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| location_ts = Cliff and foreshore exposures between [[Seven Rock Point]] and [[Golden Cap]], Dorset
| location_ts = Cliff and foreshore exposures between [[Seven Rock Point]] and [[Golden Cap]], Dorset
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The '''Charmouth Mudstone Formation''' is a [[Formation (geology)|geological formation]] in [[England]]. It preserves [[fossils]] dating back to the early part of the [[Jurassic]] [[Period (geology)|period]] ([[Sinemurian]]–[[Pliensbachian]]).<ref name="BGS_Lexicon">{{Cite web |url=http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=CHAM |title=Charmouth Mudstone Formation |last=British Geological Survey |author-link=British Geological Survey |website=BGS Lexicon of Named Rock Units |access-date=2018-03-08}}</ref> It forms part of the lower [[Lias Group]]. It is most prominently exposed at its type locality in cliff section between [[Lyme Regis]] and [[Charmouth]] (alongside the underlying [[Blue Lias]]) but onshore it extends northwards to [[Market Weighton]], [[Yorkshire]], and in the [[Bedrock|subsurface]] of the East Midlands Shelf and [[Wessex Basin]]. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including [[Mary Anning]].
The '''Charmouth Mudstone Formation''' is a [[Formation (geology)|geological formation]] in [[England]], dating to the Early [[Jurassic]] ([[Sinemurian]]–[[Pliensbachian]]).<ref name="BGS_Lexicon">{{Cite web |url=http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=CHAM |title=Charmouth Mudstone Formation |last=British Geological Survey |author-link=British Geological Survey |website=BGS Lexicon of Named Rock Units |access-date=2018-03-08}}</ref> It forms part of the lower [[Lias Group]]. It is most prominently exposed at its type locality in cliff section between [[Lyme Regis]] and [[Charmouth]] (alongside the underlying [[Blue Lias]]) but onshore it extends northwards to [[Market Weighton]], [[Yorkshire]], and in the [[Bedrock|subsurface]] of the East Midlands Shelf and [[Wessex Basin]]. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including [[Mary Anning]].


== Stratigraphy ==
== Stratigraphy ==
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=== Black Ven Marl Member ===
=== Black Ven Marl Member ===
The Black Ven Marl Member is around 43 metres thick consists of thinly bedded dark mudstones,<ref name="BGS">{{Cite web|title=Black Ven Marl Member|url=http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=BVM|website=The BGS Lexicon of Named Rock Units|publisher=[[British Geological Survey]]}}</ref> with several laterally persistent [[Calcilutite|cementstone]] horizons, notable horizons include the Lower and Upper Cement beds and the Stellare nodules.
The [[Black Ven]] Marl Member is around 43 metres thick consists of thinly bedded dark mudstones,<ref name="BGS">{{Cite web|title=Black Ven Marl Member|url=http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=BVM|website=The BGS Lexicon of Named Rock Units|publisher=[[British Geological Survey]]}}</ref> with several laterally persistent [[Calcilutite|cementstone]] horizons, notable horizons include the Lower and Upper Cement beds and the Stellare nodules.


=== Belemnite Marl Member ===
=== Belemnite Marl Member ===
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== Paleobiota ==
== Paleobiota ==
[[File:CharmouthMudstone NT.jpg|thumb|500px|Vertebrate fauna of the Charmouth Mudstone Formation, by Nobu Tamura, 2020. Click through for original file.|left]]
[[File:CharmouthMudstone NT.jpg|thumb|500px|Vertebrate fauna of the Charmouth Mudstone Formation|left]]
{{clearboth}}
{{clearboth}}


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|[[File:Echioceras_cf._raricostatum_Lyme_Regis.png|center|frameless]]
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|''[[Eoderoceras]]''
|''[[Eoderoceras]]''
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|''[[Gemmellaroceras]]''
|''[[Gemmellaroceras]]''
|''G. cf. peregrinum''
|''G. cf. peregrinum''
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|''[[Uptonia]]''
|''U. bronni''
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|[[File:Uptonia_cf._bronni_Lyme_Regis.png|center|frameless]]
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|''[[Xipheroceras]]''
|''[[Xipheroceras]]''
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''[[Archaeonectrus]]''<ref>R. Owen. 1865. Part I. Sauropterygia. Monographs on the fossil Reptilia of the Liassic formations 17(75):1-40</ref>
''[[Archaeonectrus]]''<ref>{{Cite journal |last=Owen |date=1865 |title=Monograph of the Fossil Reptilia of the Liassic Formations. Part First. Sauropterygia |url=http://dx.doi.org/10.1080/02693445.1865.12027939 |journal=Monographs of the Palaeontographical Society |volume=17 |issue=75 |pages=1–40 |doi=10.1080/02693445.1865.12027939 |bibcode=1865MPalS..17....1O |issn=0269-3445}}</ref>
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''Archaeonectrus rostratus''
''A. rostratus''
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Charmouth
Charmouth
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''[[Attenborosaurus]]''<ref>W. J. Sollas. 1881. On a new species of Plesiosaurus (P. Conybeari) from the Lower Lias of Charmouth; with observations on P megacephalus, Stutchbury, and P. brachycephalus, Owen. Quarterly Journal of the Geological Society of London 37:440-480</ref>
''[[Attenborosaurus]]''<ref>{{Cite journal |last=Sollas |first=W. J. |date=1881 |title=On a new Species of ''Plesiosaurus (P. Conybeari)'' from the Lower Lias of Charmouth; with Observations on P. ''megacephalus'', Stutchbury, and P. ''brachycephalus'', Owen |url=http://dx.doi.org/10.1144/gsl.jgs.1881.037.01-04.42 |journal=Quarterly Journal of the Geological Society |volume=37 |issue=1–4 |pages=440–481 |doi=10.1144/gsl.jgs.1881.037.01-04.42 |issn=0370-291X}}</ref>
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''Attenborosaurus conybeari''
''A. conybeari''
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*Black Ven Water, Charmouth
*Black Ven Water, Charmouth
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*BMNH 40140, partial skeleton and skull
*BMNH 40140, partial skeleton and skull
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A marine sauropterygian, early member of the family [[Pliosauridae]]. It was identified as ''Plesiosaurus conybeari''.
A marine sauropterygian, early member of the family [[Pliosauridae]]. It was first identified as ''Plesiosaurus conybeari''.
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[[File:Hul - Attenborosaurus conybeari - 2.jpg|center|thumb|Cast of the holotype]]
[[File:Hul - Attenborosaurus conybeari - 2.jpg|center|thumb|Cast of the holotype]]
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''?[[Plesiosaurus]]''<ref>G. W. Storrs. 1995. A juvenile specimen of ? Plesiosaurus sp. from the Lias (Lower Jurassic, Pliensbachian) near Charmouth, Dorset, England. Proceedings of the Dorsal Archaeological and Natural History Society 116:71-76</ref>
''?[[Plesiosaurus]]''<ref>{{Cite journal |last=Storrs |first=G. W. |date=1995 |title=A juvenile specimen of ? Plesiosaurus sp. from the Lias (Lower Jurassic, Pliensbachian) near Charmouth, Dorset, England |url=https://www.researchgate.net/publication/259822957 |journal=Proceedings of the Dorsal Archaeological and Natural History Society |volume=116 |issue=2 |pages=71–76}}</ref>
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''? Plesiosaurus sp.''
''? P. sp.''
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*Eastern point of Wear Cliffs, below Golden Cap
*Eastern point of Wear Cliffs, below Golden Cap
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*BRSMG Ce17972a-o, articulated juvenile partial postcranial skeleton
*BRSMG Ce17972a-o, articulated juvenile partial postcranial skeleton
|style="background:#E6E6E6;"|
|style="background:#E6E6E6;"|
A marine sauropterygian, type member of the family [[Plesiosauroidea]] inside [[Plesiosauria]]. A Juvenile specimen that resembles those assigned to the genus ''Plesiosaurus''. Its assignation is dubious.
A marine sauropterygian, type member of the family [[Plesiosauroidea]] inside [[Plesiosauria]]. A juvenile specimen that resembles those assigned to the genus ''Plesiosaurus''. Its assignation is dubious.
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''[[Plesiosaurus]]''<ref>W. D. Conybeare. 1824. On the discovery of an almost perfect skeleton of the Plesiosaurus. Transactions of the Geological Society of London, series 2 1:381-389</ref>
''[[Plesiosaurus]]''<ref>{{Cite journal |last=CONYBEARE |first=W. D. |date=1824 |title=XXI.—On the Discovery of an almost perfect Skeleton of the Plesiosaurus. |url=http://dx.doi.org/10.1144/transgslb.1.2.381 |journal=Transactions of the Geological Society of London |volume=1 |issue=2 |pages=381–389 |doi=10.1144/transgslb.1.2.381 |issn=2042-5295}}</ref>
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''Plesiosaurus dolichodeirus''
''P. dolichodeirus''
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*Lyme Regis
*Lyme Regis
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|[[File:Plesiosaurus dolichodeirus NHM.jpg|center|frameless]]
|[[File:Plesiosaurus dolichodeirus NHM.jpg|center|frameless]]
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|''[[Thaumatodracon]]''<ref>{{Cite journal|last1=Smith|first1=Adam S.|last2=Araújo|first2=Ricardo|date=2017-07-21|title=Thaumatodracon wiedenrothi, a morphometrically and stratigraphically intermediate new rhomaleosaurid plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis|url=http://dx.doi.org/10.1127/pala/308/2017/89|journal=Palaeontographica Abteilung A|volume=308|issue=4–6|pages=89–125|doi=10.1127/pala/308/2017/89|issn=0375-0442}}</ref>
|''[[Thaumatodracon]]''<ref>{{Cite journal|last1=Smith|first1=Adam S.|last2=Araújo|first2=Ricardo|date=2017-07-21|title=Thaumatodracon wiedenrothi, a morphometrically and stratigraphically intermediate new rhomaleosaurid plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis|url=http://dx.doi.org/10.1127/pala/308/2017/89|journal=Palaeontographica Abteilung A|volume=308|issue=4–6|pages=89–125|doi=10.1127/pala/308/2017/89|bibcode=2017PalAA.308...89S |issn=0375-0442}}</ref>
|''T. wiedenrothi''
|''T. wiedenrothi''
|Between Lyme Regis and Charmouth
|Between Lyme Regis and Charmouth
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!Notes
!Notes
!Images
!Images
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|[[Ichthyosauria]] indet.
|Indeterminate
|Lyme Regis, Charmouth
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|Indeterminate remains
|Un-attributable to any genus
|[[File:Ichthyosaur rib fragments.png|frameless]]
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''[[Ichthyosaurus]]''
''[[Ichthyosaurus]]''
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''Ichthyosaurus anningae''<ref>{{Cite journal|last1=Lomax|first1=Dean R.|last2=Massare|first2=Judy A.|date=2015-03-04|title=A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K.|journal=Journal of Vertebrate Paleontology|language=en|volume=35|issue=2|pages=e903260|doi=10.1080/02724634.2014.903260|s2cid=85745787|issn=0272-4634|url=http://osf.io/89nhw}}</ref>
''I. anningae''<ref>{{Cite journal|last1=Lomax|first1=Dean R.|last2=Massare|first2=Judy A.|date=2015-03-04|title=A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K.|journal=Journal of Vertebrate Paleontology|language=en|volume=35|issue=2|pages=e903260|doi=10.1080/02724634.2014.903260|bibcode=2015JVPal..35E3260L |s2cid=85745787|issn=0272-4634|url=http://osf.io/89nhw}}</ref>
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Charmouth
Charmouth
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DONMG:1983.98 Subadult specimen
DONMG:1983.98 Subadult specimen
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An [[Ichthyosauridae|Ichthyosaurid]] [[Ichthyosaur]]
An [[ichthyosaurid]] [[ichthyosaur]]
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[[File:Ichthyosaurus anningae trio NT small.jpg|center|thumb]]
[[File:Ichthyosaurus anningae trio NT small.jpg|center|frameless]]
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Stonebarrow Marls Member
Stonebarrow Marls Member
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NHMUK R15907, partial skeleton<ref>{{Cite journal|last1=Bennett|first1=S.P.|last2=Barrett|first2=P.M.|last3=Collinson|first3=M.E.|last4=Moore-Fay|first4=S.|last5=Davis|first5=P.G.|last6=Palmer|first6=C.P.|date=January 2012|title=A new specimen of Ichthyosaurus communis from Dorset, UK, and its bearing on the stratigraphical range of the species|journal=Proceedings of the Geologists' Association|language=en|volume=123|issue=1|pages=146–154|doi=10.1016/j.pgeola.2011.07.001}}</ref>
NHMUK R15907, partial skeleton<ref>{{Cite journal|last1=Bennett|first1=S.P.|last2=Barrett|first2=P.M.|last3=Collinson|first3=M.E.|last4=Moore-Fay|first4=S.|last5=Davis|first5=P.G.|last6=Palmer|first6=C.P.|date=January 2012|title=A new specimen of Ichthyosaurus communis from Dorset, UK, and its bearing on the stratigraphical range of the species|journal=Proceedings of the Geologists' Association|language=en|volume=123|issue=1|pages=146–154|doi=10.1016/j.pgeola.2011.07.001|bibcode=2012PrGA..123..146B }}</ref>
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Considered to belong to ''I. communis'' by Bennett et al. 2012, considered indeterminate within ''Ichthyosaurus'' by Lomax and Massare 2015
Considered to belong to ''I. communis'' by Bennett et al. 2012, considered indeterminate within ''Ichthyosaurus'' by Lomax and Massare 2015
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''[[Leptonectes]]''
''[[Leptonectes]]''
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''Leptonectes moorei''<ref>{{Cite journal|last1=McGowan|first1=Christopher|last2=Milner|first2=Angela C.|date=October 1999|title=A new Pliensbachian ichthyosaur from Dorset, England|journal=Palaeontology|language=en|volume=42|issue=5|pages=761–768|doi=10.1111/1475-4983.00096|issn=0031-0239|doi-access=free}}</ref>
''L. moorei''<ref>{{Cite journal|last1=McGowan|first1=Christopher|last2=Milner|first2=Angela C.|date=October 1999|title=A new Pliensbachian ichthyosaur from Dorset, England|journal=Palaeontology|language=en|volume=42|issue=5|pages=761–768|doi=10.1111/1475-4983.00096|issn=0031-0239|doi-access=free|bibcode=1999Palgy..42..761M }}</ref>
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Seatown
Seatown
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BMNH R14370 partial anterior skeleton with most complete skull
BMNH R14370 partial anterior skeleton with most complete skull
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A [[leptonectid]] ichthyosaur
[[Leptonectidae|Leptonectid]] Ichthyosaur
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''Leptonectes solei''<ref>{{Cite journal|last1=Lomax|first1=Dean R.|last2=Massare|first2=Judy A.|date=December 2018|title=A forefin of Leptonectes solei from the Lower Jurassic (Pliensbachian) of Dorset, UK|journal=Proceedings of the Geologists' Association|language=en|volume=129|issue=6|pages=770–773|doi=10.1016/j.pgeola.2018.07.005|url=https://www.research.manchester.ac.uk/portal/en/publications/a-forefin-of-leptonectes-solei-from-the-lower-jurassic-pliensbachian-of-dorset-uk(569a32a2-dbcf-40d9-9055-4d16351f6c19).html}}</ref>
''L. solei''<ref>{{Cite journal|last1=Lomax|first1=Dean R.|last2=Massare|first2=Judy A.|date=December 2018|title=A forefin of Leptonectes solei from the Lower Jurassic (Pliensbachian) of Dorset, UK|journal=Proceedings of the Geologists' Association|language=en|volume=129|issue=6|pages=770–773|doi=10.1016/j.pgeola.2018.07.005|bibcode=2018PrGA..129..770L |s2cid=134397018 |url=https://www.research.manchester.ac.uk/portal/en/publications/a-forefin-of-leptonectes-solei-from-the-lower-jurassic-pliensbachian-of-dorset-uk(569a32a2-dbcf-40d9-9055-4d16351f6c19).html}}</ref>
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Seatown
Seatown
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|''[[Temnodontosaurus]]''
|''T. platyodon''<ref>{{Cite web |title= PBDB|url=https://paleobiodb.org/classic/basicCollectionSearch?collection_no=98103}}</ref>
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|[[File:Temnodontosaurus plat1DB.jpg|thumb|''Temnodontosaurus platyodon''.]]
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=== Thalattosuchia ===
{| class="wikitable" width="100%" align="center"
! colspan="7" align="center" |'''Thalattosuchia of the Charmouth Mudstone Formation'''
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!Genus
!Species
!Location
!Stratigraphic position
!Material
!Notes
!Images
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| rowspan="2" |[[Thalattosuchia]]<ref>{{Cite web |last1=Young |first1=Mark T |last2=Dufeau |first2=David |last3=Bowman |first3=Charlotte |last4=Cowgill |first4=Thomas |last5=Schwab |first5=Julia A |last6=Witmer |first6=Lawrence M |last7=Herrera |first7=Yanina |last8=Katsamenis |first8=Orestis L |last9=Steel |first9=Lorna |date=2024 |title=Thalattosuchian crocodylomorphs from the Sinemurian (Early Jurassic) of the UK |url=https://academic.oup.com/zoolinnean/article-abstract/201/3/zlae079/7713265 |access-date=2024-07-15 |website=academic.oup.com |publisher=Zoological Journal of the Linnean Society}}</ref>
| rowspan="2" |Indeterminate
| rowspan="2" |
* Stonebarrow beach, Charmouth
* Conesby Quarry, Scunthorpe
| rowspan="2" |Black Ven Mudstone Member
| rowspan="2" |
* NHMUK PV R 36710, incomplete snout
* NHMUK PV R 9731, axis and four post-axial cervical vertebrae. Two incomplete axial ribs, the lef fourth cervical rib, and two cervical rib fragments
* NHMUK PV R 36 711, dorsal vertebra from the caudal end of the dorsal series, and the neural arch of another dorsal vertebra, both held together by matrix
| rowspan="2" |Early [[Thalattosuchia|Thalattosaurian]] specimens that can't be referred to any concrete genus. Includes a possible early diverging metriorhynchoid
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''[[Turnersuchus]]''<ref>{{Cite journal |last1=Wilberg |first1=E. W. |last2=Godoy |first2=P. L. |last3=Griffiths |first3=E. F. |last4=Turner |first4=A. H. |last5=Benson |first5=R. B. J. |year=2023 |title=A new early diverging thalattosuchian (Crocodylomorpha) from the Early Jurassic (Pliensbachian) of Dorset, U.K. and implications for the origin and evolution of the group |journal=Journal of Vertebrate Paleontology |volume=42 |issue=3 |at=e2161909 |doi=10.1080/02724634.2022.2161909 |s2cid=256149424 |url=https://zenodo.org/record/7594138 }}</ref>
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''T. hingleyae''
|1 kilometer west of the Charmouth Heritage Coast Centre
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Belemnite Marl Member
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LRM 2021/45, a partial skeleton including cranial material, cervical, dorsal, and caudal vertebrae, ribs, right pectoral girdle, partial limb bones, and an osteoderm
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An early diverging [[thalattosuchia]]n.
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[[File:Turnersuchus_holotype.jpg|thumb]]
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''[[Dimorphodon]]''<ref>{{Cite journal |last=Sangster |first=Sarah |date=2021-09-02 |title=THE OSTEOLOGY OF ''DIMORPHODON MACRONYX'', A NON-PTERODACTYLOID PTEROSAUR FROM THE LOWER JURASSIC OF DORSET, ENGLAND |url=http://dx.doi.org/10.1080/02693445.2021.2037868 |journal=Monographs of the Palaeontographical Society |volume=175 |issue=661 |pages=1–48 |doi=10.1080/02693445.2021.2037868 |bibcode=2021MPalS.175....1S |issn=0269-3445}}</ref>
''[[Dimorphodon]]''<ref>Martill, D. M. (2014). Dimorphodon and the Reverend George Howman's noctivagous flying dragon: the earliest restoration of a pterosaur in its natural habitat. Proceedings of the Geologists' Association, 125(1), 120-130.</ref><ref name="Steel, L. 2012">Steel, L. (2012). The pterosaur collection at the Natural History Museum, London, UK: an overview and list of specimens, with description of recent curatorial developments. Acta Geologica Sinica‐English Edition, 86(6), 1340-1355.</ref><ref name=Un2>Unwin, D. M. (2003). On the phylogeny and evolutionary history of pterosaurs. Geological Society, London, Special Publications, 217(1), 139-190.</ref>
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''Dimorphodon macronyx''
''D. macronyx''
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*Charmouth
*Charmouth
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*Semi-Complete Skeletons
*Semi-Complete Skeletons
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A basal pterosaur, type member of the family [[Dimorphodontidae]].
A basal pterosaur and the type member of the family [[Dimorphodontidae]].
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[[File:Dimorphodon.png|center|thumb]]
[[File:Dimorphodon.png|center|thumb]]
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[[Dimorphodontidae]]<ref name="Un2">{{Cite journal |last=Unwin |first=David M. |date=2003 |title=On the phylogeny and evolutionary history of pterosaurs |url=http://dx.doi.org/10.1144/gsl.sp.2003.217.01.11 |journal=Geological Society, London, Special Publications |volume=217 |issue=1 |pages=139–190 |doi=10.1144/gsl.sp.2003.217.01.11 |bibcode=2003GSLSP.217..139U |issn=0305-8719}}</ref><ref>{{Cite journal |last1=Butler |first1=Richard J. |last2=Barrett |first2=Paul M. |last3=Nowbath |first3=Stephen |last4=Upchurch |first4=Paul |date=2009 |title=Estimating the effects of sampling biases on pterosaur diversity patterns: implications for hypotheses of bird/pterosaur competitive replacement |url=http://dx.doi.org/10.1666/0094-8373-35.3.432 |journal=Paleobiology |volume=35 |issue=3 |pages=432–446 |doi=10.1666/0094-8373-35.3.432 |bibcode=2009Pbio...35..432B |issn=0094-8373}}</ref>
[[Dimorphodontidae]]<ref name=Un2/><ref>Barrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. Zitteliana, 61-107.</ref>
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Indeterminate
Indeterminate
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*NHMUK PV R1595, wing phalanges
*NHMUK PV R1595, wing phalanges
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A basal pterosaur, member of the family [[Dimorphodontidae]]. Known as "Charmouth dimorphodontid" it shares resemblance with the genus ''[[Dimorphodon]]''.
A basal [[dimorphodontid]] pterosaur. Known as "Charmouth dimorphodontid" it shares resemblance with the genus ''[[Dimorphodon]]''.
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[[Dimorphodontidae]]<ref name=Und>Unwin, D. M. (2011): A new dimorphodontid pterosaur from the Lower Jurassic of Dorset, southern England. 59th Annual Symposium of Vertebrate Palaeontology and Comparative AnatomyLyme Regis, Dorset, UK</ref>
Indeterminate<ref name="Und">{{Cite journal |last=Unwin |first=D. M. |date=2011 |title=A new dimorphodontid pterosaur from the Lower Jurassic of Dorset, southern England |journal=59th Annual Symposium of Vertebrate Palaeontology and Comparative AnatomyLyme Regis, Dorset, UK}}</ref>
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Indeterminate
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*Near Seatown
*Near Seatown
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Associated remains of the rostrum and mandibles
Associated remains of the rostrum and mandibles
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A basal Pterosaur, member of the family [[Dimorphodontidae]]. The relatively deep rostrum and extreme size dimorphism in the dentition show clear similarities to ''Dimorphodon'', also from the Lias of Dorset. The distinctive dentition, in which the first four pairs of rostral teeth and two pairs of mandibular teeth are relatively large and fang-like, while the remaining teeth are remarkably small and short.<ref name=Und/>
A basal [[dimorphodontid]] pterosaur. The relatively deep rostrum and extreme size dimorphism in the dentition show clear similarities to ''Dimorphodon'', also from the Lias of Dorset. The distinctive dentition, in which the first four pairs of rostral teeth and two pairs of mandibular teeth are relatively large and fang-like, while the remaining teeth are remarkably small and short.<ref name=Und/>
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!Images
!Images
|-
|-
| style="background:#fbdddb;" |
|rowspan="3" |
[[Coelophysidae]]<ref>{{Cite book |last=Lydekker |first=Richard |url=http://dx.doi.org/10.5962/bhl.title.61848 |title=Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural history) ... By Richard Lydekker ... |date=1888 |publisher=Printed by order of the Trustees |location=London|doi=10.5962/bhl.title.61848 }}</ref><ref name="Coelo">{{Cite journal |last1=Carrano |first1=Matthew T. |last2=Sampson |first2=Scott D. |date=2004-09-27 |title=A review of coelophysoids (Dinosauria: Theropoda) from the Early Jurassic of Europe, with comments on the late history of the Coelophysoidea |url=http://dx.doi.org/10.1127/njgpm/2004/2004/537 |journal=Neues Jahrbuch für Geologie und Paläontologie - Monatshefte |volume=2004 |issue=9 |pages=537–558 |doi=10.1127/njgpm/2004/2004/537 |issn=0028-3630}}</ref>
''[[Scelidosaurus]]''<ref name=Scelido>B.H. Newman The Jurassic dinosaur Scelidosaurus harrisoni, Owen Palaeontology, 11 (1968), pp. 40-43</ref><ref>Charig AJ, Newman BH. 1992. Scelidosaurus harrisonii Owen, 1861 (Reptilia, Ornithischia): proposed replacement of inappropriate lectotype. Bulletin of Zoological Nomenclature 49: 280–283</ref><ref>David B Norman, FLS, Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: cranial anatomy, Zoological Journal of the Linnean Society, Volume 188, Issue 1, January 2020, Pages 1–81, [https://doi.org/10.1093/zoolinnean/zlz074]</ref>
| style="background:#fbdddb;" |
''"Megalosaurus" lydekkeri''
| style="background:#fbdddb;" |
Lyme Regis
| style="background:#fbdddb;" |
Black Ven Marl Member
| style="background:#fbdddb;" |
*NHMUK OR 41352, isolated tooth
| style="background:#fbdddb;" |
A theropod, possibly a member of [[Coelophysidae]] inside [[Neotheropoda]]. Invalid and non diagnostic. It was identified as ''[[Zanclodon]](?) sp.'' b., ''Megalosaurus lydekkeri'' and reassigned as ''[[Magnosaurus]](?) lydekkeri''.<ref name="Mor2">{{Cite web |last=Mortimer |first=Mickey |date=2012-11-24 |title=The Theropod Database Blog: "New" name for Megalosaurus? lydekkeri |url=https://theropoddatabase.blogspot.com/2012/11/new-name-for-megalosaurus-lydekkeri.html |access-date=2024-04-29 |website=The Theropod Database Blog}}</ref> Mortimer found that was changed to ''"Megalosaurus" woodwardi'' later.<ref name="Mor2" /> It was found on mostly of recent works to be non diagnostic, probably a coelophysoid.<ref name="Coelo" />
| style="background:#fbdddb;" |
|-
|
''[[Dornraptor]]''<ref name=":1">{{Cite journal |last=Baron |first=Matthew G. |date=2024-04-29 |title=A new name for old bones: A reassessment of Early Jurassic theropod remains from Dorset, England |url=https://palaeo-electronica.org/content/2024/5067-on-an-early-jurassic-theropod |journal=Palaeontologia Electronica |language=English |volume=27 |issue=1 |pages=1–12 |doi=10.26879/1346 |issn=1094-8074|doi-access=free }}</ref>
|
''D. normani''
|
Lower cliff face, Charmouth
|
Black Ven Marl Member (Likely)
|
*NHMUK OR 39496, Partial hind limb
*GSM 109560, left femur
|
An Averostran theropod, possible Ceratosaur or Tetanuran. In 2024 was formally described, but referred to the older [[Blue Lias]].<ref name=":1" /> A paper non quoted on the description relocated it on the Charmouth mudstone.<ref name="Thero20" />
| [[File:Dornraptor normani.png|center|thumb]]
|-
|
[[Neotheropoda]]<ref name="Thero20">{{Cite journal |last1=Choiniere |first1=Jonah N. |last2=Wills |first2=Simon |last3=Bennett |first3=S. Christopher |last4=Barrett |first4=Paul M. |date=2020 |title=A small theropod dinosaur from the Lower Jurassic Lias Group of Charmouth, Dorset |url=http://dx.doi.org/10.1016/j.pgeola.2020.08.003 |journal=Proceedings of the Geologists' Association |volume=131 |issue=6 |pages=751–757 |doi=10.1016/j.pgeola.2020.08.003 |bibcode=2020PrGA..131..751C |issn=0016-7878}}</ref>
|
Indeterminate
|
Near Lyme Regis
|
obtusum Zone, obtusum subzone
|
*NHMUK PV R36855, Left fibula, ~ 17&nbsp;cm long
|
A theropod, possibly a basal member of [[Neotheropoda]]. Previously misidentified as a referred specimen of the pterosaur ''[[Dimorphodon]] macronyx''.<ref name="Steel, L. 2012">{{Cite journal |last=STEEL |first=Lorna |date=2012 |title=The Pterosaur Collection at the Natural History Museum, London, UK: an Overview and List of Specimens, with Description of Recent Curatorial Developments |url=http://dx.doi.org/10.1111/1755-6724.12004 |journal=Acta Geologica Sinica - English Edition |volume=86 |issue=6 |pages=1340–1355 |doi=10.1111/1755-6724.12004 |issn=1000-9515}}</ref> NHMUK PV R36855 differs from those of all other Early Jurassic-Late Triassic, theropods in its combination of features, but bears no discernible [[Autapomorphy|autapomorphies]].<ref name="Thero20" /> It is similar to ''[[Tawa (dinosaur)|Tawa]]'', ''[[Dracoraptor]]'' and ''[[Liliensternus]]'', being considered an early branching neotheropod, with a size range similar to ''[[Coelophysis]] bauri'', representing an individual of approximately 10&nbsp;kg, approximately the same size as ''[[Coelophysis bauri]]''.<ref name="Thero20" />
|[[File:Charmouth theropod fibula.jpeg|center|frameless]]
|-
|
[[Massopoda]]<ref name="Magno">{{Cite journal |last=Benson |first=Roger B. J. |date=2010-03-15 |title=The osteology of ''Magnosaurus nethercombensis'' (Dinosauria, Theropoda) from the Bajocian (Middle Jurassic) of the United Kingdom and a re-examination of the oldest records of tetanurans |url=http://dx.doi.org/10.1080/14772011003603515 |journal=Journal of Systematic Palaeontology |volume=8 |issue=1 |pages=131–146 |doi=10.1080/14772011003603515 |bibcode=2010JSPal...8..131B |issn=1477-2019}}</ref>
|
Indeterminate
|
lower cliff face, Charmouth
|
obtusum Zone, obtusum subzone
|
*GSM 109561 pedal ungual
|
A sauropodomorph, possibly a member of [[Massopoda]]. While it was not associated with BMNH 39496 or GSM 109560 and was not assigned to ''"Merosaurus"'' it was classified as coming from an indeterminate theropod. Mortimer found closer resemblance with pedal unguals of basal massopods like ''[[Blikanasaurus]]'' and ''[[Jingshanosaurus]]''.<ref name="Mor">{{Cite web |last=Mortimer |first=Mickey |date=2010-04-17 |title=The Theropod Database Blog: Pickering's taxa 4: Merosaurus newmani |url=https://theropoddatabase.blogspot.com/2010/04/pickerings-taxa-4-merosaurus-newmani.html |access-date=2024-04-29 |website=The Theropod Database Blog}}</ref>
|
|-
| rowspan="3" |
''[[Scelidosaurus]]''<ref>{{Cite journal |last=Norman |first=David B |date=2019-11-29 |title=Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: cranial anatomy |url=http://dx.doi.org/10.1093/zoolinnean/zlz074 |journal=Zoological Journal of the Linnean Society |volume=188 |issue=1 |pages=1–81 |doi=10.1093/zoolinnean/zlz074 |issn=0024-4082}}</ref><ref name=":0">{{Cite journal |last=Norman |first=David B |date=2019-12-17 |title=Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: postcranial skeleton |url=http://dx.doi.org/10.1093/zoolinnean/zlz078 |journal=Zoological Journal of the Linnean Society |volume=189 |issue=1 |pages=47–157 |doi=10.1093/zoolinnean/zlz078 |issn=0024-4082}}</ref>
|
|
''Scelidosaurus harrisonii''
''S. harrisonii''
|
|
*Charmouth, Black Ven
*Charmouth, Black Ven
Line 416: Line 538:
*BRSMG Ce12788. cervical osteoderms.
*BRSMG Ce12788. cervical osteoderms.
*BRSMG Ce12789. Vertebra and bone fragments
*BRSMG Ce12789. Vertebra and bone fragments
*BRSMG Cf2781. nine middle-distal caudal vertebrae and some haemal arches with some organic (kerogenized) remains of the skin preserved.
*BRSMG Cf2781. nine middle-distal caudal vertebrae and some haemal arches with some organic (kerogenized) remains of the skin preserved.
*BRSMG LEGL 0004. Articulated skeleton
*BRSMG LEGL 0004. Articulated skeleton
*BRSMG LEGL 0005. Articulated partial skeleton
*BRSMG LEGL 0005. Articulated partial skeleton
*DORCM G.7542. Osteoderms
*DORCM G.7542. Osteoderms
*DORCM G.10817. Osteoderms
*DORCM G.10817. Osteoderms
*CAMSM X39256. Cranial: disarticulated skull and jaw elements
*CAMSM X39256. Cranial: disarticulated skull and jaw elements
|rowspan="3"|
| rowspan="3" |
A basal [[Thyreophoran]], type member of the family [[Scelidosauridae]]. One of the best known armored dinosaurs of the lower Jurassic, also the most complete found to date, with specimens preserving even the skin and dermal armour.<ref>David B Norman, FLS, Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: the dermal skeleton, Zoological Journal of the Linnean Society, Volume 190, Issue 1, September 2020, Pages 1–53, [https://doi.org/10.1093/zoolinnean/zlz085]</ref>
A basal [[thyreophoran]], and the type member of the family [[Scelidosauridae]]. One of the best known armored dinosaurs of the lower Jurassic, also the most complete found to date, with specimens preserving even the skin and dermal armour.<ref name=":0" />
|rowspan="3" |
| rowspan="3" |
[[File:Scelidosaurus harrisonii (1).jpg|center|thumb]]
[[File:Scelidosaurus harrisonii (1).jpg|center|thumb]]
|-
|-
|
|
''S. "sp. A"''
''Scelidosaurus sp. A''<ref>P. C. Ensom. 1989. New scelidosaur remains from the Lower Lias of Dorset. Proceedings of the Dorset Natural History and Archaeological Society 110:166-167</ref>
|
|
Belemnite Marls at Seatown
Belemnite Marls at Seatown
Line 434: Line 556:
*Belemnite Marl Member, ''[[Uptonia jamesoni]] Zone''
*Belemnite Marl Member, ''[[Uptonia jamesoni]] Zone''
|
|
DORCM G.7842. Three dorsal vertebrae in articulation
* DORCM G.7842. Three dorsal vertebrae in articulation
|-
|-
|
|
''Cf. S. sp.''
''Cf. Scelidosaurus sp.''<ref>Anonymous. 1955. British Museum (Natural History). Nature 176(4487):815-816</ref><ref>Maidment, S.C.R., Porro, L.B., 2010, "Homology of the palpebral and origin of the supraorbital ossifications in ornithischian dinosaurs", Lethaia, 43: 95-111</ref><ref>Rixon AE. 1968. The development of the remains of a small
Scelidosaurus from a Lias nodule. Museums Journal 67:
315–321</ref>
|
|
Foot of Black Ven, Charmouth
Foot of Black Ven, Charmouth
Line 454: Line 574:
*NHMUK OR39518. Haemal arch (chevron).
*NHMUK OR39518. Haemal arch (chevron).
*NHMUK OR39519. Two phalanges.
*NHMUK OR39519. Two phalanges.

NHMUK OR39520. Ungual phalanx of ?pes.
* NHMUK OR39520. Ungual phalanx of ?pes.

*NHMUK OR39521. Several bone fragments.
*NHMUK OR39521. Several bone fragments.
*NHMUK OR40503. Tibia (proximal end).
*NHMUK OR40503. Tibia (proximal end).
Line 461: Line 583:
*NHMUK OR41324. Scapular blade
*NHMUK OR41324. Scapular blade
*NHMUK OR41325. Humerus
*NHMUK OR41325. Humerus
*NHMUK OR41326. Fibula
*NHMUK OR41326. Fibula
*NHMUK OR41327. Two caudal vertebrae
*NHMUK OR41327. Two caudal vertebrae
*NHMUK OR41328. Three metatarsals
*NHMUK OR41328. Three metatarsals
*NHMUK OR41329. Osteoderm
*NHMUK OR41329. Osteoderm
*NHMUK OR41330. Ischium
*NHMUK OR41330. Ischium
|-
|style="background:#fbdddb;" |
''"[[Merosaurus]]"''<ref name=Scelido/><ref name=Theropo>R.B.J. Benson, P.M. Barrett Dinosaurs of Dorset: Part 1, the carnivorous dinosaurs (Saurischia, Theropoda) Proceedings of the Dorset Natural History and Archaeological Society, 130 (2009), pp. 133-147</ref><ref>M.T. Carrano, R.B.J. Benson, S.D. Sampson The phylogeny of Tetanurae (Dinosauria: Theropoda) Journal of Systematic Palaeontology, 10 (2012), pp. 211-300</ref>
|style="background:#fbdddb;" |
''"Merosaurus" newmani''
|style="background:#fbdddb;" |
lower cliff face, Charmouth
|style="background:#fbdddb;" |
Black Ven Marl Member
|style="background:#fbdddb;" |
*GSM 109560, Left femur
*NHMUK OR 39496, Partial hind limb
|style="background:#fbdddb;" |
A Theropod, possible basal member of [[Tetanurae]]. Initially considered part of ''Scelidosaurus'' holotype, considered Tetanurae indet by Carrano 2012. It was named "Merosaurus" by Welles, Powell and Pickering in 1995.<ref>Pickering, 1995. Jurassic Park: Unauthorized Jewish Fractals in Philopatry. A Fractal Scaling in Dinosaurology Project, 2nd revised printing. Capitola, California. 478 pp.</ref> But due to lack description was considered invalid.<ref name=Mor>Mortimer, M.(2010): Pickering's taxa 4: Merosaurus newmani. The Theropod Database Blog</ref>
|style="background:#fbdddb;" |
[[File:Merosaurus.jpg|center|thumb]]
|-
|style="background:#fbdddb;" |
''[[Coelophysidae]]''<ref>R. Lydekker Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part I. British Museum (Natural History), London 309 pp
(1888)</ref><ref name=Coelo>M.T. Carrano, S.D. Sampson A review of coelophysoids (Dinosauria: Theropoda) from the Early Jurassic of Europe, with comments on the late history of the Coelophysoidea Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 2004 (2004), pp. 537-558</ref>
|style="background:#fbdddb;" |
''"Megalosaurus" lydekkeri''
|style="background:#fbdddb;" |
Lyme Regis
|style="background:#fbdddb;" |
Black Ven Marl Member
|style="background:#fbdddb;" |
*NHMUK OR 41352, isolated tooth
|style="background:#fbdddb;" |
A Theropod, possible member of [[Coelophysidae]] inside [[Neotheropoda]]. Invalid and non diagnostic. It was identified as ''[[Zanclodon]](?) sp.'' b., ''Megalosaurus lydekkeri'' and reassigned as ''[[Magnosaurus]](?) lydekkeri''.<ref name=Mor2>Mortimer, M.(2012):"New" name for Megalosaurus? lydekkeri. The Theropod Database Blog</ref> Mortimer found that was changed to ''"Megalosaurus" woodwardi'' later.<ref name=Mor2/> It was found on mostly of recent works to be non diagnostic, probably a Coelophysoid.<ref name=Coelo/>
|style="background:#fbdddb;" |
|-
|
[[Neotheropoda]]<ref name=Thero20>Choinierea J. N., Wills, S., Bennett, S.C., Barrett P.M.(2020):A small theropod dinosaur from the Lower Jurassic Lias Group of Charmouth, Dorset. Proceedings of the Geologists' Association. 14 September 2020</ref>
|
Indeterminate
|
Near Lyme Regis
|
obtusum Zone, obtusum subzone
|
*NHMUK PV R36855, Left fibula, ~ 17&nbsp;cm long
|
A theropod, possible basal member of [[Neotheropoda]]. Previously misidentified as a referred specimen of the pterosaur ''[[Dimorphodon]] macronyx''.<ref name="Steel, L. 2012"/> NHMUK PV R36855 differs from those of all other Early Jurassic-Late Triassic, theropods in its combination of features, but bears no discernible [[Autapomorphy|autapomorphies]].<ref name=Thero20/> It is similar to ''[[Tawa (dinosaur)|Tawa]]'', ''[[Dracoraptor]]'' and ''[[Liliensternus]]'', being considered an early branching neotheropod, with a size range similar to ''[[Coelophysis]] bauri'', representing an individual of approximately 10&nbsp;kg, approximately the same size as ''[[Coelophysis bauri]]''.<ref name=Thero20/>
|[[File:Charmouth theropod fibula.jpeg|center|frameless]]
|-
|
[[Massopoda]]<ref name=Magno>Benson, 2010. The osteology of Magnosaurus nethercombensis (Dinosauria, Theropoda) from the Bajocian (Middle Jurassic) of the United Kingdom and a re-examination of the oldest records of tetanurans. Journal of Systematic Palaeontology. 8(1), 131-146.</ref>
|
Indeterminate
|
lower cliff face, Charmouth
|
obtusum Zone, obtusum subzone
|
*GSM 109561 pedal ungual
|
A Sauropodomorph, possible member of [[Massopoda]]. While it was not associated with BMNH 39496 or GSM 109560 and was not assigned to ''"Merosaurus"'' it was classified as coming from an indeterminate theropod. Mortimer found closer resemblance with pedal unguals of basal massopods like [[Blikanasaurus]] and [[Jingshanosaurus]].<ref name=Mor/>
|
|-
|-
|}
|}


=== Fish ===
=== Fish ===
Numerous fish species are known from the Charmouth Mudstone, from such horizons as the "Fish Bed" of the Shales With Beef Member.
Numerous fish species are known from the Charmouth Mudstone and underlying Blue Lias, from such horizons as the "Fish Bed" of the Shales With Beef Member.


{| class="wikitable" width="100%" align="center"
{| class="wikitable" width="100%" align="center"
Line 588: Line 651:
|
|
|A stem-group [[teleost]]
|A stem-group [[teleost]]
|[[File:Pholidophorus bechei Lyme Regis Teylers.JPG|center|frameless|320x320px]]
|
|-
|-
|''"[[Coccolepis]]"''
|''"[[Coccolepis]]"''
Line 612: Line 675:
|
|
|A [[Chondrosteidae|chondrosteid]] [[Acipenseriformes|acipenseriform]] fish, related to [[sturgeon]] and [[paddlefish]]
|A [[Chondrosteidae|chondrosteid]] [[Acipenseriformes|acipenseriform]] fish, related to [[sturgeon]] and [[paddlefish]]
|[[File:FMIB 51725 Chondrosteus acipeenseroides Egerton Family Chondrosteidae.jpeg|center|frameless|340x340px]]
|
|-
|-
|''[[Oxygnathus]]''
|''Oxygnathus''
|''O. ornatus''
|''O. ornatus''
|
|
Line 647: Line 710:
|-
|-
|''[[Caturus]]''
|''[[Caturus]]''
|Spp.
|
|
|
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Line 672: Line 735:


=== Insects ===
=== Insects ===
Numerous species of insect are known from concretions, predominately in the Black Ven Marl Member.<ref>{{Cite web|title=Flatstones, Charmouth (Jackson collection) (Jurassic of the United Kingdom)|url=http://www.fossilworks.org/cgi-bin/bridge.pl?a=collectionSearch&collection_no=114637|access-date=17 December 2021|website=fossilworks.org}}</ref><ref>{{Cite web|title=Flatstones, Stonebarrow, Charmouth (Jackson collection) (Jurassic of the United Kingdom)|url=http://www.fossilworks.org/cgi-bin/bridge.pl?a=collectionSearch&collection_no=114638|access-date=17 December 2021|website=fossilworks.org}}</ref><ref>{{Cite web|title=Flatstones, Black Ven, Charmouth (Jackson collection)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114639|access-date=2020-08-26|website=paleobiodb.org}}</ref><ref>{{Cite web|title=Woodstones, Black Ven, Charmouth (Jackson collection)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114640|access-date=2020-08-26|website=paleobiodb.org}}</ref>
Numerous species of insect are known from concretions, predominately in the Black Ven Marl Member.<ref>{{Cite web|title=Flatstones, Charmouth (Jackson collection) (Jurassic of the United Kingdom)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114637|access-date=17 December 2021|website=fossilworks.org}}</ref><ref>{{Cite web|title=Flatstones, Stonebarrow, Charmouth (Jackson collection) (Jurassic of the United Kingdom)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114638|access-date=17 December 2021|website=fossilworks.org}}</ref><ref>{{Cite web|title=Flatstones, Black Ven, Charmouth (Jackson collection)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114639|access-date=2020-08-26|website=[[Paleobiology Database]]}}</ref><ref>{{Cite web|title=Woodstones, Black Ven, Charmouth (Jackson collection)|url=https://paleobiodb.org/classic/displayCollResults?collection_no=114640|access-date=2020-08-26|website=[[Paleobiology Database]]}}</ref>
{| class="wikitable" width="100%" align="center"
{| class="wikitable" width="100%" align="center"
! colspan="7" align="center" |'''Insects of the Charmouth Mudstone Formation'''
! colspan="7" align="center" |'''Insects of the Charmouth Mudstone Formation'''
Line 684: Line 747:
!Images
!Images
|-
|-
|''[[Anglophlebia]]''<ref name=":4">{{Cite journal |last=Zeuner |first=F. E. |date=1962 |title=Fossil insects from the Lower Lias of Charmouth, Dorset |url=https://www.biodiversitylibrary.org/part/83511 |journal=Bulletin of the British Museum (Natural History), Geology |volume=7 |issue=1 |pages=155–171}}</ref>
|''[[Anglophlebia]]''
|''A. gigantea''
|''A. gigantea''
|Woodstones, Black Ven
|
|Black Ven Marl Member
|
|NHMUK In.51030, part and counterpart of partial forewing
|NHMUK In.51030, part and counterpart of partial forewing
|Damsel-dragonfly
|Damsel-dragonfly
|[[File:Anglophlebia gigantea.jpg|frameless]]
|[[File:Anglophlebia gigantea.jpg|frameless]]
|-
|-
|''[[Archaeolepis]]<ref name=":5">{{Cite journal |last=Whalley |first=P. E. S. |date=1985 |title=The systematics and palaeogeography of the Lower Jurassic insects of Dorset, England |url=https://www.biodiversitylibrary.org/part/83348 |journal=Bulletin of the British Museum (Natural History), Geology |volume=39 |issue=3 |pages=107–189}}</ref>''
|''[[Archaeolepis]]''
|''A. mane''
|''A. mane''
|Birchi Nodules, Black Ven
|Birchi Nodules, Black Ven
Line 700: Line 763:
|
|
|-
|-
|''[[Archelcana]]''
|''[[Archelcana]]<ref name=":5" />''
|''A. durnovaria''
|''A. durnovaria''
|Woodstones
|Woodstones
|Black Ven Marl Member
|Black Ven Marl Member
|BMNH In 59162
|
|[[Elcanidae|Elcanid]] [[orthoptera]]n
|[[Elcanidae|Elcanid]] [[orthoptera]]n
|
|
|-
|-
|''[[Austaulius]]''<ref>{{Cite journal |last1=Kelly |first1=Richard S. |last2=Ross |first2=Andrew J. |last3=Coram |first3=Robert A. |date=2018 |title=A Review of Necrotauliids from the Triassic/Jurassic of England (Trichoptera: Necrotauliidae) |journal=Psyche: A Journal of Entomology |volume=2018 |pages=1–12 |doi=10.1155/2018/6706120 |doi-access=free |issn=0033-2615|hdl=1983/06ae01fb-06bc-41d0-bcbd-f7519cb07df6 |hdl-access=free }}</ref>
|''[[Austaulius]]''
|''A. haustrum''
|''A. haustrum''
|Monmouth Beach, Lyme Regis
|Monmouth Beach, Lyme Regis
|Black Ven Marl Member
|
|NHMUK II 3103
|NHMUK II 3103
|[[Necrotauliidae|Necrotauliid]] [[caddisfly]]
|[[Necrotauliidae|Necrotauliid]] [[caddisfly]]
|
|
|-
|-
|''[[Brevicula]]''<ref name=":5" /><ref name=":6">{{Cite journal |last1=Thomson |first1=Ulysses |last2=Ross |first2=Andrew J. |last3=Davidson |first3=Philip |date=2017 |title=Mesozoic Holcoptera (Coleoptera: Coptoclavidae) from England and the United States |url=http://dx.doi.org/10.1016/j.pgeola.2017.05.009 |journal=Proceedings of the Geologists' Association |volume=128 |issue=4 |pages=659–674 |doi=10.1016/j.pgeola.2017.05.009 |bibcode=2017PrGA..128..659T |issn=0016-7878|hdl=1983/f54090ef-e495-4f35-a375-b845583021e3 |hdl-access=free }}</ref><ref name=":7">{{Cite journal |last1=Kelly |first1=Richard S. |last2=Ross |first2=Andrew J. |last3=Jarzembowski |first3=Edmund A. |date=2016 |title=Earwigs (Dermaptera) from the Mesozoic of England and Australia, described from isolated tegmina, including the first species to be named from the Triassic |url=http://dx.doi.org/10.1017/s1755691017000329 |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |volume=107 |issue=2–3 |pages=129–143 |doi=10.1017/s1755691017000329 |bibcode=2016EESTR.107..129K |issn=1755-6910|hdl=1983/d363e702-e4cb-4285-8ee8-393e808c890e |hdl-access=free }}</ref>
|''[[Brevicula]]''
|''B. gradus, B. maculata''
|''B. gradus, B. maculata''
|Flatstones, Woodstones
|Flatstones, Woodstones;Monmouth Beach, Lyme Regis
|Black Ven Marl Member
|
|B. gradus: Holotype: NHMUK In.53993, female with ovipositor, B. maculata: Holotype: NHMUK II.3086, isolated Tegmina
|B. gradus: Holotype: NHMUK In.53993, female with ovipositor, B. maculata: Holotype: NHMUK II.3086, isolated Tegmina
|[[Dermapteridae|Dermapterid]] earwig
|[[Dermapteridae|Dermapterid]] earwig
|[[File:Brevicula.jpg|center|frameless]]
|[[File:Brevicula.jpg|center|frameless]]
|-
|-
|''[[Brochocoleus]]''
|''[[Brochocoleus]]<ref name=":5" />''
|''B. maculatus''
|''B. maculatus''
|Flatstones, Stonebarrow
|
|Black Ven Marl Member
|
|BMNH In 49577, 59133, 59361, 59154
|
|[[Ommatidae|Ommatid]] beetle
|[[Ommatidae|Ommatid]] beetle
|
|
|-
|-
|''[[Chrismooreia]]''<ref name=":8">{{Cite journal |last=Bechly |first=Guenter |date=2018-04-27 |title=Chrismooreia michaelbehei gen. et sp. nov. (Insecta: Odonata: Asiopteridae), a new fossil damsel-dragonfly from the Early Jurassic of England |url=http://dx.doi.org/10.5048/bio-c.2018.1 |journal=BIO-Complexity |volume=2018 |issue=1 |doi=10.5048/bio-c.2018.1 |issn=2151-7444}}</ref>
|''[[Chrismooreia]]''
|''C. michaelbehei''
|''C. michaelbehei''
|Charmouth
|Charmouth
Line 740: Line 803:
|[[File:108-750-2-SP.tif|center|frameless]]
|[[File:108-750-2-SP.tif|center|frameless]]
|-
|-
|''[[Dacryoderma]]''
|''[[Dacryoderma]]''<ref name=":7" />
|''B. teres''
|''B. teres''
|Charmouth-by-pass construction site
|Charmouth-by-pass construction site
Line 748: Line 811:
|
|
|-
|-
|''[[Daniilacheta]]''<ref name=":9">{{Cite journal |last1=GOROCHOV |first1=ANDREI V. |last2=CORAM |first2=ROBERT A. |date=2023-04-28 |title=New and little known taxa of the order Orthoptera (Insecta) from the Upper Triassic and Lower Jurassic of England |url=http://dx.doi.org/10.11646/palaeoentomology.6.2.11 |journal=Palaeoentomology |volume=6 |issue=2 |doi=10.11646/palaeoentomology.6.2.11 |issn=2624-2834}}</ref>
|''[[Dorsettia]]''
|''D. laeta''
|''D. aristovi''
|Monmouth Beach, Brooki bed
|Flatstones
|Shales with Beef Member
|ZIN LD/Orth 19
|[[Protogryllidae]] cricket
|
|
|-
|''[[Dorsettia]]<ref name=":5" />''
|''D. laeta''
|Flatstones
|Black Ven Marl Member
|BMNH In 59375, a male hindwing fragment
|BMNH In 59375, a male hindwing fragment
|[[Campterophlebiidae|Campterophlebiid]] damsel-dragonfly, genus also known from China
|[[Campterophlebiidae|Campterophlebiid]] damsel-dragonfly, genus also known from China
|
|
|-
|-
|''[[Durnovaria (insect)|Durnovaria]]''
|''[[Durnovaria (insect)|Durnovaria]]<ref name=":5" />''
|''D. parallela''
|''D. parallela''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 59171
|
|[[Aerophasmatidae|Aerophasmatid]], Stem group of [[Phasmatodea]]
|[[Aerophasmatidae|Aerophasmatid]], Stem group of [[Phasmatodea]]
|
|
|-
|-
|''[[Elaterina]]''
|''[[Elaterina]]''<ref name=":5" />
|''E. liassica''
|''E. liassica''
|Lyme Regis
|Lyme Regis
|Black Ven Marl Member
|
|BMNH In 59369
|
|[[Click beetle]]
|[[Click beetle]]
|
|
|-
|-
|''[[Elaterophanes]]''
|''[[Elaterophanes]]''<ref name=":6" />
|''E. regius''
|''E. regius''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 59385, 53952
|
|Click beetle
|Click beetle
|
|
|-
|-
|''[[Eoptychoptera]]''
|''[[Eoptychoptera]]<ref name=":5" />''
|''E. spectra''
|''E. spectra''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 49226
|
|[[Ptychopteridae|Ptychopterid]] fly, formerly referred to the genus ''Prodocidia''
|[[Ptychopteridae|Ptychopterid]] fly, formerly referred to the genus ''Prodocidia''
|
|
|-
|-
|''[[Holcoptera]]''
|''[[Hagla]]''<ref name=":5" />
|''H. giebeli, H. alisonae''
|''H. cf. gracilis''
|Flatstones
|Flatstones
|Bed 83 of the ''Obtusum'' Subchronozone, Black Ven Marl Member
|Black Ven Marl Member
|BMNH In 59167
|[[Haglidae]] hump-winged cricket
|
|-
|''[[Holcoptera]]''<ref name=":4" /><ref name=":5" /><ref name=":6" />
|''H. giebeli, H. alisonae; H. schlotheimi''
|Flatstones; Bed 75, Black Ven; Catherstone Lane; Monmouth Beach, Lyme Regis
|Black Ven Marl Member
|Complete specimen, isolated wings
|Complete specimen, isolated wings
|[[Coptoclavidae|Coptoclavid]] beetle
|[[Coptoclavidae|Coptoclavid]] beetle
|[[File:Holcoptera giebeli NHMUK PI II.3101 a specimen.jpg|center|frameless]]
|[[File:Holcoptera giebeli NHMUK PI II.3101 a specimen.jpg|center|frameless]]
|-
|-
|''[[Hypsothemis]]''
|''[[Hypsothemis]]<ref name=":5" />''
|''H. fraseri''
|''H. fraseri''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 59109, a hindwing
|BMNH In 59109, a hindwing
|Campterophlebiid damsel-dragonfly, genus also known from China and Kazakhstan
|Campterophlebiid damsel-dragonfly, genus also known from China and Kazakhstan
|
|
|-
|-
|''[[Jurachorista]]''<ref>{{Cite journal |last1=Soszyńska-Maj |first1=Agnieszka |last2=Krzemiński |first2=Wiesław |last3=Kopeć |first3=Katarzyna |last4=Coram |first4=Robert A. |date=2016-02-05 |title=Phylogenetic relationships within the relict family Eomeropidae (Insecta, Mecoptera) based on the oldest fossil from the Early Jurassic (Sinemurian) of Dorset, southern England |url=http://dx.doi.org/10.1080/14772019.2016.1139007 |journal=Journal of Systematic Palaeontology |volume=14 |issue=12 |pages=1025–1031 |doi=10.1080/14772019.2016.1139007 |bibcode=2016JSPal..14.1025S |issn=1477-2019}}</ref>
|''[[Jurachorista]]''
|''J. bashkuevi''
|''J. bashkuevi''
|Monmouth Beach, Brooki bed
|Monmouth Beach, Brooki bed
|Shales with Beef Member
|Shales with Beef Member
|ISEA I-F/MP/4/1637/14
|
|[[Eomeropidae|Eomeropid]] [[scorpionfly]]
|[[Eomeropidae|Eomeropid]] [[scorpionfly]]
|
|
|-
|-
|''[[Lateophlebia]]''
|''[[Lateophlebia]]<ref name=":5" />''
|''L. anglicanopsis''
|''L. anglicanopsis''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 49573, 59376
|
|Campterophlebiid damsel-dragonfly
|Campterophlebiid damsel-dragonfly
|
|
|-
|-
|''[[Lethonectes]]''<ref name=":10">{{Cite journal |last1=Popov |first1=Yuri A. |last2=Dolling |first2=W. R. |last3=Whalley |first3=P. E. S. |date=1994 |title=British Upper Triassic and Lower Jurassic Heteroptera and Coleorrhyncha (Insecta: Hemiptera) |url=https://archive.org/details/genus-0867-1710-5-4-307-347 |journal=Genus |volume=5 |issue=4 |pages=307–347 |doi= |issn=}}</ref>
|''[[Liassocorixa]]''
|''L. dorsetica''
|''L. naucoroides''
|Flatstones
|Flatstones
|Black Ven Marl Member
|BMNH In 49615, 51014(59383)
|[[Belostomatidae]] giant water bug
|
|
|
|-
|''[[Liassocorixa]]''<ref name=":10" />
|''L. dorsetica''
|Flatstones
|Black Ven Marl Member
|BMNH In 59166
|[[Corixidae|Corixid]] bug
|[[Corixidae|Corixid]] bug
|
|
|-
|-
|''[[Liassocupes]]''
|''[[Liassocupes]]''<ref name=":4" />
|''L. parvus''
|''L. parvus''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 64008, 49210
|
|Ommatid beetle
|Ommatid beetle
|
|
|-
|-
|''[[Liassophlebia]]''
|''[[Liassophlebia]]<ref name=":5" />''
|''L. pseudomagnifica''
|''L. pseudomagnifica''
|Stonebarrow
|Stonebarrow
|Black Ven Marl Member
|
|NHMUK In.64000, partial hindwing
|NHMUK In.64000, partial hindwing
|[[Liassophlebiidae|Liassophlebiid]] damsel-dragonfly
|[[Liassophlebiidae|Liassophlebiid]] damsel-dragonfly
|[[File:Liassophlebia pseudomagnifica.jpg|center|frameless]]
|[[File:Liassophlebia pseudomagnifica.jpg|center|frameless]]
|-
|-
|''[[Locustopsis]]''
|''[[Locustopsis]]''<ref name=":5" /><ref name=":9" />
|''L. spectabilis''
|''L. spectabilis''
|Flatstones
|Flatstones; Monmouth Beach, Brooki bed
|Black Ven Marl Member
|
|ZIN LD/Orth 1, Orth 4, Orth 10, Orth 18, Orth 31
|
|[[Locustopsidae|Locustopsid]] grasshopper
|[[Locustopsidae|Locustopsid]] grasshopper
|
|
|-
|-
|''[[Megarhyphus]]''<ref>{{Cite journal |last1=EWA |first1=Krzemińska |last2=ROBERT |first2=Coram A. |last3=WIESŁAW |first3=Krzeminski |date=2010 |title=A New Species of ''Megarhyphus'', an Interesting Discovery from the Lower Jurassic of England (Diptera, Anisopodidae) |url=http://dx.doi.org/10.1111/j.1755-6724.2010.00231.x |journal=Acta Geologica Sinica - English Edition |volume=84 |issue=4 |pages=693–695 |doi=10.1111/j.1755-6724.2010.00231.x |bibcode=2010AcGlS..84..693E |issn=1000-9515}}</ref>
|''"Mesocixiella"''
|''M. amberae''
|Monmouth Beach, Brooki bed
|Shales with Beef Member
|NHML In 28447
|[[Anisopodidae]] wood gnat
|
|-
|''"Mesocixiella"''<ref name=":5" />
|''"M." fennahi''
|''"M." fennahi''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 53942
|
|[[Fulgoridiidae|Fulgoridiid]] [[planthopper]], does not belong to the genus ''[[Mesocixiella]]''
|[[Fulgoridiidae|Fulgoridiid]] [[planthopper]], does not belong to the genus ''[[Mesocixiella]]''
|
|
|-
|-
|''[[Metaraphidia]]''
|''[[Metaraphidia]]''<ref name=":5" />
|''M. confusa''
|''M. confusa''
|Charmouth (GSM)
|
|Black Ven Marl Member
|Black Ven Marl Member
|GSM 117552
|
|[[Snakefly]]
|[[Snakefly]]
|
|
|-
|-
|''[[Micromacula]]''
|''[[Micromacula]]<ref name=":5" />''
|''M. gracilis''
|''M. gracilis''
|Flatstones, Birchi Nodules
|Flatstones, Birchi Nodules; Flatstones, Black Ven
|Shales with Beef; Black Ven Marl Member
|
|BMNH In 49230
|
|[[Regiatidae|Regiatid]] orthopteran
|[[Regiatidae|Regiatid]] orthopteran
|
|
|-
|-
|''[[Mimemala]]''
|''[[Mimemala]]<ref name=":5" />''
|''M. giganteum''
|''M. giganteum''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 51026
|
|[[Schizocoleidae|Schizocoleid]] beetle
|[[Schizocoleidae|Schizocoleid]] beetle
|
|
|-
|-
|''[[Nannoblattina]]''
|''[[Nannoblattina]]''<ref name=":5" />
|''N. petulantia''
|''N. petulantia''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 53929, 51004
|
|[[Mesoblattinidae|Mesoblattinid]] cockroach
|[[Mesoblattinidae|Mesoblattinid]] cockroach
|
|
|-
|-
|''[[Nannotanyderus]]''<ref>{{Cite journal |last1=Skibińska |first1=Kornelia |last2=Krzemiński |first2=Wiesław |last3=Coram |first3=Robert A. |date=2014-08-28 |title=Discovery of the most ancient member of family Tanyderidae (Diptera) from the Lower Jurassic (Sinemurian) of England |url=http://dx.doi.org/10.11646/zootaxa.3857.1.6 |journal=Zootaxa |volume=3857 |issue=1 |pages=125–130 |doi=10.11646/zootaxa.3857.1.6 |pmid=25283100 |issn=1175-5334}}</ref>
|''[[Nannotanyderus]]''
|''N. oliviae''
|''N. oliviae''
|Monmouth Beach, Brooki bed
|Monmouth Beach, Brooki bed
|Shales with Beef Member
|Shales with Beef Member
|ISEA I-F/MP/2/1621/13, 12
|
|[[Tanyderidae|Tanyderid]] fly
|[[Tanyderidae|Tanyderid]] fly
|
|
|-
|-
|''[[Neomeridium]]''
|''[[Neomeridium]]''<ref name=":10" />
|''N. trifurcum''
|''N. trifurcum''
|Woodstones
|Woodstones
|Black Ven Marl Member
|
|BMNH In 59151, 51039, 53901, 64014
|
|[[Pachymeridiidae|Pachymeridiid]] [[Lygaeoidea|Lygaeoid]] [[Hemiptera|bug]]
|[[Pachymeridiidae|Pachymeridiid]] [[Lygaeoidea|Lygaeoid]] [[Hemiptera|bug]]
|
|
|-
|-
|''[[Oligophryne]]''<ref>{{Cite journal |last1=Ansorge |first1=J. |last2=Krzemiński |first2=W. |date=1994 |title=Oligophrynidae, a Lower Jurassic dipteran family (Diptera, Brachycera). |url=https://www.researchgate.net/publication/268518046 |journal=Acta Zoologica Cracoviensia |volume=37 |issue=2 |pages=115–119}}</ref>
|''[[Omma]]''
|''O. britannica''
|Stonebarrow Hill
|Black Ven Marl Member
|Coll. Ansorge fly No 1
|[[Oligophrynidae]] fly
|
|-
|''[[Omma]]''<ref name=":5" />
|''O. liassicum''
|''O. liassicum''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH 59132
|BMNH 59132
|Ommatid beetle, genus extant
|Ommatid beetle, genus extant
|
|
|-
|-
|''[[Orthophlebia]]''
|''[[Orthophlebia]]''<ref name=":5" />
|''O. capillata''
|''O. capillata''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 53924, an exoskeleton (wings & thorax)
|BMNH In 53924, an exoskeleton (wings & thorax)
|[[Orthophlebiidae|Orthophlebiid]] scorpionfly
|[[Orthophlebiidae|Orthophlebiid]] scorpionfly
|
|
|-
|-
|''[[Paraprosbole]]''
|''[[Orichalcum]]<ref name=":5" />''
|''P. rotruda''
|''O. ornatum''
|Flatstones
|Flatstones, Birchi Nodules
|Shales with Beef
|
|BMNH In 53983
|[[Triassomanteidae]] grasshopper
|
|
|-
|''[[Paraprosbole]]''<ref name=":5" />
|''P. rotruda''
|Flatstones
|Black Ven Marl Member
|BMNH In 59374, 48162, 64395
|[[Tettigarctidae|Tettigarctid]] cicada
|[[Tettigarctidae|Tettigarctid]] cicada
|
|
|-
|-
|''[[Propreocoris]]''
|''[[Propreocoris]]''<ref name=":10" />
|''P. maculatus''
|''P. maculatus''
|Woodstones
|Woodstones
|Black Ven Marl Member
|
|BMNH In 59152
|
|[[Stem-group]] to [[Ochteridae]] and [[Gelastocoridae]]
|[[Stem-group]] to [[Ochteridae]] and [[Gelastocoridae]]
|
|
|-
|-
|''[[Protohagla]]''
|''[[Protogryllus]]''<ref name=":5" />
|''P. langi''
|''P. (Archaegryllodes) magnus''
|Flatstones
|Flatstones
|Black Ven Marl Member
|BMNH In 44003, 51016, 51022, 59373
|[[Protogryllidae]] cricket
|
|
|
|-
|''[[Protohagla]]''<ref name=":4" /><ref name=":5" />
|''P. langi''
|Flatstones
|Black Ven Marl Member
|BMNH In 59018
|[[Haglidae|Haglid]] cricket
|[[Haglidae|Haglid]] cricket
|
|
|-
|-
|''[[Protorthophlebia]]''
|''[[Protomyrmeleon]]''<ref name=":8" />
|''P. cf. brunonis''
|Charmouth burden pile
|Black Ven Marl Member
|AR 428
|[[Protomyrmeleontidae]] winged insect
|
|-
|''[[Protorthophlebia]]''<ref name=":5" />
|''P. latipennis''
|''P. latipennis''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 49575, 53954, 53963, 59120, 59365, 59371, 59384
|
|[[Protorthophlebia|Protorthophlebiid]] scorpionfly
|[[Protorthophlebia|Protorthophlebiid]] scorpionfly
|
|
|-
|-
|''[[Priscaenigma]]''
|''[[Priscaenigma]]''<ref name=":5" />
|''P. obtusa''
|''P. obtusa''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 53898
|
|Snakefly
|Snakefly
|
|
|-
|-
|''[[Pseudopolycentropus]]''
|''[[Pseudopolycentropus]]''<ref name=":4" />
|''P. triangularis''
|''P. triangularis''
|Woodstones
|Woodstones
|Black Ven Marl Member
|Black Ven Marl Member
|BMNH In 53915
|
|[[Pseudopolycentropodidae|Pseudopolycentropodid]] [[Mecoptera|scorpionfly]]
|[[Pseudopolycentropodidae|Pseudopolycentropodid]] [[Mecoptera|scorpionfly]]
|
|
|-
|-
|''[[Pterocimex]]''
|''[[Pterocimex]]''<ref name=":10" />
|''P. jacksoni''
|''P. jacksoni''
|Woodstones
|Woodstones
|Black Ven Marl Member
|Black Ven Marl Member
|BMNH in 53909, 49589, 51048, 59359
|
|[[Nepomorpha]]n
|[[Nepomorpha]]n
|
|
|-
|-
|''[[Regiata]]''
|''[[Regiata]]''<ref name=":5" />
|''R. scutra''
|''R. scutra''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|BMNH In 64027
|
|Regiatid orthopteran
|Regiatid orthopteran
|
|
|-
|-
|''[[Rossiphlebia]]''
|''[[Rhipidoblattina]]''<ref name=":5" />
|''R. jacksoni''
|''R. ssp.''
|Flatstones
|Flatstones
|Black Ven Marl Member
|BMNH In 49212, 49249, 53909, 53920
|[[Caloblattinidae]] cockroach
|
|
|-
|''[[Rossiphlebia]]''<ref name=":5" />
|''R. jacksoni''
|Flatstones
|Black Ven Marl Member
|NHMUK In.53999, part and counterpart of a partial hindwing
|NHMUK In.53999, part and counterpart of a partial hindwing
|Liassophlebiid damsel-dragonfly
|Liassophlebiid damsel-dragonfly
|[[File:Rossiphlebia jacksoni.jpg|center|frameless]]
|[[File:Rossiphlebia jacksoni.jpg|center|frameless]]
|-
|-
|''[[Tersus]]''
|''[[Tarsabedus]]''<ref name=":10" />
|''T. menkei''
|0.6 km east Charmouth
|Black Ven Marl Member
|BRSMG Cc 688
|[[Belostomatidae]] giant water bug
|
|-
|''[[Tersus]]''<ref name=":5" />
|''T. crowsoni''
|''T. crowsoni''
|Flatstones
|Flatstones
|Black Ven Marl Member
|
|NHM, In. 53949
|BMNH In 53949, 59038
|[[Schizophoridae|Schizophorid]] beetle
|[[Schizophoridae|Schizophorid]] beetle
|
|
|}

=== Plants ===
{| class="wikitable"
!Genus
!Species
!Stratigraphic position
!Material
!Notes
!Images
|-
|style="background:#D1FFCF;" |''[[Ctenozamites]]''<ref name=":2">{{Cite journal |last=van Konijnenburg-van Cittert |first=Johanna H.A. |date=2008 |title=The Jurassic fossil plant record of the UK area |url=http://dx.doi.org/10.1016/s0016-7878(08)80258-1 |journal=Proceedings of the Geologists' Association |volume=119 |issue=1 |pages=59–72 |doi=10.1016/s0016-7878(08)80258-1 |bibcode=2008PrGA..119...59V |issn=0016-7878}}</ref>
|style="background:#D1FFCF;" |
* ''C. cycadea''
|style="background:#D1FFCF;" |
* Lyme regis
* Charmouth
|style="background:#D1FFCF;" |Isolated shoots
|style="background:#D1FFCF;" |A "Seed Fern" of the group with [[Corystospermales]]
|style="background:#D1FFCF;" |
|-
|style="background:#D1FFCF;" |''[[Cycadites]]''<ref name=":2" />
|style="background:#D1FFCF;" |
* ''C. rectangularis''
|style="background:#D1FFCF;" |
* Lyme regis
|style="background:#D1FFCF;" |Isolated shoots
|style="background:#D1FFCF;" |A possible Cycadophyte leaf, related with [[Cycadales]]. Alternatively, it can be of Bennettite origin
|style="background:#D1FFCF;" |
|-
|style="background:#D1FFCF;" |''[[Cycadeoidea]]''<ref name=":2" />
|style="background:#D1FFCF;" |
* ''C. gracilis''
|style="background:#D1FFCF;" |
* Lyme regis
|style="background:#D1FFCF;" |Stems
|style="background:#D1FFCF;" |Bennettite Trunks of the family [[Cycadeoidaceae]]
|style="background:#D1FFCF;" |[[File:Cycadeoidea fossil cropped.png|thumb|Example of specimen of the genus]]
|-
|style="background:#D1FFCF;" |''[[Cycadopteris]]''<ref name=":2" />
|style="background:#D1FFCF;" |
* ''C. jurensis''
|style="background:#D1FFCF;" |
* Lyme regis
|style="background:#D1FFCF;" |Isolated shoots
|style="background:#D1FFCF;" |A "Seed Fern" of the group with [[Corystospermales]]
|style="background:#D1FFCF;" |[[File:Cycadopteris jurensis 44.JPG|thumb|169x169px|Shoot of the same species]]
|-
|style="background:#D1FFCF;" |''[[Otozamites]]''<ref name=":3" />
|style="background:#D1FFCF;" |
* ''O. obtusus''
|style="background:#D1FFCF;" |
* Lyme regis
* Charmouth
* Seatown
|style="background:#D1FFCF;" |Isolated shoots
|style="background:#D1FFCF;" |Bennettite leafs of the family [[Williamsoniaceae]]
|style="background:#D1FFCF;" |[[File:Otozamites Anina jurassic (15586380432).jpg|thumb|240x240px|Shoot of the same species]]
|-
| style="background:#D1FFCF;" |''[[Pagiophyllum]]''<ref name=":3">{{Cite journal |last=Chamberlain |first=C. J. |date=1919 |title=Fossil Plants ''Fossil Plants, a Textbook for Students of Botany and Geology.'' A. C. Seward |url=http://dx.doi.org/10.1086/332400 |journal=Botanical Gazette |volume=67 |issue=1 |pages=93–95 |doi=10.1086/332400 |issn=0006-8071}}</ref>
| style="background:#D1FFCF;" |
* ''P. peregrinum''
* ''P. sewardii''
| style="background:#D1FFCF;" |
* Lyme regis
* Charmouth
* Seatown
* Wear Cliffs
| style="background:#D1FFCF;" |Isolated shoots
| style="background:#D1FFCF;" |[[Conifer]] shoots of the family [[Hirmeriellaceae]] or [[Araucariaceae]]
| style="background:#D1FFCF;" |[[File:Pagiophyllum sp. - Naturhistorisches Museum, Braunschweig, Germany - DSC05070 cropped.jpg|thumb|186x186px|Shoot of the same species]]
|}
|}


== See also ==
== See also ==
* [[List of fossiliferous stratigraphic units in England]]
* [[List of fossiliferous stratigraphic units in England]]
* [[List of dinosaur-bearing rock formations]]
{{div col|colwidth=29em}}
* [[Blue Lias]], England
* [[Sorthat Formation]], Denmark
* [[Hasle Formation]], Denmark
* [[Zagaje Formation]], Poland
* [[Drzewica Formation]], Poland
* [[Ciechocinek Formation]], Poland
* [[Borucice Formation]], Poland
* [[Rotzo Formation]], Italy
* [[Saltrio Formation]], Italy
* [[Moltrasio Formation]], Italy
* [[Marne di Monte Serrone]], Italy
* [[Calcare di Sogno]], Italy
* [[Podpeč Limestone]], Slovenia
* [[Coimbra Formation]], Portugal
* [[El Pedregal Formation]], Spain
* [[Fernie Formation]], Canada
* [[Whiteaves Formation]], British Columbia
* [[Navajo Sandstone]], Utah
* [[Aganane Formation]], Morocco
* [[Tafraout Group]], Morocco
* [[Azilal Formation]], Morocco
* [[Budoš Limestone]], Montenegro
* [[Kota Formation]], India
* [[Cañadón Asfalto Formation]], Argentina
* [[Los Molles Formation]], Argentina
* [[Kandreho Formation]], Madagascar
* [[Elliot Formation]], South Africa
* [[Clarens Formation]], South Africa
* [[Evergreen Formation]], Australia
* [[Cattamarra Coal Measures]], Australia
* [[Hanson Formation]], Antarctica
* [[Mawson Formation]], Antarctica
{{div col end}}


== References ==
== References ==

Revision as of 17:19, 28 November 2024

Charmouth Mudstone Formation
Stratigraphic range: Lower Sinemurian– Lower Pliensbachian Semicostatum-lowermost Margaritatus 198–188 Ma
Cliffs of the Charmouth Mudstone Formation at Charmouth, Dorset
TypeFormation
Unit ofLias Group
Sub-unitsShales With Beef Member, Black Ven Marl Member, Belemnite Marl Member, Green Ammonite Member
UnderliesDyrham Formation, Marlstone Rock Formation, unconformity with Gault (Charmouth area)
OverliesBlue Lias Formation, Scunthorpe Mudstone
AreaWessex Basin, Worcester Basin, East Midland Shelf
Thickness~335 m
Lithology
PrimaryShale, mudstone
OtherLimestone, sandstone
Location
RegionEngland
Country United Kingdom
Type section
Named forCharmouth
LocationCliff and foreshore exposures between Seven Rock Point and Golden Cap, Dorset

The Charmouth Mudstone Formation is a geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian).[1] It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cliff section between Lyme Regis and Charmouth (alongside the underlying Blue Lias) but onshore it extends northwards to Market Weighton, Yorkshire, and in the subsurface of the East Midlands Shelf and Wessex Basin. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including Mary Anning.

Stratigraphy

Shales With Beef Member

The Shales With Beef Member is around 28–30 metres thick in the Lyme Regis-Charmouth region and predominantly consists of thinly bedded medium to dark grey mudstone, blocky calcareous pale-weathering mudstone and brown-grey organic-rich mudstones with frequent bedding parallel veins of fibrous calcite ("beef"), that are usually less than 10 centimetres thick. Several beds of nodular and tabular limestone are also present. It is the lowest unit of the formation and directly overlies the Blue Lias Formation, with the boundary being marked by a prominent bioturbated horizon. Notable persistent marker beds within the member include the laminated calcareous siltstone "Fish Bed", "Table Ledge", which consists of lens beds of limestones with mud content with nests of rhynchonellid brachiopods, the Devonshire Head and the Spittles limestones and the Birchi Nodules (which are septarian concretions) The upper boundary with the Black Ven Marl Member is marked by the prominent laterally persistent limestone Birchi Tabular Bed.[2]

Black Ven Marl Member

The Black Ven Marl Member is around 43 metres thick consists of thinly bedded dark mudstones,[3] with several laterally persistent cementstone horizons, notable horizons include the Lower and Upper Cement beds and the Stellare nodules.

Belemnite Marl Member

The Belemnite Marl Member is around 20 to 27 metres thick, and consists of interbedded pale and dark grey calcareous mudstone, with numerous belemnites, hence the name. The top of the member is marked by the Belemnite Stone Bed[4]

Green Ammonite Member

The Green Ammonite Member is up to 31 metres thick predominantly consists of medium grey mudstones, with 3 limestone horizons, Lower Limestone; Red Band, and Upper Limestone, it is conformably overlain by the Dyrham Formation in some areas,[5] but in the Charmouth area there is an erosive unconformable boundary with the much younger Early Cretaceous (Albian) aged Gault clays.

Paleobiota

Vertebrate fauna of the Charmouth Mudstone Formation

Ammonites

Ammonites of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images
Apoderoceras A. cf. dunrobinense, A. subtriangulare
Arnioceras A. semicostatum Semicostatum Zone
Asteroceras A. obtusum Obtusum zone
Caenisites
Cymbites
Echioceras E. raricostatum Raricostatum Zone
Eoderoceras E. bispinigerum
Gemmellaroceras G. cf. peregrinum Belemnite Marls
Oxynoticeras O. williamsi
Phricodoceras P. taylori, P. lamellosum Belemnite Marls
Promicroceras P. planicosta Obtusum Zone
Radstockiceras R. complicatum
Tragophylloceras T. ibex, T. loscombi Green Ammonite Member
Uptonia U. bronni
Xipheroceras

Sauropterygia

Sauropterygia of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images

Archaeonectrus[6]

A. rostratus

Charmouth

Arnioceras semicostatum-Echioceras raricostatum ammonoid zone

BMNH 38525 (holotype skeleton)

A marine sauropterygian, member of the family Rhomaleosauridae

Attenborosaurus[7]

A. conybeari

  • Black Ven Water, Charmouth
  • Northwest corner of Black Ven Water

Asteroceras obtusum, Arnioceras semicostatum-Echioceras raricostatum ammonoid zones

  • Holotype skeleton (now destroyed)
  • BMNH 40140, partial skeleton and skull

A marine sauropterygian, early member of the family Pliosauridae. It was first identified as Plesiosaurus conybeari.

Cast of the holotype

?Plesiosaurus[8]

? P. sp.

  • Eastern point of Wear Cliffs, below Golden Cap

Prodactylioceras davoei ammonoid zone

  • BRSMG Ce17972a-o, articulated juvenile partial postcranial skeleton

A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. A juvenile specimen that resembles those assigned to the genus Plesiosaurus. Its assignation is dubious.

Plesiosaurus[9]

P. dolichodeirus

  • Lyme Regis

Echioceras raricostatum ammonoid zone

  • Holotype (BMNH 22656)
  • Numerous specimens referred

A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. The typical plesiosaur, and one of the most common found on the formation.

Thaumatodracon[10] T. wiedenrothi Between Lyme Regis and Charmouth Black Ven NLMH 106.058, "a partial skeleton comprising a complete cranium, mandible, articulated cervical series, and indeterminate fragmentary remains" A rhomaleosaurid

Ichthyosauria

Ichthyosauria of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images
Ichthyosauria indet. Indeterminate Lyme Regis, Charmouth Indeterminate remains Un-attributable to any genus

Ichthyosaurus

I. anningae[11]

Charmouth

Stonebarrow Marls Member

DONMG:1983.98 Subadult specimen

An ichthyosaurid ichthyosaur

Ichthyosaurus sp.

West of Westhay Water, Stonebarrow

Stonebarrow Marls Member

NHMUK R15907, partial skeleton[12]

Considered to belong to I. communis by Bennett et al. 2012, considered indeterminate within Ichthyosaurus by Lomax and Massare 2015

Leptonectes

L. moorei[13]

Seatown

Belemnite Marls Member

BMNH R14370 partial anterior skeleton with most complete skull

A leptonectid ichthyosaur

L. solei[14]

Seatown

Acanthopleuroceras valdani Subzone, Tragophylloceras ibex Zone, lower Pliensbachian

NMW 91.296.2.2 "a large, isolated and incomplete forefin"

Temnodontosaurus T. platyodon[15]
Temnodontosaurus platyodon.

Thalattosuchia

Thalattosuchia of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images
Thalattosuchia[16] Indeterminate
  • Stonebarrow beach, Charmouth
  • Conesby Quarry, Scunthorpe
 Black Ven Mudstone Member
  • NHMUK PV R 36710, incomplete snout
  • NHMUK PV R 9731, axis and four post-axial cervical vertebrae. Two incomplete axial ribs, the lef fourth cervical rib, and two cervical rib fragments
  • NHMUK PV R 36 711, dorsal vertebra from the caudal end of the dorsal series, and the neural arch of another dorsal vertebra, both held together by matrix
 Early Thalattosaurian specimens that can't be referred to any concrete genus. Includes a possible early diverging metriorhynchoid

Turnersuchus[17]

T. hingleyae

1 kilometer west of the Charmouth Heritage Coast Centre

Belemnite Marl Member

LRM 2021/45, a partial skeleton including cranial material, cervical, dorsal, and caudal vertebrae, ribs, right pectoral girdle, partial limb bones, and an osteoderm

An early diverging thalattosuchian.

Pterosauria

Pterosauria of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images

Dimorphodon[18]

D. macronyx

  • Charmouth
  • Lyme Regis

Black Ven Marl Member

  • Tail Fragments
  • Tibia
  • Femur
  • Humerus
  • Phalanax
  • Metacarps
  • Semi-Complete Skeletons

A basal pterosaur and the type member of the family Dimorphodontidae.

Dimorphodontidae[19][20]

Indeterminate

  • Charmouth, Black Ven
  • Lyme Regis

Black Ven Marl Member

  • OUM J.53070
  • NHMUK PV R1595, wing phalanges

A basal dimorphodontid pterosaur. Known as "Charmouth dimorphodontid" it shares resemblance with the genus Dimorphodon.

Indeterminate[21]

  • Near Seatown

Lower Pliensbachian member

Associated remains of the rostrum and mandibles

A basal dimorphodontid pterosaur. The relatively deep rostrum and extreme size dimorphism in the dentition show clear similarities to Dimorphodon, also from the Lias of Dorset. The distinctive dentition, in which the first four pairs of rostral teeth and two pairs of mandibular teeth are relatively large and fang-like, while the remaining teeth are remarkably small and short.[21]

Dinosauria

Dinosauria of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images

Coelophysidae[22][23]

"Megalosaurus" lydekkeri

Lyme Regis

Black Ven Marl Member

  • NHMUK OR 41352, isolated tooth

A theropod, possibly a member of Coelophysidae inside Neotheropoda. Invalid and non diagnostic. It was identified as Zanclodon(?) sp. b., Megalosaurus lydekkeri and reassigned as Magnosaurus(?) lydekkeri.[24] Mortimer found that was changed to "Megalosaurus" woodwardi later.[24] It was found on mostly of recent works to be non diagnostic, probably a coelophysoid.[23]

Dornraptor[25]

D. normani

Lower cliff face, Charmouth

Black Ven Marl Member (Likely)

  • NHMUK OR 39496, Partial hind limb
  • GSM 109560, left femur

An Averostran theropod, possible Ceratosaur or Tetanuran. In 2024 was formally described, but referred to the older Blue Lias.[25] A paper non quoted on the description relocated it on the Charmouth mudstone.[26]

Neotheropoda[26]

Indeterminate

Near Lyme Regis

obtusum Zone, obtusum subzone

  • NHMUK PV R36855, Left fibula, ~ 17 cm long

A theropod, possibly a basal member of Neotheropoda. Previously misidentified as a referred specimen of the pterosaur Dimorphodon macronyx.[27] NHMUK PV R36855 differs from those of all other Early Jurassic-Late Triassic, theropods in its combination of features, but bears no discernible autapomorphies.[26] It is similar to Tawa, Dracoraptor and Liliensternus, being considered an early branching neotheropod, with a size range similar to Coelophysis bauri, representing an individual of approximately 10 kg, approximately the same size as Coelophysis bauri.[26]

Massopoda[28]

Indeterminate

lower cliff face, Charmouth

obtusum Zone, obtusum subzone

  • GSM 109561 pedal ungual

A sauropodomorph, possibly a member of Massopoda. While it was not associated with BMNH 39496 or GSM 109560 and was not assigned to "Merosaurus" it was classified as coming from an indeterminate theropod. Mortimer found closer resemblance with pedal unguals of basal massopods like Blikanasaurus and Jingshanosaurus.[29]

Scelidosaurus[30][31]

S. harrisonii

  • Charmouth, Black Ven
  • Black Ven landslip

Black Ven Marl Member

  • NHMUK OR42068. Six osteoderms
  • NHMUK OR42070. Phalanx.
  • NHMUK OR42072. Large bone fragment – indet.
  • NHMUK OR42074. Ulna (paired with NHMOR41323).
  • NHMUK OR43062. Shaft of a large humerus, crushed proximal femur, fibula, fragmentary radius?, other large bone fragments
  • NHMUK OR46011. Probable rib fragment – indet.
  • BRSMG Ce12785. Partial skull and some disarticulated postcranial elements
  • BRSMG Ce12787. Partly eroded femur and vertebra
  • BRSMG Ce12788. cervical osteoderms.
  • BRSMG Ce12789. Vertebra and bone fragments
  • BRSMG Cf2781. nine middle-distal caudal vertebrae and some haemal arches with some organic (kerogenized) remains of the skin preserved.
  • BRSMG LEGL 0004. Articulated skeleton
  • BRSMG LEGL 0005. Articulated partial skeleton
  • DORCM G.7542. Osteoderms
  • DORCM G.10817. Osteoderms
  • CAMSM X39256. Cranial: disarticulated skull and jaw elements

A basal thyreophoran, and the type member of the family Scelidosauridae. One of the best known armored dinosaurs of the lower Jurassic, also the most complete found to date, with specimens preserving even the skin and dermal armour.[31]

S. "sp. A"

Belemnite Marls at Seatown

  • DORCM G.7842. Three dorsal vertebrae in articulation

Cf. S. sp.

Foot of Black Ven, Charmouth

Asteroceras obtusum ammonoid zone, Black Ven Marl Member

  • NHMUK R6704, juvenile specimen
  • NHMUK R12019. A posterior fragment of the occiput and an articulated series of six cervical vertebrae
  • NHMUK R10103. Indeterminate bone fragments
  • NHMUK OR28333. Two scapulae
  • NHMUK OR32396. Radius
  • NHMUK OR39516. Osteoderms.
  • NHMUK OR39517. Caudal or?sacral vertebra, small distal caudal vertebra, small distal caudal
  • NHMUK OR39518. Haemal arch (chevron).
  • NHMUK OR39519. Two phalanges.
  • NHMUK OR39520. Ungual phalanx of ?pes.
  • NHMUK OR39521. Several bone fragments.
  • NHMUK OR40503. Tibia (proximal end).
  • NHMUK OR41322. Femur
  • NHMUK OR41323. Radius
  • NHMUK OR41324. Scapular blade
  • NHMUK OR41325. Humerus
  • NHMUK OR41326. Fibula
  • NHMUK OR41327. Two caudal vertebrae
  • NHMUK OR41328. Three metatarsals
  • NHMUK OR41329. Osteoderm
  • NHMUK OR41330. Ischium

Fish

Numerous fish species are known from the Charmouth Mudstone and underlying Blue Lias, from such horizons as the "Fish Bed" of the Shales With Beef Member.

Fish of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images
Acrodus A hybodont shark
Hybodus A hybodont shark
Palidiplospinax A synechodontiform shark
Squaloraja S. tenuispina, S. polyspondyla Closely related to modern chimaeras
Myriacanthus M. paradoxus, M. granulatus Black Ven Marl Member A myriacanthid closely related to modern chimaeras
Dorsetichthys D. bechei A stem-group teleost
"Coccolepis" "C." liassicus A coccolepidid fish, probably does not belong to the genus
Holophagus H. gulo A coelacanth
Chondrosteus C. acipenseroides A chondrosteid acipenseriform fish, related to sturgeon and paddlefish
Oxygnathus O. ornatus A palaeonisciform fish
Saurorhynchus S. brevirostris, S. anningae A member of Saurichthyiformes
Ptycholepis P. gracilis, P. curtus A palaeonisciform fish
Dapedium Spp. A dapediiform fish
Caturus Spp. An amiiform fish related to bowfins
Platysiagum P. sclerocephalum A platysiagid fish
Furo F. orthostomus A member of Ionoscopiformes within Halecomorphi

Insects

Numerous species of insect are known from concretions, predominately in the Black Ven Marl Member.[32][33][34][35]

Insects of the Charmouth Mudstone Formation
Genus Species Location Stratigraphic position Material Notes Images
Anglophlebia[36] A. gigantea Woodstones, Black Ven Black Ven Marl Member NHMUK In.51030, part and counterpart of partial forewing Damsel-dragonfly
Archaeolepis[37] A. mane Birchi Nodules, Black Ven Shales with Beef Isolated wing Among the oldest known lepidopterans
Archelcana[37] A. durnovaria Woodstones Black Ven Marl Member BMNH In 59162 Elcanid orthopteran
Austaulius[38] A. haustrum Monmouth Beach, Lyme Regis Black Ven Marl Member NHMUK II 3103 Necrotauliid caddisfly
Brevicula[37][39][40] B. gradus, B. maculata Flatstones, Woodstones;Monmouth Beach, Lyme Regis Black Ven Marl Member B. gradus: Holotype: NHMUK In.53993, female with ovipositor, B. maculata: Holotype: NHMUK II.3086, isolated Tegmina Dermapterid earwig
Brochocoleus[37] B. maculatus Flatstones, Stonebarrow Black Ven Marl Member BMNH In 49577, 59133, 59361, 59154 Ommatid beetle
Chrismooreia[41] C. michaelbehei Charmouth Obtusum Zone Part and counterpart of a mostly complete specimen Asiopterid damsel-dragonfly
Dacryoderma[40] B. teres Charmouth-by-pass construction site Obtusum Zone Isolated tegmen Dermapterid earwig
Daniilacheta[42] D. aristovi Monmouth Beach, Brooki bed Shales with Beef Member ZIN LD/Orth 19 Protogryllidae cricket
Dorsettia[37] D. laeta Flatstones Black Ven Marl Member BMNH In 59375, a male hindwing fragment Campterophlebiid damsel-dragonfly, genus also known from China
Durnovaria[37] D. parallela Flatstones Black Ven Marl Member BMNH In 59171 Aerophasmatid, Stem group of Phasmatodea
Elaterina[37] E. liassica Lyme Regis Black Ven Marl Member BMNH In 59369 Click beetle
Elaterophanes[39] E. regius Flatstones Black Ven Marl Member BMNH In 59385, 53952 Click beetle
Eoptychoptera[37] E. spectra Flatstones Black Ven Marl Member BMNH In 49226 Ptychopterid fly, formerly referred to the genus Prodocidia
Hagla[37] H. cf. gracilis Flatstones Black Ven Marl Member BMNH In 59167 Haglidae hump-winged cricket
Holcoptera[36][37][39] H. giebeli, H. alisonae; H. schlotheimi Flatstones; Bed 75, Black Ven; Catherstone Lane; Monmouth Beach, Lyme Regis Black Ven Marl Member Complete specimen, isolated wings Coptoclavid beetle
Hypsothemis[37] H. fraseri Flatstones Black Ven Marl Member BMNH In 59109, a hindwing Campterophlebiid damsel-dragonfly, genus also known from China and Kazakhstan
Jurachorista[43] J. bashkuevi Monmouth Beach, Brooki bed Shales with Beef Member ISEA I-F/MP/4/1637/14 Eomeropid scorpionfly
Lateophlebia[37] L. anglicanopsis Flatstones Black Ven Marl Member BMNH In 49573, 59376 Campterophlebiid damsel-dragonfly
Lethonectes[44] L. naucoroides Flatstones Black Ven Marl Member BMNH In 49615, 51014(59383) Belostomatidae giant water bug
Liassocorixa[44] L. dorsetica Flatstones Black Ven Marl Member BMNH In 59166 Corixid bug
Liassocupes[36] L. parvus Flatstones Black Ven Marl Member BMNH In 64008, 49210 Ommatid beetle
Liassophlebia[37] L. pseudomagnifica Stonebarrow Black Ven Marl Member NHMUK In.64000, partial hindwing Liassophlebiid damsel-dragonfly
Locustopsis[37][42] L. spectabilis Flatstones; Monmouth Beach, Brooki bed Black Ven Marl Member ZIN LD/Orth 1, Orth 4, Orth 10, Orth 18, Orth 31 Locustopsid grasshopper
Megarhyphus[45] M. amberae Monmouth Beach, Brooki bed Shales with Beef Member NHML In 28447 Anisopodidae wood gnat
"Mesocixiella"[37] "M." fennahi Flatstones Black Ven Marl Member BMNH In 53942 Fulgoridiid planthopper, does not belong to the genus Mesocixiella
Metaraphidia[37] M. confusa Charmouth (GSM) Black Ven Marl Member GSM 117552 Snakefly
Micromacula[37] M. gracilis Flatstones, Birchi Nodules; Flatstones, Black Ven Shales with Beef; Black Ven Marl Member BMNH In 49230 Regiatid orthopteran
Mimemala[37] M. giganteum Flatstones Black Ven Marl Member BMNH In 51026 Schizocoleid beetle
Nannoblattina[37] N. petulantia Flatstones Black Ven Marl Member BMNH In 53929, 51004 Mesoblattinid cockroach
Nannotanyderus[46] N. oliviae Monmouth Beach, Brooki bed Shales with Beef Member ISEA I-F/MP/2/1621/13, 12 Tanyderid fly
Neomeridium[44] N. trifurcum Woodstones Black Ven Marl Member BMNH In 59151, 51039, 53901, 64014 Pachymeridiid Lygaeoid bug
Oligophryne[47] O. britannica Stonebarrow Hill Black Ven Marl Member Coll. Ansorge fly No 1 Oligophrynidae fly
Omma[37] O. liassicum Flatstones Black Ven Marl Member BMNH 59132 Ommatid beetle, genus extant
Orthophlebia[37] O. capillata Flatstones Black Ven Marl Member BMNH In 53924, an exoskeleton (wings & thorax) Orthophlebiid scorpionfly
Orichalcum[37] O. ornatum Flatstones, Birchi Nodules Shales with Beef BMNH In 53983 Triassomanteidae grasshopper
Paraprosbole[37] P. rotruda Flatstones Black Ven Marl Member BMNH In 59374, 48162, 64395 Tettigarctid cicada
Propreocoris[44] P. maculatus Woodstones Black Ven Marl Member BMNH In 59152 Stem-group to Ochteridae and Gelastocoridae
Protogryllus[37] P. (Archaegryllodes) magnus Flatstones Black Ven Marl Member BMNH In 44003, 51016, 51022, 59373 Protogryllidae cricket
Protohagla[36][37] P. langi Flatstones Black Ven Marl Member BMNH In 59018 Haglid cricket
Protomyrmeleon[41] P. cf. brunonis Charmouth burden pile Black Ven Marl Member AR 428 Protomyrmeleontidae winged insect
Protorthophlebia[37] P. latipennis Flatstones Black Ven Marl Member BMNH In 49575, 53954, 53963, 59120, 59365, 59371, 59384 Protorthophlebiid scorpionfly
Priscaenigma[37] P. obtusa Flatstones Black Ven Marl Member BMNH In 53898 Snakefly
Pseudopolycentropus[36] P. triangularis Woodstones Black Ven Marl Member BMNH In 53915 Pseudopolycentropodid scorpionfly
Pterocimex[44] P. jacksoni Woodstones Black Ven Marl Member BMNH in 53909, 49589, 51048, 59359 Nepomorphan
Regiata[37] R. scutra Flatstones Black Ven Marl Member BMNH In 64027 Regiatid orthopteran
Rhipidoblattina[37] R. ssp. Flatstones Black Ven Marl Member BMNH In 49212, 49249, 53909, 53920 Caloblattinidae cockroach
Rossiphlebia[37] R. jacksoni Flatstones Black Ven Marl Member NHMUK In.53999, part and counterpart of a partial hindwing Liassophlebiid damsel-dragonfly
Tarsabedus[44] T. menkei 0.6 km east Charmouth Black Ven Marl Member BRSMG Cc 688 Belostomatidae giant water bug
Tersus[37] T. crowsoni Flatstones Black Ven Marl Member BMNH In 53949, 59038 Schizophorid beetle

Plants

Genus Species Stratigraphic position Material Notes Images
Ctenozamites[48]
  • C. cycadea
  • Lyme regis
  • Charmouth
 Isolated shoots A "Seed Fern" of the group with Corystospermales
Cycadites[48]
  • C. rectangularis
  • Lyme regis
 Isolated shoots A possible Cycadophyte leaf, related with Cycadales. Alternatively, it can be of Bennettite origin
Cycadeoidea[48]
  • C. gracilis
  • Lyme regis
 Stems Bennettite Trunks of the family Cycadeoidaceae
Example of specimen of the genus
Cycadopteris[48]
  • C. jurensis
  • Lyme regis
 Isolated shoots A "Seed Fern" of the group with Corystospermales
Shoot of the same species
Otozamites[49]
  • O. obtusus
  • Lyme regis
  • Charmouth
  • Seatown
 Isolated shoots Bennettite leafs of the family Williamsoniaceae
Shoot of the same species
Pagiophyllum[49]
  • P. peregrinum
  • P. sewardii
  • Lyme regis
  • Charmouth
  • Seatown
  • Wear Cliffs
 Isolated shoots Conifer shoots of the family Hirmeriellaceae or Araucariaceae
Shoot of the same species

See also

References

  1. ^ British Geological Survey. "Charmouth Mudstone Formation". BGS Lexicon of Named Rock Units. Retrieved 2018-03-08.
  2. ^ Gallois, R.W. (2008). The lithostratigraphy of the Shales-with-Beef Member of the Charmouth Mudstone Formation, Lower Jurassic (PDF). Ussher Society. OCLC 703253954.
  3. ^ "Black Ven Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
  4. ^ "Belemnite Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
  5. ^ "Green Ammonite Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
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