Jump to content

Dinosaur size: Difference between revisions

From Wikipedia, the free encyclopedia
Content deleted Content added
No edit summary
Tag: Reverted
Minor typographical errors
 
(20 intermediate revisions by 14 users not shown)
Line 1: Line 1:
{{pp|small=yes}}
{{Short description|Dinosaur mass and length estimates}}
{{Short description|Dinosaur mass and length estimates}}
{{pp|small=yes}}
{{cleanup|reason=Outdated information and unreliable overestimates|date = August 2022}}
{{cleanup|reason=Outdated information and unreliable overestimates|date = August 2022}}
[[File:Longest dinosaur by clade.svg|alt=|thumb|Scale diagram comparing a human and the longest-known dinosaurs of five major clades]]
[[File:Longest dinosaur by clade.svg|alt=|thumb|Scale diagram comparing a human and the longest-known dinosaurs of five major clades]]
[[File:Bee hummingbird (Mellisuga helenae) adult male in flight-cropped.jpg|thumb|An adult male [[bee hummingbird]], the smallest known and the smallest [[Neontology|living]] dinosaur]]
[[File:Bee hummingbird (Mellisuga helenae) adult male in flight-cropped.jpg|thumb|An adult male [[bee hummingbird]], the smallest known and the smallest [[Neontology|living]] dinosaur]]
Size is an important aspect of [[dinosaur]] [[paleontology]], of interest to both the general public and professional scientists. Dinosaurs show some of the most extreme variations in size of any land animal group, ranging from tiny [[hummingbird]]s, which can weigh as little as two grams, to the extinct [[titanosaur]]s, which could weigh as much as {{convert|50-100|t|ST|abbr=on}}.
Size is an important aspect of [[dinosaur]] [[paleontology]], of interest to both the general public and professional scientists. Dinosaurs show some of the most extreme variations in size of any land animal group, ranging from tiny [[hummingbird]]s, which can weigh as little as two grams, to the extinct [[titanosaur]]s, such as ''[[Argentinosaurus]]'' and ''[[Bruhathkayosaurus]]''<ref>{{Cite journal |last=Paul |first=Gregory S. |last2=Larramendi |first2=Asier |date=2023-04-11 |title=Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales |url=https://www.idunn.no/doi/10.18261/let.56.2.5 |journal=Lethaia |language=en |volume=56 |issue=2 |pages=1–11 |doi=10.18261/let.56.2.5 |issn=0024-1164}}</ref> which could weigh as much as {{convert|50-130|t|ST|abbr=on}}.


The latest evidence suggests that dinosaurs' average size varied through the Triassic, early Jurassic, late Jurassic and Cretaceous periods, and dinosaurs probably only became widespread during the early or mid Jurassic.<ref name="Sereno1999">{{cite journal|author=Sereno PC|title=The evolution of dinosaurs|year=1999|journal=Science|volume=284|issue=5423|pages=2137–2147|doi=10.1126/science.284.5423.2137|pmid=10381873}}</ref> Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the {{Convert|100-1000|kg|lbs|abbr=on}} category when sorted by estimated weight into categories based on [[order of magnitude]], whereas [[Holocene|recent]] predatory [[carnivora]]n mammals peak in the range of {{Convert|10-100|kg|lbs|abbr=on}}.<ref name="JF93">{{cite book |author=Farlow JA|year=1993 |title=Functional Morphology and Evolution |editor=Dodson, Peter |editor2=Gingerich, Philip |series=American Journal of Science, Special Volume |volume=293-A |chapter=On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large, carnivorous dinosaurs |pages=167–199}}</ref> The [[Mode (statistics)|mode]] of Mesozoic dinosaur body masses is between one and ten metric tonnes.<ref name="Peczkis1994">{{Cite journal | doi = 10.1080/02724634.1995.10011575 | author = Peczkis, J. | year = 1994 | title = Implications of body-mass estimates for dinosaurs |journal = Journal of Vertebrate Paleontology | volume = 14 | issue = 4 | pages = 520–33 }}</ref> This contrasts sharply with the size of [[Cenozoic]] mammals, estimated by the [[National Museum of Natural History]] as about {{convert|2|to|5|kg|abbr=on}}.<ref name="NMNH">{{cite web |url=http://paleobiology.si.edu/dinosaurs/info/everything/evo_1.html |title=Anatomy and evolution |access-date=2007-11-21 |publisher=National Museum of Natural History |archive-url=https://web.archive.org/web/20071111204903/http://paleobiology.si.edu/dinosaurs/info/everything/evo_1.html |archive-date=2007-11-11 |url-status=live }}</ref>
The latest evidence suggests that dinosaurs' average size varied through the [[Triassic]], [[early Jurassic]], [[late Jurassic]] and [[Cretaceous]] periods, and dinosaurs probably only became widespread during the early or [[Middle Jurassic|mid Jurassic]].<ref name="Sereno1999">{{cite journal|author=Sereno PC|title=The evolution of dinosaurs|year=1999|journal=Science|volume=284|issue=5423|pages=2137–2147|doi=10.1126/science.284.5423.2137|pmid=10381873}}</ref> Predatory [[Theropoda|theropod]] dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the {{Convert|100-1000|kg|lbs|abbr=on}} category when sorted by estimated weight into categories based on [[order of magnitude]], whereas [[Holocene|recent]] predatory [[carnivora]]n mammals peak in the range of {{Convert|10-100|kg|lbs|abbr=on}}.<ref name="JF93">{{cite book |author=Farlow JA|year=1993 |title=Functional Morphology and Evolution |editor=Dodson, Peter |editor2=Gingerich, Philip |series=American Journal of Science, Special Volume |volume=293-A |chapter=On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large, carnivorous dinosaurs |pages=167–199}}</ref> The [[Mode (statistics)|mode]] of Mesozoic dinosaur body masses is between one and ten metric tonnes.<ref name="Peczkis1994">{{Cite journal | doi = 10.1080/02724634.1995.10011575 | author = Peczkis, J. | year = 1994 | title = Implications of body-mass estimates for dinosaurs |journal = Journal of Vertebrate Paleontology | volume = 14 | issue = 4 | pages = 520–33 }}</ref> This contrasts sharply with the size of [[Cenozoic]] mammals, estimated by the [[National Museum of Natural History]] as about {{convert|2|to|5|kg|abbr=on}}.<ref name="NMNH">{{cite web |url=http://paleobiology.si.edu/dinosaurs/info/everything/evo_1.html |title=Anatomy and evolution |access-date=2007-11-21 |publisher=National Museum of Natural History |archive-url=https://web.archive.org/web/20071111204903/http://paleobiology.si.edu/dinosaurs/info/everything/evo_1.html |archive-date=2007-11-11 |url-status=live }}</ref>


==Size estimation==
==Size estimation==
Scientists will probably never be certain of the [[largest organism|largest and smallest dinosaurs]]. This is because only a small fraction of animals ever fossilize, and most of these remains will either never be uncovered, or will be unintentionally destroyed as a result of human activity. Of the specimens that are recovered, few are even relatively complete skeletons, and impressions of skin and other soft tissues are rarely discovered. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species is an inexact art (though governed by some established [[allometry|allometric]] trends), and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork, and never perfect.<ref name="GSP10">{{cite book|last=Paul|first=Gregory S.|title=Princeton Field Guide to Dinosaurs|url=https://archive.org/details/princetonfieldgu0000paul|url-access=registration|year=2010|publisher=Princeton University Press|isbn=978-0-691-13720-9}}</ref> Mass estimates for dinosaurs are much more variable than length estimates given the lack of soft tissue preservation in the fossilization process. Modern mass estimation is often done with the laser scan skeleton technique that puts a "virtual" skin over the known or implied skeleton, but the limitations inherent in previous mass estimation techniques remain.<ref>[http://dinosaurs.about.com/od/dinosaurevolution/a/bigdinos.htm Strauss, Bob."Why Were Dinosaurs So Big? The Facts and Theories Behind Dinosaur Gigantism". About Education]{{Webarchive|url=https://web.archive.org/web/20141006142543/http://dinosaurs.about.com/od/dinosaurevolution/a/bigdinos.htm |date=2014-10-06 }}</ref>
Scientists will probably never be certain of the [[largest organism|largest]] and smallest dinosaurs. This is because only a small fraction of animals ever [[Fossilization|fossilize]], and most of these remains will either never be uncovered, or will be unintentionally destroyed as a result of human activity. Of the specimens that are recovered, few are even relatively complete skeletons, and impressions of skin and other soft tissues are rarely discovered. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species is an inexact art (though governed by some established [[allometry|allometric]] trends), and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork, and never perfect.<ref name="GSP10">{{cite book|last=Paul|first=Gregory S.|title=Princeton Field Guide to Dinosaurs|url=https://archive.org/details/princetonfieldgu0000paul|url-access=registration|year=2010|publisher=Princeton University Press|isbn=978-0-691-13720-9}}</ref> Mass estimates for dinosaurs are much more variable than length estimates given the lack of soft tissue preservation in the fossilization process. Modern mass estimation is often done with the laser scan skeleton technique that puts a "virtual" skin over the known or implied skeleton, but the limitations inherent in previous mass estimation techniques remain.<ref>[http://dinosaurs.about.com/od/dinosaurevolution/a/bigdinos.htm Strauss, Bob."Why Were Dinosaurs So Big? The Facts and Theories Behind Dinosaur Gigantism". About Education]{{Webarchive|url=https://web.archive.org/web/20141006142543/http://dinosaurs.about.com/od/dinosaurevolution/a/bigdinos.htm |date=2014-10-06 }}</ref>


==Sauropodomorphs==
==Sauropodomorphs==
{{Main|Sauropodomorpha}}
{{Main|Sauropodomorpha}}
[[File:Longest dinosaurs2.svg|alt=|thumb|upright=1.5|size comparison of selected giant sauropod dinosaurs]]
[[File:Longest dinosaurs2.svg|alt=|thumb|upright=1.5|Size comparison of selected giant sauropod dinosaurs]]
Sauropodomorph size is difficult to estimate given their usually fragmentary state of preservation. Sauropods are often preserved without their tails, so the margin of error in overall length estimates is high. Mass is calculated using the cube of the length, so for species in which the length is particularly uncertain, the weight is even more so. Estimates that are particularly uncertain (due to very fragmentary or lost material) are preceded by a question mark. Each number represents the highest estimate of a given research paper. One large sauropod, ''[[Maraapunisaurus fragillimus]]'', was based on particularly scant remains that have been lost since their description by paleontologists in 1878. Analysis of the illustrations included in the original report suggested that ''M. fragillimus'' may have been the largest land animal of all time, possibly weighing {{convert|100|-|150|MT|ST|abbr=on}} and measuring between {{convert|40|-|60|m|abbr=on}} long.<ref name="G.S.Paul2010"/><ref name="Paul1997">{{cite conference|last=Paul|first=G.S.|year=1997|editor2=Stump, E.|editor3=Rosenberg, G.D.|title=Dinosaur models: the good, the bad, and using them to estimate the mass of dinosaurs|url=http://gspauldino.com/Models.pdf|conference=Dinofest International|publisher=The Academy of Natural Sciences|pages=129–154|archive-url=https://web.archive.org/web/20160304084114/http://gspauldino.com/Models.pdf|archive-date=March 4, 2016|editor1=Wolberg, D.L.|book-title=DinoFest International Proceedings|url-status=live}}</ref> One later analysis of the surviving evidence, and the biological plausibility of such a large land animal, suggested that the enormous size of this animal was an over-estimate due partly to typographical errors in the original report.<ref name="woodruf&foster2015">{{cite journal |last1=Woodruff |first1=C |last2=Foster |first2=JR |date=2015 |title=The fragile legacy of ''Amphicoelias fragillimus'' (Dinosauria: Sauropoda; Morrison Formation - Latest Jurassic) |journal=PeerJ PrePrints |doi=10.7287/peerj.preprints.838v1|doi-access=free }}</ref> This would later be challenged by a different study, which argued Cope's measurements were genuine and there's no basis for assuming typographical errors. The study, however, also reclassified the species and correspondingly gave a much lower length estimate of {{convert|30.3|m|ft}} and a mass of {{convert|78.5|MT|ST|abbr=on}}.<ref>{{Cite journal|last=Carpenter|first=Kenneth|date=2018-10-19|title=''Maraapunisaurus fragillimus'', N.G. (formerly ''Amphicoelias fragillimus''), a basal Rebbachisaurid from the Morrison Formation (Upper Jurassic) of Colorado|url=https://giw.utahgeology.org/giw/index.php/GIW/article/view/32|journal=Geology of the Intermountain West|language=en|volume=5|pages=227–244|doi=10.31711/giw.v5.pp227-244|issn=2380-7601|doi-access=free}}</ref> This in itself would later be disputed as being too small for an animal of such size, with some believing it to be even larger at around {{convert|35-40|m|ft}} and weighing around {{convert|80-120|MT|ST|abbr=on}}.<ref>{{Cite journal|last=Paul|first=Gregory S.|date=2019|title=Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals|url=http://www.gspauldino.com/Titanomass.pdf|journal=Annals of the Carnegie Museum|volume=85|issue=4|pages=335–358|doi=10.2992/007.085.0403|s2cid=210840060}}</ref>
Sauropodomorph size is difficult to estimate given their usually fragmentary state of preservation. Sauropods are often preserved without their tails, so the margin of error in overall length estimates is high. Mass is calculated using the cube of the length, so for species in which the length is particularly uncertain, the weight is even more so. Estimates that are particularly uncertain (due to very fragmentary or lost material) are preceded by a question mark. Each number represents the highest estimate of a given research paper. One large sauropod, ''[[Maraapunisaurus fragillimus]]'', was based on particularly scant remains that have been lost since their description by paleontologists in 1878. Analysis of the illustrations included in the original report suggested that ''M. fragillimus'' may have been the largest land animal of all time, possibly weighing {{convert|100|-|150|MT|ST|abbr=on}} and measuring between {{convert|40|-|60|m|abbr=on}} long.<ref name="G.S.Paul2010"/><ref name="Paul1997">{{cite conference|last=Paul|first=G.S.|year=1997|editor2=Stump, E.|editor3=Rosenberg, G.D.|title=Dinosaur models: the good, the bad, and using them to estimate the mass of dinosaurs|url=http://gspauldino.com/Models.pdf|conference=Dinofest International|publisher=The Academy of Natural Sciences|pages=129–154|archive-url=https://web.archive.org/web/20160304084114/http://gspauldino.com/Models.pdf|archive-date=March 4, 2016|editor1=Wolberg, D.L.|book-title=DinoFest International Proceedings|url-status=live}}</ref> One later analysis of the surviving evidence, and the biological plausibility of such a large land animal, suggested that the enormous size of this animal was an over-estimate due partly to typographical errors in the original report.<ref name="woodruf&foster2015">{{cite journal |last1=Woodruff |first1=C |last2=Foster |first2=JR |date=2015 |title=The fragile legacy of ''Amphicoelias fragillimus'' (Dinosauria: Sauropoda; Morrison Formation - Latest Jurassic) |journal=PeerJ PrePrints |doi=10.7287/peerj.preprints.838v1|doi-access=free }}</ref> This would later be challenged by a different study, which argued Cope's measurements were genuine and that there was no basis for assuming typographical errors. The study, however, also reclassified the species and correspondingly gave a much lower length estimate of {{convert|30.3|m|ft}} and a mass of {{convert|78.5|MT|ST|abbr=on}}.<ref>{{Cite journal|last=Carpenter|first=Kenneth|date=2018-10-19|title=''Maraapunisaurus fragillimus'', N.G. (formerly ''Amphicoelias fragillimus''), a basal Rebbachisaurid from the Morrison Formation (Upper Jurassic) of Colorado|url=https://giw.utahgeology.org/giw/index.php/GIW/article/view/32|journal=Geology of the Intermountain West|language=en|volume=5|pages=227–244|doi=10.31711/giw.v5.pp227-244|issn=2380-7601|doi-access=free}}</ref> This in itself would later be disputed as being too small for an animal of such size, with some believing it to be even larger at around {{convert|35-40|m|ft}} and weighing around {{convert|80-120|MT|ST|abbr=on}}.<ref>{{Cite journal|last=Paul|first=Gregory S.|date=2019|title=Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals|url=http://www.gspauldino.com/Titanomass.pdf|journal=Annals of the Carnegie Museum|volume=85|issue=4|pages=335–358|doi=10.2992/007.085.0403|s2cid=210840060}}</ref>


Another absurdly large but even more controversial sauropod is [[Bruhathkayosaurus]] which had a calculated weight ranging between {{convert|126|-|220|MT|ST|abbr=on}} and a length of {{convert|44.1|m|abbr=on}}<ref name="mortimer2001a">Mortimer, M. (2001), [http://dml.cmnh.org/2001Jun/msg00665.html "Re: Bruhathkayosaurus"], discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.</ref><ref name="mortimer2004">Mortimer, M. (2004), [http://dml.cmnh.org/2004Sep/msg00086.html "Re: Largest Dinosaurs"], discussion group, The Dinosaur Mailing List, 7 September 2004. Accessed 23 May 2008.</ref><ref name="mortimer2001b">Mortimer, M. (2001), [http://dml.cmnh.org/2001Sep/msg00402.html "Titanosaurs too large?"], discussion group, The Dinosaur Mailing List, 12 September 2001. Accessed 23 May 2008.</ref> Although the existence of this sauropod had long been dismissed as a potential fake or a misidentification of a petrified tree trunk, recent photographic evidence emerged, confirming its existence.<ref>{{Cite journal |last1=Pal |first1=Saurabh |last2=Ayyasami |first2=Krishnan |date=27 June 2022 |title=The lost titan of Cauvery |journal=[[Geology Today]] |language=en |volume=38 |issue=3 |pages=112–116 |doi=10.1111/gto.12390 |s2cid=250056201 |issn=0266-6979}}</ref> More recent and reliable estimates in 2023 have rescaled Bruhathkayosaurus to weigh around {{convert|110|-|170|MT|ST|abbr=on}}, making it incredibly massive for such an animal.<ref>{{Cite journal |last1=Paul |first1=Gregory S. |last2=Larramendi |first2=Asier |date=11 April 2023 |title=Body mass estimate of ''Bruhathkayosaurus'' and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales |journal=Lethaia |language=en |volume=56 |issue=2 |pages=1–11 |doi=10.18261/let.56.2.5 |issn=0024-1164}}</ref> If the upper size estimates were to be taken at face value, Bruhathkayosaurus would not only be the largest dinosaur to have ever lived, but also the largest animal to have lived, exceeding even the largest [[blue whale]] recorded.
Another large but even more controversial sauropod is ''[[Bruhathkayosaurus]]'', which had a calculated weight ranging between {{convert|126|-|220|MT|ST|abbr=on}} and a length of {{convert|44.1|m|abbr=on}}<ref name="mortimer2001a">Mortimer, M. (2001), [http://dml.cmnh.org/2001Jun/msg00665.html "Re: Bruhathkayosaurus"], discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.</ref><ref name="mortimer2004">Mortimer, M. (2004), [http://dml.cmnh.org/2004Sep/msg00086.html "Re: Largest Dinosaurs"] {{Webarchive|url=https://web.archive.org/web/20190913171408/http://dml.cmnh.org/2004Sep/msg00086.html |date=2019-09-13 }}, discussion group, The Dinosaur Mailing List, 7 September 2004. Accessed 23 May 2008.</ref><ref name="mortimer2001b">Mortimer, M. (2001), [http://dml.cmnh.org/2001Sep/msg00402.html "Titanosaurs too large?"] {{Webarchive|url=https://web.archive.org/web/20160303203315/http://dml.cmnh.org/2001Sep/msg00402.html |date=2016-03-03 }}, discussion group, The Dinosaur Mailing List, 12 September 2001. Accessed 23 May 2008.</ref> Although the existence of this sauropod had long been dismissed as a potential fake or a misidentification of a petrified tree trunk, recent photographic evidence emerged, confirming its existence.<ref>{{Cite journal |last1=Pal |first1=Saurabh |last2=Ayyasami |first2=Krishnan |date=27 June 2022 |title=The lost titan of Cauvery |journal=[[Geology Today]] |language=en |volume=38 |issue=3 |pages=112–116 |doi=10.1111/gto.12390 |s2cid=250056201 |issn=0266-6979}}</ref> More recent and reliable estimates in 2023 have rescaled ''Bruhathkayosaurus'' to weigh around {{convert|110|-|130|MT|ST|abbr=on}} with its most liberal estimate being {{convert|240|MT|ST|abbr=on}}, making it incredibly massive for such an animal.<ref name="Bruhathkayosaurus2023">{{Cite journal |last1=Paul |first1=Gregory S. |last2=Larramendi |first2=Asier |date=11 April 2023 |title=Body mass estimate of ''Bruhathkayosaurus'' and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales |journal=Lethaia |language=en |volume=56 |issue=2 |pages=1–11 |doi=10.18261/let.56.2.5 |s2cid=259782734 |issn=0024-1164}}</ref> If the upper unlikely size estimates were to be taken at face value, ''Bruhathkayosaurus'' would not only be the largest dinosaur to have ever lived, but also the largest animal to have lived, exceeding even the largest [[blue whale]] recorded. According to [[Gregory S. Paul]], 'super-sauropods' or 'land-whales' such as ''Maraapunisaurus'', ''Bruhathkayosaurus'' and the "Broome Titanosaur footprints," as he calls them, should not be surprising as sauropods were more heat tolerant and grew rapidly, which allowed them to reach truly titanic sizes that rivaled the largest whales in mass despite the prevalence of air sacs.<ref name="Bruhathkayosaurus2023"/> Other potential factors for such extreme sauropod sizes include increasing bone robustness and load-distributing cartilaginous features to better redistribute and support such massive weights.<ref name="Bruhathkayosaurus2023"/>


Generally, the giant sauropods can be divided into two categories: the shorter but stockier and more massive forms (mainly [[titanosaur]]s and some [[Brachiosauridae|brachiosaurids]]), and the longer but slenderer and more light-weight forms (mainly [[diplodocid]]s).
Generally, the giant sauropods can be divided into two categories: the shorter but stockier and more massive forms (mainly [[titanosaur]]s and some [[Brachiosauridae|brachiosaurids]]), and the longer but slenderer and more light-weight forms (mainly [[diplodocid]]s).
Line 26: Line 26:
One of the tallest and heaviest dinosaurs known from good skeletons is ''[[Giraffatitan|Giraffatitan brancai]]'' (previously classified as a species of ''[[Brachiosaurus]]''). Its remains were discovered in [[Tanzania]] between 1907 and 1912. Bones from several similar-sized individuals were incorporated into the skeleton now mounted and on display at the [[Museum für Naturkunde|Museum für Naturkunde Berlin]];<ref name="EC68">{{cite book |author=Colbert, Edwin Harris |title=Men and dinosaurs: the search in field and laboratory |publisher=Penguin |place=Harmondsworth [Eng.] |year=1971 |isbn=978-0-14-021288-4}}</ref> this mount is {{convert|12-13.27|m|}} tall and {{convert|21.8|-|22.5|m}} long,<ref>{{cite journal|last=Mazzetta|first=G.V.|title=Giants and Bizarres: Body Size of Some Southern South American Cretaceous Dinosaurs|journal=Historical Biology|year=2004|pages=1–13|display-authors=etal|doi=10.1080/08912960410001715132|volume=16|issue=2–4|citeseerx=10.1.1.694.1650|s2cid=56028251}}</ref><ref>{{cite journal|last=Janensch|first=W.|title=The Skeleton Reconstruction of ''Brachiosaurus brancai''|year=1950|pages=97–103}}</ref><ref>{{Cite news|url=https://www.museumfuernaturkunde.berlin/en/museum/ausstellungen/world-dinosaurs|title=The World of Dinosaurs|work=Museum für Naturkunde|access-date=2018-11-16|language=en|archive-url=https://web.archive.org/web/20181116215935/https://www.museumfuernaturkunde.berlin/en/museum/ausstellungen/world-dinosaurs|archive-date=2018-11-16|url-status=live}}</ref> and would have belonged to an animal that weighed between {{convert|30000|to|60000|kg|lb}}. One of the longest complete dinosaurs is the {{convert|27|m|ft|adj=mid|-long}} ''[[Diplodocus]]'', which was discovered in [[Wyoming]] in the United States and displayed in [[Pittsburgh, Pennsylvania|Pittsburgh's]] [[Carnegie Natural History Museum]] in 1907.<ref name="hecket04">{{cite journal | last1 = Lucas | first1 = H. | last2 = Hecket | first2 = H. | year = 2004 | title = Reappraisal of ''Seismosaurus'', a Late Jurassic Sauropod | journal = Proceeding, Annual Meeting of the Society of Paleontology | volume = 36 | issue = 5| page = 422 }}</ref>
One of the tallest and heaviest dinosaurs known from good skeletons is ''[[Giraffatitan|Giraffatitan brancai]]'' (previously classified as a species of ''[[Brachiosaurus]]''). Its remains were discovered in [[Tanzania]] between 1907 and 1912. Bones from several similar-sized individuals were incorporated into the skeleton now mounted and on display at the [[Museum für Naturkunde|Museum für Naturkunde Berlin]];<ref name="EC68">{{cite book |author=Colbert, Edwin Harris |title=Men and dinosaurs: the search in field and laboratory |publisher=Penguin |place=Harmondsworth [Eng.] |year=1971 |isbn=978-0-14-021288-4}}</ref> this mount is {{convert|12-13.27|m|}} tall and {{convert|21.8|-|22.5|m}} long,<ref>{{cite journal|last=Mazzetta|first=G.V.|title=Giants and Bizarres: Body Size of Some Southern South American Cretaceous Dinosaurs|journal=Historical Biology|year=2004|pages=1–13|display-authors=etal|doi=10.1080/08912960410001715132|volume=16|issue=2–4|citeseerx=10.1.1.694.1650|s2cid=56028251}}</ref><ref>{{cite journal|last=Janensch|first=W.|title=The Skeleton Reconstruction of ''Brachiosaurus brancai''|year=1950|pages=97–103}}</ref><ref>{{Cite news|url=https://www.museumfuernaturkunde.berlin/en/museum/ausstellungen/world-dinosaurs|title=The World of Dinosaurs|work=Museum für Naturkunde|access-date=2018-11-16|language=en|archive-url=https://web.archive.org/web/20181116215935/https://www.museumfuernaturkunde.berlin/en/museum/ausstellungen/world-dinosaurs|archive-date=2018-11-16|url-status=live}}</ref> and would have belonged to an animal that weighed between {{convert|30000|to|60000|kg|lb}}. One of the longest complete dinosaurs is the {{convert|27|m|ft|adj=mid|-long}} ''[[Diplodocus]]'', which was discovered in [[Wyoming]] in the United States and displayed in [[Pittsburgh, Pennsylvania|Pittsburgh's]] [[Carnegie Natural History Museum]] in 1907.<ref name="hecket04">{{cite journal | last1 = Lucas | first1 = H. | last2 = Hecket | first2 = H. | year = 2004 | title = Reappraisal of ''Seismosaurus'', a Late Jurassic Sauropod | journal = Proceeding, Annual Meeting of the Society of Paleontology | volume = 36 | issue = 5| page = 422 }}</ref>


There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest [[herbivore|herbivorous]] specimens on record were discovered in the 1970s or later, and include the massive [[titanosaur]] ''[[Argentinosaurus huinculensis]]'', which is the largest dinosaur known from uncontroversial and relatively substantial evidence, estimated to have been {{convert|70-80|t|ST|abbr=on}} and {{convert|36|m|ft|abbr=on}} long.<ref name=":5">{{Cite book|last1=Molina-Pérez|first1=Rubén|title=Dinosaur Facts and Figures The Sauropods and Other Sauropodomorphs.|last2=Larramendi|first2=Asier|publisher=Princeton University Press|others=Illustrated by Andrey Atuchin and Sante Mazzei|year=2020|isbn=978-0-691-20297-6|location=Princeton|translator-last=Donaghey|translator-first=Joan|oclc=1157079384}}</ref><ref name="G.S.Paul2010">{{cite book|last=Paul|first=Gregory S.|title=The Princeton Field Guide to Dinosaurs: 2nd Edition|publisher=Princeton University Press|year=2016|isbn=978-0-691-16766-4|place=United States of America}}</ref> Some of the longest sauropods were those with exceptionally long, whip-like tails, such as the {{convert|29-30|m|ft|abbr=on}} ''[[Diplodocus|Diplodocus hallorum]]''<ref name="G.S.Paul2010" /><ref name=":5" /> (formerly ''Seismosaurus'') and the {{Convert|45|m|ft|abbr=on}} ''Barosaurus''.<ref name=":5" />
There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest [[herbivore|herbivorous]] specimens on record were discovered in the 1970s or later, and include the massive [[titanosaur]] ''[[Argentinosaurus huinculensis]]'', which is the largest dinosaur known from uncontroversial and relatively substantial evidence, estimated to have been {{convert|70-80|t|ST|abbr=on}} and {{convert|36|m|ft|abbr=on}} long.<ref name=":5">{{Cite book|last1=Molina-Pérez|first1=Rubén|title=Dinosaur Facts and Figures The Sauropods and Other Sauropodomorphs.|last2=Larramendi|first2=Asier|publisher=Princeton University Press|others=Illustrated by Andrey Atuchin and Sante Mazzei|year=2020|isbn=978-0-691-20297-6|location=Princeton|translator-last=Donaghey|translator-first=Joan|oclc=1157079384}}</ref><ref name="G.S.Paul2010">{{cite book|last=Paul|first=Gregory S.|title=The Princeton Field Guide to Dinosaurs: 2nd Edition|publisher=Princeton University Press|year=2016|isbn=978-0-691-16766-4|place=United States of America}}</ref> Some of the longest sauropods were those with exceptionally long, whip-like tails, such as the {{convert|29-30|m|ft|abbr=on}} ''[[Diplodocus|Diplodocus hallorum]]''<ref name="G.S.Paul2010" /><ref name=":5" /> (formerly ''Seismosaurus'') and the 39 m ''[[Supersaurus]]''.<ref>{{Cite web |title=SVP 2021 Symposium |url=https://vertpaleo.org/svp-2021-symposium/ |access-date=2023-12-16 |website=Society of Vertebrate Paleontology |language=en-US}}</ref>


In 2014, the fossilized remains of a previously unknown species of sauropod were discovered in Argentina.<ref>{{Cite news|url=https://www.bbc.co.uk/news/science-environment-27441156|title='Biggest dinosaur ever' discovered|last=Morgan|first=James|date=2014-05-17|work=BBC News|access-date=2017-03-21|language=en-GB|archive-url=https://web.archive.org/web/20170218032155/http://www.bbc.co.uk/news/science-environment-27441156|archive-date=2017-02-18|url-status=live}}</ref> The titanosaur, named ''[[Patagotitan mayorum]]'', was estimated to have been around {{Convert|40|m|ft|abbr=on}} long weighing around {{Convert|77|t|ST|abbr=on}}, larger than any other previously found sauropod. The specimens found were remarkably complete, significantly more so than previous titanosaurs. It since been suggested that ''Patagotitan'' was not necessarily larger than ''Argentinosaurus'' and ''Puertasaurus''.<ref>{{Cite web|url=https://svpow.com/2017/08/09/dont-believe-the-hype-patagotitan-was-not-bigger-than-argentinosaurus/|title=Don't believe the hype: ''Patagotitan'' was not bigger than ''Argentinosaurus''|date=2017-08-09|website=Sauropod Vertebra Picture of the Week|language=en|access-date=2019-02-19|archive-url=https://web.archive.org/web/20190414075959/https://svpow.com/2017/08/09/dont-believe-the-hype-patagotitan-was-not-bigger-than-argentinosaurus/|archive-date=2019-04-14|url-status=live}}</ref> In 2019, ''Patagotitan'' was estimated to have been {{Convert|31|m|ft|abbr=}} long and {{Convert|50-77|t|lb|abbr=}} massive.<ref name=":3">{{Cite journal|last=Paul|first=Gregory|date=December 2019|title=Determining the Largest Known Land Animal: A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals|url=http://gspauldino.com/Titanomass.pdf|journal=Annals of Carnegie Museum|volume=85|issue=4|pages=335–358|doi=10.2992/007.085.0403|issn=0097-4463|s2cid=210840060}}</ref>
In 2014, the fossilized remains of a previously unknown species of sauropod were discovered in Argentina.<ref>{{Cite news|url=https://www.bbc.co.uk/news/science-environment-27441156|title='Biggest dinosaur ever' discovered|last=Morgan|first=James|date=2014-05-17|work=BBC News|access-date=2017-03-21|language=en-GB|archive-url=https://web.archive.org/web/20170218032155/http://www.bbc.co.uk/news/science-environment-27441156|archive-date=2017-02-18|url-status=live}}</ref> The titanosaur, named ''[[Patagotitan mayorum]]'', was estimated to have been around {{Convert|40|m|ft|abbr=on}} long weighing around {{Convert|77|t|ST|abbr=on}}, larger than any other previously found sauropod. The specimens found were remarkably complete, significantly more so than previous titanosaurs. It since been suggested that ''Patagotitan'' was not necessarily larger than ''Argentinosaurus'' and ''[[Puertasaurus]]''.<ref>{{Cite web|url=https://svpow.com/2017/08/09/dont-believe-the-hype-patagotitan-was-not-bigger-than-argentinosaurus/|title=Don't believe the hype: ''Patagotitan'' was not bigger than ''Argentinosaurus''|date=2017-08-09|website=Sauropod Vertebra Picture of the Week|language=en|access-date=2019-02-19|archive-url=https://web.archive.org/web/20190414075959/https://svpow.com/2017/08/09/dont-believe-the-hype-patagotitan-was-not-bigger-than-argentinosaurus/|archive-date=2019-04-14|url-status=live}}</ref> In 2019, ''Patagotitan'' was estimated to have been {{Convert|31|m|ft|abbr=}} long and about {{Convert|55|t|lb|abbr=}}.<ref name=":3">{{Cite journal|last=Paul|first=Gregory|date=December 2019|title=Determining the Largest Known Land Animal: A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals|url=http://gspauldino.com/Titanomass.pdf|journal=Annals of Carnegie Museum|volume=85|issue=4|pages=335–358|doi=10.2992/007.085.0403|issn=0097-4463|s2cid=210840060}}</ref>


The largest of non-sauropod [[Sauropodomorpha|sauropodomorphs]] was ''[[Euskelosaurus]]''. It reached {{cvt|12.2|m}} in length and {{cvt|2|MT|ST}} in weight.<ref>{{cite book
The largest of non-sauropod [[Sauropodomorpha|sauropodomorphs]] was the unnamed {{Convert|16|m|ft}} long {{Convert|10|t|lb}} unnamed Elliot giant.<ref name=":5" /> Another large sauropodomorph was ''[[Euskelosaurus]]''. It reached {{cvt|12.2|m}} in length and {{cvt|2|MT|ST}} in weight.<ref>{{cite book
| url = https://books.google.com/books?id=QCC9DwAAQBAJ
| url = https://books.google.com/books?id=QCC9DwAAQBAJ
| title = The Fossil Book: A Record of Prehistoric Life
| title = The Fossil Book: A Record of Prehistoric Life
Line 39: Line 39:
| author = [[Patricia Vickers-Rich|Patricia Vickers Rich]], Thomas Hewitt Rich, [[Mildred Adams Fenton]], Carroll Lane
| author = [[Patricia Vickers-Rich|Patricia Vickers Rich]], Thomas Hewitt Rich, [[Mildred Adams Fenton]], Carroll Lane
| isbn = 9780486838557
| isbn = 9780486838557
}}</ref> Another large sauropodomorph ''[[Yunnanosaurus youngi]]'' reached {{cvt|13|m}} long.<ref>Lu, J., Li, T., Zhong, S., Azuma, Y., Fujita, M., Dong, Z., and Ji, Q. (2007). "[https://www.semanticscholar.org/paper/NEW-YUNNANOSAURID-DINOSAUR-(DINOSAURIA%2C-FROM-THE-OF-Junchang-Li/a8549bec983b40bcb561a5bc3ff4e1bc0b0f6979 New yunnanosaurid dinosaur (Dinosauria, Prosauropoda) from the Middle Jurassic Zhanghe Formation of Yuanmou, Yunnan Province of China]". Memoir of the Fukui Prefectural Dinosaur Museum, 6: 1-15.</ref>
}}</ref> ''[[Yunnanosaurus youngi]]'' also reached a length of {{cvt|13|m}}.<ref>Lu, J., Li, T., Zhong, S., Azuma, Y., Fujita, M., Dong, Z., and Ji, Q. (2007). "[https://www.semanticscholar.org/paper/NEW-YUNNANOSAURID-DINOSAUR-(DINOSAURIA%2C-FROM-THE-OF-Junchang-Li/a8549bec983b40bcb561a5bc3ff4e1bc0b0f6979 New yunnanosaurid dinosaur (Dinosauria, Prosauropoda) from the Middle Jurassic Zhanghe Formation of Yuanmou, Yunnan Province of China]". Memoir of the Fukui Prefectural Dinosaur Museum, 6: 1-15.</ref>


==Theropods==
==Theropods==
Line 48: Line 48:
The largest extant theropod is the [[common ostrich]], up to {{convert|2.74|m}} tall and weighs between {{convert|63.5|and|145.15|kg|lb}}.<ref>{{Cite web| url=http://www.awf.org/wildlife-conservation/ostrich| title=See what African Wildlife Foundation is doing to protect these iconic flightless birds| date=2013-02-25| access-date=2015-04-18| archive-url=https://web.archive.org/web/20150429040924/http://www.awf.org/wildlife-conservation/ostrich| archive-date=2015-04-29| url-status=live}}</ref>
The largest extant theropod is the [[common ostrich]], up to {{convert|2.74|m}} tall and weighs between {{convert|63.5|and|145.15|kg|lb}}.<ref>{{Cite web| url=http://www.awf.org/wildlife-conservation/ostrich| title=See what African Wildlife Foundation is doing to protect these iconic flightless birds| date=2013-02-25| access-date=2015-04-18| archive-url=https://web.archive.org/web/20150429040924/http://www.awf.org/wildlife-conservation/ostrich| archive-date=2015-04-29| url-status=live}}</ref>


The smallest non-avialan theropod known from adult specimens may be ''[[Anchiornis|Anchiornis huxleyi]]'', at {{convert|110|g|oz|abbr=off}} in weight and {{convert|34|cm}} in length,<ref name="anchiadvance">Xu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." ''Chinese Science Bulletin'', 6 pages, accepted November 15, 2008.</ref> although later study discovered larger specimen reaching {{convert|62|cm}}.<ref>{{Cite journal |last1=Pei |first1=Rui |last2=Li |first2=Quanguo |last3=Meng |first3=Qingjin |last4=Norell |first4=Mark A. |last5=Gao |first5=Ke-Qin |date=2017 |title=New Specimens of<i>Anchiornis huxleyi</i>(Theropoda: Paraves) from the Late Jurassic of Northeastern China |url=http://dx.doi.org/10.1206/0003-0090-411.1.1 |journal=Bulletin of the American Museum of Natural History |volume=411 |pages=1–67 |doi=10.1206/0003-0090-411.1.1 |issn=0003-0090}}</ref> However, some studies suggest that ''Anchiornis'' was actually an avialan.<ref name="Nature">{{Cite journal|author1=Pascal Godefroit |author2=Andrea Cau |author3=Hu Dong-Yu |author4=François Escuillié |author5=Wu Wenhao |author6=Gareth Dyke |year=2013 |title=A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds |journal=Nature |volume= 498|issue= 7454|pages= 359–62|doi=10.1038/nature12168 |pmid=23719374|bibcode = 2013Natur.498..359G |s2cid=4364892 }}</ref> The smallest dinosaur known from adult specimens which is definitely not an avialan is ''[[Parvicursor remotus]]'', at {{convert|162|g|oz|abbr=}} and measuring {{convert|39|cm|in|abbr=}} long.<ref>[http://www.technosaurs.ca/tag/default.aspx?id=114 Which was the smallest dinosaur?] {{webarchive|url=https://web.archive.org/web/20110706204350/http://www.technosaurs.ca/tag/default.aspx?id=114 |date=2011-07-06 }} Royal Tyrrell Museum. Last accessed 22 September 2022.</ref> However, in 2022 its [[holotype]] was recognized as a juvenile individual.<ref>{{cite journal| vauthors = Averianov AO, Lopatin AV |title=A re-appraisal of ''Parvicursor remotus'' from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs |journal=Journal of Systematic Palaeontology |year=2022 |volume=19 |issue=16 |pages=1097–1128 |doi=10.1080/14772019.2021.2013965 |s2cid=247222017 }}</ref> Among living dinosaurs, the [[bee hummingbird]] (''Mellisuga helenae'') is smallest at {{convert|1.9|g|abbr=on}} and {{convert|5.5|cm|abbr=on}} long.<ref name="eoe">[http://www.eoearth.org/article/Biological_diversity_in_the_Caribbean_Islands Conservation International (Content Partner); Mark McGinley (Topic Editor). 2008. "Biological diversity in the Caribbean Islands." In: Encyclopedia of Earth. Eds. Cutler J. Cleveland (Washington, D.C.: Environmental Information Coalition, National Council for Science and the Environment). &#91;First published in the Encyclopedia of Earth May 3, 2007; Last revised August 22, 2008; Retrieved November 9, 2009&#93;.] {{Webarchive|url=https://web.archive.org/web/20130523191902/http://www.eoearth.org/article/Biological_diversity_in_the_Caribbean_Islands |date=2013-05-23 }}></ref>
The smallest non-[[Avialae|avialan]] theropod known from adult specimens may be ''[[Anchiornis|Anchiornis huxleyi]]'', at {{convert|110|g|oz|abbr=off}} in weight and {{convert|34|cm}} in length,<ref name="anchiadvance">Xu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." ''Chinese Science Bulletin'', 6 pages, accepted November 15, 2008.</ref> although later study discovered larger specimen reaching {{convert|62|cm}}.<ref>{{Cite journal |last1=Pei |first1=Rui |last2=Li |first2=Quanguo |last3=Meng |first3=Qingjin |last4=Norell |first4=Mark A. |last5=Gao |first5=Ke-Qin |date=2017 |title=New Specimens of''Anchiornis huxleyi''(Theropoda: Paraves) from the Late Jurassic of Northeastern China |url=http://dx.doi.org/10.1206/0003-0090-411.1.1 |journal=Bulletin of the American Museum of Natural History |volume=411 |pages=1–67 |doi=10.1206/0003-0090-411.1.1 |s2cid=90650697 |issn=0003-0090}}</ref> However, some studies suggest that ''Anchiornis'' was actually an avialan.<ref name="Nature">{{Cite journal|author1=Pascal Godefroit |author2=Andrea Cau |author3=Hu Dong-Yu |author4=François Escuillié |author5=Wu Wenhao |author6=Gareth Dyke |year=2013 |title=A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds |journal=Nature |volume= 498|issue= 7454|pages= 359–62|doi=10.1038/nature12168 |pmid=23719374|bibcode = 2013Natur.498..359G |s2cid=4364892 }}</ref> The smallest dinosaur known from adult specimens which is definitely not an avialan is ''[[Parvicursor remotus]]'', at {{convert|162|g|oz|abbr=}} and measuring {{convert|39|cm|in|abbr=}} long.<ref>[http://www.technosaurs.ca/tag/default.aspx?id=114 Which was the smallest dinosaur?] {{webarchive|url=https://web.archive.org/web/20110706204350/http://www.technosaurs.ca/tag/default.aspx?id=114 |date=2011-07-06 }} Royal Tyrrell Museum. Last accessed 22 September 2022.</ref> However, in 2022 its [[holotype]] was recognized as a juvenile individual.<ref>{{cite journal| vauthors = Averianov AO, Lopatin AV |title=A re-appraisal of ''Parvicursor remotus'' from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs |journal=Journal of Systematic Palaeontology |year=2022 |volume=19 |issue=16 |pages=1097–1128 |doi=10.1080/14772019.2021.2013965 |s2cid=247222017 }}</ref> Among living dinosaurs, the [[bee hummingbird]] (''Mellisuga helenae'') is smallest at {{convert|1.9|g|abbr=on}} and {{convert|5.5|cm|abbr=on}} long.<ref name="eoe">[http://www.eoearth.org/article/Biological_diversity_in_the_Caribbean_Islands Conservation International (Content Partner); Mark McGinley (Topic Editor). 2008. "Biological diversity in the Caribbean Islands." In: Encyclopedia of Earth. Eds. Cutler J. Cleveland (Washington, D.C.: Environmental Information Coalition, National Council for Science and the Environment). &#91;First published in the Encyclopedia of Earth May 3, 2007; Last revised August 22, 2008; Retrieved November 9, 2009&#93;.] {{Webarchive|url=https://web.archive.org/web/20130523191902/http://www.eoearth.org/article/Biological_diversity_in_the_Caribbean_Islands |date=2013-05-23 }}></ref> The smallest theropod overall (including avians) is the currently extant bee hummingbird at 6.12 cm long and 2.6g for females, and 5.51 cm long and 3.25g for the males.<ref>Glick, Adrienne [https://web.archive.org/web/20200508225701/https://animaldiversity.org/accounts/Mellisuga_helenae/ "Mellisuga helenae"] ''Animal Diversity Web'', Retrieved 04 October 2023</ref>


In [[evolution of birds|the theropod lineage leading to birds]], body size shrank continuously over a period of 50 million years, from an average of {{convert|163|kg}} down to {{convert|0.8|kg|abbr=on}}. This was the only dinosaur lineage to get continuously smaller over such an extended time period, and their skeletons developed adaptations at about four times the average rate for dinosaurs.<ref name="AP-20140731">
In [[evolution of birds|the theropod lineage leading to birds]], body size shrank continuously over a period of 50 million years, from an average of {{convert|163|kg}} down to {{convert|0.8|kg|abbr=on}}. This was the only dinosaur lineage to get continuously smaller over such an extended time period, and their skeletons developed adaptations at about four times the average rate for dinosaurs.<ref name="AP-20140731">
Line 59: Line 59:
*[[Megafauna]]
*[[Megafauna]]
*[[Pterosaur size]]
*[[Pterosaur size]]
*[[Argentinosaurus]]


==References==
==References==

Latest revision as of 20:25, 24 November 2024

Scale diagram comparing a human and the longest-known dinosaurs of five major clades
An adult male bee hummingbird, the smallest known and the smallest living dinosaur

Size is an important aspect of dinosaur paleontology, of interest to both the general public and professional scientists. Dinosaurs show some of the most extreme variations in size of any land animal group, ranging from tiny hummingbirds, which can weigh as little as two grams, to the extinct titanosaurs, such as Argentinosaurus and Bruhathkayosaurus[1] which could weigh as much as 50–130 t (55–143 short tons).

The latest evidence suggests that dinosaurs' average size varied through the Triassic, early Jurassic, late Jurassic and Cretaceous periods, and dinosaurs probably only became widespread during the early or mid Jurassic.[2] Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the 100–1,000 kg (220–2,200 lb) category when sorted by estimated weight into categories based on order of magnitude, whereas recent predatory carnivoran mammals peak in the range of 10–100 kg (22–220 lb).[3] The mode of Mesozoic dinosaur body masses is between one and ten metric tonnes.[4] This contrasts sharply with the size of Cenozoic mammals, estimated by the National Museum of Natural History as about 2 to 5 kg (4.4 to 11.0 lb).[5]

Size estimation

Scientists will probably never be certain of the largest and smallest dinosaurs. This is because only a small fraction of animals ever fossilize, and most of these remains will either never be uncovered, or will be unintentionally destroyed as a result of human activity. Of the specimens that are recovered, few are even relatively complete skeletons, and impressions of skin and other soft tissues are rarely discovered. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species is an inexact art (though governed by some established allometric trends), and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork, and never perfect.[6] Mass estimates for dinosaurs are much more variable than length estimates given the lack of soft tissue preservation in the fossilization process. Modern mass estimation is often done with the laser scan skeleton technique that puts a "virtual" skin over the known or implied skeleton, but the limitations inherent in previous mass estimation techniques remain.[7]

Sauropodomorphs

Size comparison of selected giant sauropod dinosaurs

Sauropodomorph size is difficult to estimate given their usually fragmentary state of preservation. Sauropods are often preserved without their tails, so the margin of error in overall length estimates is high. Mass is calculated using the cube of the length, so for species in which the length is particularly uncertain, the weight is even more so. Estimates that are particularly uncertain (due to very fragmentary or lost material) are preceded by a question mark. Each number represents the highest estimate of a given research paper. One large sauropod, Maraapunisaurus fragillimus, was based on particularly scant remains that have been lost since their description by paleontologists in 1878. Analysis of the illustrations included in the original report suggested that M. fragillimus may have been the largest land animal of all time, possibly weighing 100–150 t (110–170 short tons) and measuring between 40–60 m (130–200 ft) long.[8][9] One later analysis of the surviving evidence, and the biological plausibility of such a large land animal, suggested that the enormous size of this animal was an over-estimate due partly to typographical errors in the original report.[10] This would later be challenged by a different study, which argued Cope's measurements were genuine and that there was no basis for assuming typographical errors. The study, however, also reclassified the species and correspondingly gave a much lower length estimate of 30.3 metres (99 ft) and a mass of 78.5 t (86.5 short tons).[11] This in itself would later be disputed as being too small for an animal of such size, with some believing it to be even larger at around 35–40 metres (115–131 ft) and weighing around 80–120 t (88–132 short tons).[12]

Another large but even more controversial sauropod is Bruhathkayosaurus, which had a calculated weight ranging between 126–220 t (139–243 short tons) and a length of 44.1 m (145 ft)[13][14][15] Although the existence of this sauropod had long been dismissed as a potential fake or a misidentification of a petrified tree trunk, recent photographic evidence emerged, confirming its existence.[16] More recent and reliable estimates in 2023 have rescaled Bruhathkayosaurus to weigh around 110–130 t (120–140 short tons) with its most liberal estimate being 240 t (260 short tons), making it incredibly massive for such an animal.[17] If the upper unlikely size estimates were to be taken at face value, Bruhathkayosaurus would not only be the largest dinosaur to have ever lived, but also the largest animal to have lived, exceeding even the largest blue whale recorded. According to Gregory S. Paul, 'super-sauropods' or 'land-whales' such as Maraapunisaurus, Bruhathkayosaurus and the "Broome Titanosaur footprints," as he calls them, should not be surprising as sauropods were more heat tolerant and grew rapidly, which allowed them to reach truly titanic sizes that rivaled the largest whales in mass despite the prevalence of air sacs.[17] Other potential factors for such extreme sauropod sizes include increasing bone robustness and load-distributing cartilaginous features to better redistribute and support such massive weights.[17]

Generally, the giant sauropods can be divided into two categories: the shorter but stockier and more massive forms (mainly titanosaurs and some brachiosaurids), and the longer but slenderer and more light-weight forms (mainly diplodocids).

Because different methods of estimation sometimes give conflicting results, mass estimates for sauropods can vary widely causing disagreement among scientists over the accurate number. For example, the titanosaur Dreadnoughtus was originally estimated to weigh 59.3 tonnes by the allometric scaling of limb-bone proportions, whereas more recent estimates, based on three-dimensional reconstructions, yield a much smaller figure of 22.1–38.2 tonnes.[18]

Reconstructed skeleton of the titanosaur Argentinosaurus huinculensis, often considered the largest-known dinosaur

The sauropods were the longest and heaviest dinosaurs. For much of the dinosaur era, the smallest sauropods were larger than almost anything else in their habitat, and the largest were an order of magnitude more massive than anything else known to have walked the Earth since. Giant prehistoric mammals such as Paraceratherium and Palaeoloxodon (the largest land mammals ever discovered[19]) were dwarfed by the giant sauropods, and only modern whales approach or surpass them in weight, though they live in the oceans.[20] There are several proposed advantages for the large size of sauropods, including protection from predation, reduction of energy use, and longevity, but it may be that the most important advantage was dietary. Large animals are more efficient at digestion than small animals, because food spends more time in their digestive systems. This also permits them to subsist on food with lower nutritive value than smaller animals. Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry, and the ability to eat large quantities of low-nutrient browse would have been advantageous in such environments.[21]

One of the tallest and heaviest dinosaurs known from good skeletons is Giraffatitan brancai (previously classified as a species of Brachiosaurus). Its remains were discovered in Tanzania between 1907 and 1912. Bones from several similar-sized individuals were incorporated into the skeleton now mounted and on display at the Museum für Naturkunde Berlin;[22] this mount is 12–13.27 metres (39.4–43.5 ft) tall and 21.8–22.5 metres (72–74 ft) long,[23][24][25] and would have belonged to an animal that weighed between 30,000 to 60,000 kilograms (66,000 to 132,000 lb). One of the longest complete dinosaurs is the 27-metre-long (89 ft) Diplodocus, which was discovered in Wyoming in the United States and displayed in Pittsburgh's Carnegie Natural History Museum in 1907.[26]

There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were discovered in the 1970s or later, and include the massive titanosaur Argentinosaurus huinculensis, which is the largest dinosaur known from uncontroversial and relatively substantial evidence, estimated to have been 70–80 t (77–88 short tons) and 36 m (118 ft) long.[27][8] Some of the longest sauropods were those with exceptionally long, whip-like tails, such as the 29–30 m (95–98 ft) Diplodocus hallorum[8][27] (formerly Seismosaurus) and the 39 m Supersaurus.[28]

In 2014, the fossilized remains of a previously unknown species of sauropod were discovered in Argentina.[29] The titanosaur, named Patagotitan mayorum, was estimated to have been around 40 m (130 ft) long weighing around 77 t (85 short tons), larger than any other previously found sauropod. The specimens found were remarkably complete, significantly more so than previous titanosaurs. It since been suggested that Patagotitan was not necessarily larger than Argentinosaurus and Puertasaurus.[30] In 2019, Patagotitan was estimated to have been 31 metres (102 ft) long and about 55 tonnes (121,000 lb).[31]

The largest of non-sauropod sauropodomorphs was the unnamed 16 metres (52 ft) long 10 tonnes (22,000 lb) unnamed Elliot giant.[27] Another large sauropodomorph was Euskelosaurus. It reached 12.2 m (40 ft) in length and 2 t (2.2 short tons) in weight.[32] Yunnanosaurus youngi also reached a length of 13 m (43 ft).[33]

Theropods

Size comparison of selected giant theropod dinosaurs

Tyrannosaurus was for many decades the largest and best-known theropod to the general public. Since its discovery, however, a number of other giant carnivorous dinosaurs have been described, including Spinosaurus, Carcharodontosaurus, and Giganotosaurus.[34] These large theropod dinosaurs are estimated to rival or even exceeded Tyrannosaurus rex in size, though more recent studies and reconstructions show that Tyrannosaurus, although shorter, was the bulkier animal overall. Specimens such as Sue and Scotty are both estimated to be the most massive theropods known to science. There is still no clear explanation for exactly why these animals grew so bulky and heavy compared to the land predators that came before and after them.

Skeleton of Giganotosaurus, one of the largest theropods known.

The largest extant theropod is the common ostrich, up to 2.74 metres (9 ft 0 in) tall and weighs between 63.5 and 145.15 kilograms (140.0 and 320.0 lb).[35]

The smallest non-avialan theropod known from adult specimens may be Anchiornis huxleyi, at 110 grams (3.9 ounces) in weight and 34 centimetres (13 in) in length,[36] although later study discovered larger specimen reaching 62 centimetres (24 in).[37] However, some studies suggest that Anchiornis was actually an avialan.[38] The smallest dinosaur known from adult specimens which is definitely not an avialan is Parvicursor remotus, at 162 grams (5.7 oz) and measuring 39 centimetres (15 in) long.[39] However, in 2022 its holotype was recognized as a juvenile individual.[40] Among living dinosaurs, the bee hummingbird (Mellisuga helenae) is smallest at 1.9 g (0.067 oz) and 5.5 cm (2.2 in) long.[41] The smallest theropod overall (including avians) is the currently extant bee hummingbird at 6.12 cm long and 2.6g for females, and 5.51 cm long and 3.25g for the males.[42]

In the theropod lineage leading to birds, body size shrank continuously over a period of 50 million years, from an average of 163 kilograms (359 lb) down to 0.8 kg (1.8 lb). This was the only dinosaur lineage to get continuously smaller over such an extended time period, and their skeletons developed adaptations at about four times the average rate for dinosaurs.[43][44]

See also

References

  1. ^ Paul, Gregory S.; Larramendi, Asier (2023-04-11). "Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales". Lethaia. 56 (2): 1–11. doi:10.18261/let.56.2.5. ISSN 0024-1164.
  2. ^ Sereno PC (1999). "The evolution of dinosaurs". Science. 284 (5423): 2137–2147. doi:10.1126/science.284.5423.2137. PMID 10381873.
  3. ^ Farlow JA (1993). "On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large, carnivorous dinosaurs". In Dodson, Peter; Gingerich, Philip (eds.). Functional Morphology and Evolution. American Journal of Science, Special Volume. Vol. 293-A. pp. 167–199.
  4. ^ Peczkis, J. (1994). "Implications of body-mass estimates for dinosaurs". Journal of Vertebrate Paleontology. 14 (4): 520–33. doi:10.1080/02724634.1995.10011575.
  5. ^ "Anatomy and evolution". National Museum of Natural History. Archived from the original on 2007-11-11. Retrieved 2007-11-21.
  6. ^ Paul, Gregory S. (2010). Princeton Field Guide to Dinosaurs. Princeton University Press. ISBN 978-0-691-13720-9.
  7. ^ Strauss, Bob."Why Were Dinosaurs So Big? The Facts and Theories Behind Dinosaur Gigantism". About EducationArchived 2014-10-06 at the Wayback Machine
  8. ^ a b c Paul, Gregory S. (2016). The Princeton Field Guide to Dinosaurs: 2nd Edition. United States of America: Princeton University Press. ISBN 978-0-691-16766-4.
  9. ^ Paul, G.S. (1997). "Dinosaur models: the good, the bad, and using them to estimate the mass of dinosaurs" (PDF). In Wolberg, D.L.; Stump, E.; Rosenberg, G.D. (eds.). DinoFest International Proceedings. Dinofest International. The Academy of Natural Sciences. pp. 129–154. Archived (PDF) from the original on March 4, 2016.
  10. ^ Woodruff, C; Foster, JR (2015). "The fragile legacy of Amphicoelias fragillimus (Dinosauria: Sauropoda; Morrison Formation - Latest Jurassic)". PeerJ PrePrints. doi:10.7287/peerj.preprints.838v1.
  11. ^ Carpenter, Kenneth (2018-10-19). "Maraapunisaurus fragillimus, N.G. (formerly Amphicoelias fragillimus), a basal Rebbachisaurid from the Morrison Formation (Upper Jurassic) of Colorado". Geology of the Intermountain West. 5: 227–244. doi:10.31711/giw.v5.pp227-244. ISSN 2380-7601.
  12. ^ Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID 210840060.
  13. ^ Mortimer, M. (2001), "Re: Bruhathkayosaurus", discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.
  14. ^ Mortimer, M. (2004), "Re: Largest Dinosaurs" Archived 2019-09-13 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 7 September 2004. Accessed 23 May 2008.
  15. ^ Mortimer, M. (2001), "Titanosaurs too large?" Archived 2016-03-03 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 12 September 2001. Accessed 23 May 2008.
  16. ^ Pal, Saurabh; Ayyasami, Krishnan (27 June 2022). "The lost titan of Cauvery". Geology Today. 38 (3): 112–116. doi:10.1111/gto.12390. ISSN 0266-6979. S2CID 250056201.
  17. ^ a b c Paul, Gregory S.; Larramendi, Asier (11 April 2023). "Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales". Lethaia. 56 (2): 1–11. doi:10.18261/let.56.2.5. ISSN 0024-1164. S2CID 259782734.
  18. ^ Bates, Karl T.; Falkingham, Peter L.; Macaulay, Sophie; Brassey, Charlotte; Maidment, Susannah C.R. (2015). "Downsizing a giant: re-evaluating Dreadnoughtus body mass". Biol Lett. 11 (6): 20150215. doi:10.1098/rsbl.2015.0215. PMC 4528471. PMID 26063751.
  19. ^ Larramendi, A. (2016). "Shoulder height, body mass and shape of proboscideans" (PDF). Acta Palaeontologica Polonica. 61. doi:10.4202/app.00136.2014. S2CID 2092950. Archived (PDF) from the original on 2016-08-24.
  20. ^ Sander, P. Martin; Christian, Andreas; Clauss, Marcus; Fechner, Regina; Gee, Carole T.; Griebeler, Eva-Maria; Gunga, Hanns-Christian; Hummel, Jürgen; Mallison, Heinrich; et al. (2011). "Biology of the sauropod dinosaurs: the evolution of gigantism". Biological Reviews. 86 (1): 117–155. doi:10.1111/j.1469-185X.2010.00137.x. PMC 3045712. PMID 21251189.
  21. ^ Carpenter, K. (2006). "Biggest of the big: a critical re-evaluation of the mega-sauropod Amphicoelias fragillimus." In Foster, J.R. and Lucas, S.G., eds., 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36: 131–138.
  22. ^ Colbert, Edwin Harris (1971). Men and dinosaurs: the search in field and laboratory. Harmondsworth [Eng.]: Penguin. ISBN 978-0-14-021288-4.
  23. ^ Mazzetta, G.V.; et al. (2004). "Giants and Bizarres: Body Size of Some Southern South American Cretaceous Dinosaurs". Historical Biology. 16 (2–4): 1–13. CiteSeerX 10.1.1.694.1650. doi:10.1080/08912960410001715132. S2CID 56028251.
  24. ^ Janensch, W. (1950). "The Skeleton Reconstruction of Brachiosaurus brancai": 97–103. {{cite journal}}: Cite journal requires |journal= (help)
  25. ^ "The World of Dinosaurs". Museum für Naturkunde. Archived from the original on 2018-11-16. Retrieved 2018-11-16.
  26. ^ Lucas, H.; Hecket, H. (2004). "Reappraisal of Seismosaurus, a Late Jurassic Sauropod". Proceeding, Annual Meeting of the Society of Paleontology. 36 (5): 422.
  27. ^ a b c Molina-Pérez, Rubén; Larramendi, Asier (2020). Dinosaur Facts and Figures The Sauropods and Other Sauropodomorphs. Translated by Donaghey, Joan. Illustrated by Andrey Atuchin and Sante Mazzei. Princeton: Princeton University Press. ISBN 978-0-691-20297-6. OCLC 1157079384.
  28. ^ "SVP 2021 Symposium". Society of Vertebrate Paleontology. Retrieved 2023-12-16.
  29. ^ Morgan, James (2014-05-17). "'Biggest dinosaur ever' discovered". BBC News. Archived from the original on 2017-02-18. Retrieved 2017-03-21.
  30. ^ "Don't believe the hype: Patagotitan was not bigger than Argentinosaurus". Sauropod Vertebra Picture of the Week. 2017-08-09. Archived from the original on 2019-04-14. Retrieved 2019-02-19.
  31. ^ Paul, Gregory (December 2019). "Determining the Largest Known Land Animal: A Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals" (PDF). Annals of Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. ISSN 0097-4463. S2CID 210840060.
  32. ^ Patricia Vickers Rich, Thomas Hewitt Rich, Mildred Adams Fenton, Carroll Lane (January 15, 2020). The Fossil Book: A Record of Prehistoric Life. Dover Publications. p. 444. ISBN 9780486838557. Retrieved 2022-08-25.{{cite book}}: CS1 maint: multiple names: authors list (link)
  33. ^ Lu, J., Li, T., Zhong, S., Azuma, Y., Fujita, M., Dong, Z., and Ji, Q. (2007). "New yunnanosaurid dinosaur (Dinosauria, Prosauropoda) from the Middle Jurassic Zhanghe Formation of Yuanmou, Yunnan Province of China". Memoir of the Fukui Prefectural Dinosaur Museum, 6: 1-15.
  34. ^ Therrien, F.; Henderson, D. M. (2007). "My theropod is bigger than yours...or not: estimating body size from skull length in theropods". Journal of Vertebrate Paleontology. 27 (1): 108–115. doi:10.1671/0272-4634(2007)27[108:MTIBTY]2.0.CO;2. S2CID 86025320.
  35. ^ "See what African Wildlife Foundation is doing to protect these iconic flightless birds". 2013-02-25. Archived from the original on 2015-04-29. Retrieved 2015-04-18.
  36. ^ Xu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." Chinese Science Bulletin, 6 pages, accepted November 15, 2008.
  37. ^ Pei, Rui; Li, Quanguo; Meng, Qingjin; Norell, Mark A.; Gao, Ke-Qin (2017). "New Specimens ofAnchiornis huxleyi(Theropoda: Paraves) from the Late Jurassic of Northeastern China". Bulletin of the American Museum of Natural History. 411: 1–67. doi:10.1206/0003-0090-411.1.1. ISSN 0003-0090. S2CID 90650697.
  38. ^ Pascal Godefroit; Andrea Cau; Hu Dong-Yu; François Escuillié; Wu Wenhao; Gareth Dyke (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds". Nature. 498 (7454): 359–62. Bibcode:2013Natur.498..359G. doi:10.1038/nature12168. PMID 23719374. S2CID 4364892.
  39. ^ Which was the smallest dinosaur? Archived 2011-07-06 at the Wayback Machine Royal Tyrrell Museum. Last accessed 22 September 2022.
  40. ^ Averianov AO, Lopatin AV (2022). "A re-appraisal of Parvicursor remotus from the Late Cretaceous of Mongolia: implications for the phylogeny and taxonomy of alvarezsaurid theropod dinosaurs". Journal of Systematic Palaeontology. 19 (16): 1097–1128. doi:10.1080/14772019.2021.2013965. S2CID 247222017.
  41. ^ Conservation International (Content Partner); Mark McGinley (Topic Editor). 2008. "Biological diversity in the Caribbean Islands." In: Encyclopedia of Earth. Eds. Cutler J. Cleveland (Washington, D.C.: Environmental Information Coalition, National Council for Science and the Environment). [First published in the Encyclopedia of Earth May 3, 2007; Last revised August 22, 2008; Retrieved November 9, 2009]. Archived 2013-05-23 at the Wayback Machine>
  42. ^ Glick, Adrienne "Mellisuga helenae" Animal Diversity Web, Retrieved 04 October 2023
  43. ^ Borenstein, Seth (July 31, 2014). "Study traces dinosaur evolution into early birds". AP News. Archived from the original on August 8, 2014. Retrieved August 3, 2014.
  44. ^ Zoe Gough (31 July 2014). "Dinosaurs 'shrank' regularly to become birds". BBC. Archived from the original on 20 February 2015. Retrieved 20 April 2015.