are almost never more than three adult females in a group.<ref>Patry M , Leus K Macdonald AA (1995) Group structure and behaviour of babirusa (''Babyrousa babyrussa) in northern Sulawesi. Australian Journal of Zoology 43, 643-655.</ref> The tusks of the adult males are used in intraspecific fighting. The upper tusks are for defense while the lower tusks are offensive weapons.<ref>J. MacKinnon, (1981) "The structure and function of the tusks of babirusa", ''Mammal Review'', 11(1):37-40.</ref> If a male babirusa does not grind his tusks (achievable through regular activity), they can eventually keep growing so as to penetrate the individual's own skull.<!-- at which time, does it kill them? --><ref>{{cite web|url=http://scienceblogs.com/tetrapodzoology/2010/03/08/babirusa-impales-own-head/|title=Babirusas can get impaled by their own teeth: that most sought-after of objects does exist! (babirusas, part VIII)|work=Tetrapod Zoology}}</ref>
Males tend to live solitarily while adult females can be found in groups with young.<ref name=Patry>Patry, Maurice; Leus, Kristin; Macdonald, Alastair A (1995) "Group Structure and Behaviour of Babirusa (''Babyrousa babyrussa'') in Northern Sulawesi", Australian Journal of ''Zoology'', 43:643–655.</ref> Groups of females and young may number up to 84 individuals, most of which contain no adult males. Males rarely travel in pairs or trios.<ref name=Patry /> There are almost never more than three adult females in a group.<ref>Patry M , Leus K Macdonald AA (1995) Group structure and behaviour of babirusa (''Babyrousa babyrussa) in northern Sulawesi. Australian Journal of Zoology 43, 643–655.</ref> The tusks of the adult males are used in intraspecific fighting. The upper tusks are for defense while the lower tusks are offensive weapons.<ref>J. MacKinnon, (1981) "The structure and function of the tusks of babirusa", ''Mammal Review'', 11(1):37-40.</ref> If a male babirusa does not grind his tusks (achievable through regular activity), they can eventually keep growing and, rarely, penetrate the individual's skull.<!-- at which time, does it kill them? --><ref>{{cite web|url=http://scienceblogs.com/tetrapodzoology/2010/03/08/babirusa-impales-own-head/|title=Babirusas can get impaled by their own teeth: that most sought-after of objects does exist! (babirusas, part VIII)|work=Tetrapod Zoology}}</ref>


Female babirusa cycle lengths are between 28 and 42 days and estrus last 2–3 days.<ref name=MacDonald1993 /> The litter size for a babirusa is usually one or two piglets.<ref name=MacDonald1993 />
Female babirusa cycle lengths are between 28 and 42 days and estrus last 2–3 days.<ref name=MacDonald1993 /> The litter size for a babirusa is usually one or two piglets.<ref name=MacDonald1993 />
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The [[Bali]]nese Hindu-era Court of Justice pavilion and the "floating pavilion" of [[Klungkung]] palace ruins are notable for painted babirusa ''[[Rakshasa|raksasa]]'' ([[grotesque]]s) on the ceilings.<ref name=Umberto />
The [[Bali]]nese Hindu-era Court of Justice pavilion and the "floating pavilion" of [[Klungkung]] palace ruins are notable for painted babirusa ''[[Rakshasa|raksasa]]'' ([[grotesque]]s) on the ceilings.<ref name=Umberto />


Prehistoric paintings of babirusa found in caves on the island of Sulawesi in Indonesia have been dated back at least 35,400 years (to the ice age Pleistocene epoch). [[Adam Brumm]], who co-authored the 2014 study dating the paintings, said "The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves."<ref>{{cite news|last1=Sample|first1=Ian|title=35,000-year-old Indonesian cave paintings suggest art came out of Africa|url=https://www.theguardian.com/science/2014/oct/08/cave-art-indonesia-sulawesi|access-date=14 November 2015|work=The Guardian|date=9 October 2014|quote="The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves", said Brumm.}}</ref>
Prehistoric paintings of babirusa found in caves on the island of Sulawesi in Indonesia have been dated back at least 35,400 years (to the ice age Pleistocene epoch). {{ill|Adam Brumm|wd=Q88329455|s=1|v=sup}}, who co-authored the 2014 study dating the paintings, said "The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves."<ref>{{cite news|last1=Sample|first1=Ian|title=35,000-year-old Indonesian cave paintings suggest art came out of Africa|url=https://www.theguardian.com/science/2014/oct/08/cave-art-indonesia-sulawesi|access-date=14 November 2015|work=The Guardian|date=9 October 2014|quote="The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves", said Brumm.}}</ref>

The babirusa has sparked debate among Jewish scholars and animal researchers about whether it is considered [[Kosher foods|kosher]], or permissible to be consumed by Jews, according to [[Kashrut|Jewish dietary laws]]. The debate centers around whether the animal chews its [[cud]], which is a requirement according to the [[Old Testament]] for an animal to be considered kosher. Some experts, like [[J. David Bleich]], a professor of [[Halakha|Jewish law]] and ethics at [[Yeshiva University]], believe that the babirusa does not meet the physical criteria to be considered kosher, challenging the assertion that the babirusa chews its cud by citing a report from 1940 that found that true rumination could not take place in the animal's stomach. However, he also notes that Jews can eat any food that is not expressly forbidden and that "the babirusa's resemblance to a pig in appearance and taste is not sufficient grounds for banning its consumption as kosher meat." Others, such as Fuller Bazer, an animal science professor at the University of Florida, believe that the animal is kosher due to its [[cloven hoof]] and cud chewing. Additionally, it has been noted that the babirusa is an endangered species and that most Muslims, who face similar dietary restrictions, would avoid eating the meat of any animal whose status in religious law is uncertain.<ref>{{Cite web |last=Dart |first=John |date=1985-02-19 |title=This Little Piggy Not Likely to Be on Kosher Menus |url=https://www.latimes.com/archives/la-xpm-1985-02-19-mn-376-story.html |access-date=2023-01-24 |website=Los Angeles Times |language=en-US}}</ref><ref>{{Cite web |title=Scholar Questions Whether The Babirusa Chews Its Cud |url=https://apnews.com/article/9cc9d764daef8fbae8f78b6a7c50f61e |access-date=2023-01-24 |website=AP NEWS |language=en}}</ref>


== Conservation status ==
== Conservation status ==
Babirusas are protected in Indonesia and killing them is illegal in most cases. However, poaching remains a significant threat to the babirusa. Additionally, commercial logging operations threaten the babirusa by [[habitat loss]], and also reduce cover, making the babirusa more exposed to poachers.<ref name=ARKive /> All [[extant taxon|extant]] species of babirusa are listed as [[Vulnerable species|vulnerable]] or [[Endangered species|endangered]] by the [[IUCN]].<ref name=RedList2010>{{cite web |url=http://www.iucnredlist.org/search |title=IUCN Red List of Threatened Species Version 2010.1 |year=2010 |access-date= 18 April 2010}}</ref>
Babirusas are protected in Indonesia and killing them is illegal in most cases. However, [[poaching]] remains a significant threat to the babirusa. Additionally, commercial logging operations threaten the babirusa by [[habitat loss]], and also reduce cover, making the babirusa more exposed to poachers.<ref name=ARKive /> All [[extant taxon|extant]] species of babirusa are listed as [[Vulnerable species|vulnerable]] or [[Endangered species|endangered]] by the [[IUCN]].<ref name=RedList2010>{{cite web |url=http://www.iucnredlist.org/search |title=IUCN Red List of Threatened Species Version 2010.1 |year=2010 |access-date= 18 April 2010}}</ref>


== References ==
== References ==
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[[Category:Suidae]]
[[Category:Suidae]]
[[Category:Wallacea]]
[[Category:Wallacea]]
[[Category:Indonesian words and phrases]]
[[Category:Endemic fauna of Indonesia]]
[[Category:Endemic fauna of Indonesia]]
[[Category:Mammals of Indonesia]]
[[Category:Mammals of Indonesia]]

Latest revision as of 07:16, 24 November 2024

Babirusa
A male North Sulawesi babirusa; only the adult males possess the distinctive tusks.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Suidae
Subfamily: Babyrousinae
Thenius, 1970
Genus: Babyrousa
Perry, 1811
Type species
Babyrousa quadricornua [1]
Perry, 1811
Species

3–4, See text.

The babirusas, also called deer-pigs (Indonesian: babi rusa[2]), are a genus, Babyrousa, in the swine family found in the Indonesian islands of Sulawesi, Togian, Sula and Buru.[3] All members of this genus were considered part of a single species until 2002, the babirusa, B. babyrussa, but following that was split into several species. This scientific name is restricted to the Buru babirusa from Buru and Sula, whereas the best-known species, the North Sulawesi babirusa, is named B. celebensis.[4] The males have prominent upwards incurving canine tusks, which pierce the flesh in the snout.[5]

All species of babirusa are listed as threatened by the International Union for Conservation of Nature (IUCN).[6]

Classification

[edit]

The genus is monotypic within the subfamily Babyrousinae, or alternatively considered to form a tribe, Babyrousini, of the subfamily Suinae. To date, only one fossil skull has been found to suggest a larger ancestor.[7]

All members of the genus were considered part of a single species, the babirusa or pig-deer, B. babyrussa. After they were split into several species, this scientific name is restricted to the Buru babirusa from Buru and the Sula Islands, whereas the best-known species, the north Sulawesi babirusa, is named B. celebensis.[4] The split, which uses the phylogenetic species concept, is based on differences in size, amount of hair on the body and tail-tuft, and measurements of the skull and teeth.[4]

Species

[edit]
Image Name Distribution
Buru babirusa (B. babyrussa), also known as the hairy or golden babirusa Sula Islands of Mangole and Taliabu.
Bola Batu babirusa (B. bolabatuensis) Sulawesi.
North Sulawesi babirusa (B. celebensis), also known as the Sulawesi babirusa Sulawesi
Togian babirusa (B. togeanensis) Togian Islands

B. babyrussa beruensis was described as an extinct, Pleistocene subspecies from southwestern Sulawesi before babirusas were split into multiple species.

Description

[edit]
Skull of a male North Sulawesi babirusa (37 cm or 15 inches long).

Babirusas are notable for the long upper canines in the males. The upper canines of males emerge vertically from the alveolar process, penetrating through the skin and curving backward over the front of the face and towards the forehead.[8][9] The lower canines also grow upwards. The canines of females are either reduced or absent.[8] The structure of the male's canines varies by species. In the golden babirusa, the upper canines are short and slender with the alveolar rotated forward to allow the lower canines to cross the lateral view.[8] The Togian babirusa also has the same characteristics and the upper canines always converge. The North Sulawesi babirusa has long and thick upper canines with a vertically implanted alveolar. This causes the upper canines to emerge vertically and not cross with the lower canines.[10]

Babirusas also vary by species in other characteristics. The golden babirusa has a long, thick pelage that is white, creamy gold, black or gold overall, and black at the rump.[8] The pelage of the Togian babirusa is also long but not as that of the golden babirusa. The Togian babirusa has a tawny, brown, or black pelage that is darker on the upper parts than in the lower parts.[8] The North Sulawesi babirusa has very short hair and appears bald. The female babirusa has only one pair of teats.[11]

Biology and ecology

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Two babirusas fighting in Indonesia.
North Sulawesi babirusa skeleton (Museum of Osteology)

Babirusas are native to Sulawesi, some of the Togian Islands, the Sula Islands, and Buru.[8] In Sulawesi, they range from the Minahasa Peninsula to the provinces of South Sulawesi and Southeast Sulawesi. Although they are present on both Sulawesi and Sula, they are not found on the large islands between the two, the Banggai Archipelago. It has been hypothesized that the unusual distribution may be due to their being transported by humans as gifts bestowed by native royalty.[12] The preferred habitat of babirusa is tropical rainforest along river banks.[8] It appears that they have been confined to the higher grounds in the interior despite occurring in lowland areas near coasts in the past.[8] They are active during the daytime. Like all pig species, babirusa has an omnivorous diet with an intestinal tract similar to that of the domestic pig.[13] The stomach diverticulum of a babirusa is enlarged which may indicate that it is a ruminant but evidence shows otherwise.[8] Because it does not have a rostral bone in the nose, a babirusa does not dig with its snout like other pigs do except in mud and swampy grounds. The diet of the babirusa includes leaves, roots, fruits and animal material. The strong jaws of a babirusa are capable of easily cracking hard nuts.[8]

Males tend to live solitarily while adult females can be found in groups with young.[14] Groups of females and young may number up to 84 individuals, most of which contain no adult males. Males rarely travel in pairs or trios.[14] There are almost never more than three adult females in a group.[15] The tusks of the adult males are used in intraspecific fighting. The upper tusks are for defense while the lower tusks are offensive weapons.[16] If a male babirusa does not grind his tusks (achievable through regular activity), they can eventually keep growing and, rarely, penetrate the individual's skull.[17]

Female babirusa cycle lengths are between 28 and 42 days and estrus last 2–3 days.[8] The litter size for a babirusa is usually one or two piglets.[8]

Relationship with humans

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In Indonesia, the striking appearance of the babirusa has inspired demonic masks.[18]

The Balinese Hindu-era Court of Justice pavilion and the "floating pavilion" of Klungkung palace ruins are notable for painted babirusa raksasa (grotesques) on the ceilings.[12]

Prehistoric paintings of babirusa found in caves on the island of Sulawesi in Indonesia have been dated back at least 35,400 years (to the ice age Pleistocene epoch). Adam Brumm [d], who co-authored the 2014 study dating the paintings, said "The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves."[19]

The babirusa has sparked debate among Jewish scholars and animal researchers about whether it is considered kosher, or permissible to be consumed by Jews, according to Jewish dietary laws. The debate centers around whether the animal chews its cud, which is a requirement according to the Old Testament for an animal to be considered kosher. Some experts, like J. David Bleich, a professor of Jewish law and ethics at Yeshiva University, believe that the babirusa does not meet the physical criteria to be considered kosher, challenging the assertion that the babirusa chews its cud by citing a report from 1940 that found that true rumination could not take place in the animal's stomach. However, he also notes that Jews can eat any food that is not expressly forbidden and that "the babirusa's resemblance to a pig in appearance and taste is not sufficient grounds for banning its consumption as kosher meat." Others, such as Fuller Bazer, an animal science professor at the University of Florida, believe that the animal is kosher due to its cloven hoof and cud chewing. Additionally, it has been noted that the babirusa is an endangered species and that most Muslims, who face similar dietary restrictions, would avoid eating the meat of any animal whose status in religious law is uncertain.[20][21]

Conservation status

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Babirusas are protected in Indonesia and killing them is illegal in most cases. However, poaching remains a significant threat to the babirusa. Additionally, commercial logging operations threaten the babirusa by habitat loss, and also reduce cover, making the babirusa more exposed to poachers.[18] All extant species of babirusa are listed as vulnerable or endangered by the IUCN.[6]

References

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  1. ^ Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. ^ Skeat, Walter W. (1901). A Concise Etymological Dictionary of the English Language. Clarendon Press. p. 35. Retrieved 17 July 2012.
  3. ^ Grubb, P. (2005). "Order Artiodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 637. ISBN 978-0-8018-8221-0. OCLC 62265494.
  4. ^ a b c Meijaard, E., & C. Groves. (2002). Proposal for taxonomic changes within the genus Babyrousa. Asian Wild Pig News 2 (1), 9–10.
  5. ^ "Babyrousa babyrussa". Animal Diversity Web. Retrieved 2 December 2015.
  6. ^ a b "IUCN Red List of Threatened Species Version 2010.1". 2010. Retrieved 18 April 2010.
  7. ^ Ian Metcalfe, Faunal and floral migrations and evolution in SE Asia-AustralasiaCRC Press, 2001. ISBN 978-90-5809-349-3. 416 pages: pp338.
  8. ^ a b c d e f g h i j k l MacDonald, A. A. 1993. The Babirusa (Babyrousa babyrussa). In: W. L. R. Oliver (ed.), Pigs, Peccaries, and Hippos: Status Survey and Conservation Action Plan, IUCN, Gland, Switzerland.
  9. ^ Nasoori, A (2020). "Tusks, the extra-oral teeth". Archives of Oral Biology. 117: 104835. doi:10.1016/j.archoralbio.2020.104835. PMID 32668361. S2CID 220585014.
  10. ^ Groves, C. 1980. Notes on the systematics of Babyrousa (Artiodactyla, Suidae). Zoologische Mededelingen 55:29–46.
  11. ^ BBC (2010). Babirusa. Downloaded 18 April 2010.
  12. ^ a b Umberto Albarella, Keith Dobney, Anton Ervynck: 2007. Pigs and humans: 10,000 years of interaction Oxford University Press 2007. ISBN 978-0-19-920704-6. 454 pages: pp20.
  13. ^ Langer, P. (1988): The Mammalian Herbivore Stomach – Comparative Anatomy, Function and Evolution. Gustav Fischer, Stuttgart and New York: 136–161.
  14. ^ a b Patry, Maurice; Leus, Kristin; Macdonald, Alastair A (1995) "Group Structure and Behaviour of Babirusa (Babyrousa babyrussa) in Northern Sulawesi", Australian Journal of Zoology, 43:643–655.
  15. ^ Patry M , Leus K Macdonald AA (1995) Group structure and behaviour of babirusa (Babyrousa babyrussa) in northern Sulawesi. Australian Journal of Zoology 43, 643–655.
  16. ^ J. MacKinnon, (1981) "The structure and function of the tusks of babirusa", Mammal Review, 11(1):37-40.
  17. ^ "Babirusas can get impaled by their own teeth: that most sought-after of objects does exist! (babirusas, part VIII)". Tetrapod Zoology.
  18. ^ a b "Sulawesi babirusa (Babyrousa celebensis)". ARKive. Archived from the original on 11 April 2012. Retrieved 17 July 2012.
  19. ^ Sample, Ian (9 October 2014). "35,000-year-old Indonesian cave paintings suggest art came out of Africa". The Guardian. Retrieved 14 November 2015. "The paintings of the wild animals are most fascinating because it is clear they were of particular interest to the artists themselves", said Brumm.
  20. ^ Dart, John (19 February 1985). "This Little Piggy Not Likely to Be on Kosher Menus". Los Angeles Times. Retrieved 24 January 2023.
  21. ^ "Scholar Questions Whether The Babirusa Chews Its Cud". AP NEWS. Retrieved 24 January 2023.
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