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{{Short description|Subfamily of moths}}
{{Taxobox
{{Automatic taxobox
| name = Cossinae
| name = Cossinae
| image = Prionoxystusrobiniae.JPG
| image = Prionoxystusrobiniae.JPG
| image_caption = [[Prionoxystus robiniae|Carpenterworm]] moth<br/>(''Prionoxystus robiniae'')
| image_width = 200px
| taxon = Cossinae
| image_caption = [[Prionoxystus robiniae|Carpenterworm]] moth<br/>(''[[Prionoxystus robiniae]]'')
| authority = Leach, 1830
| regnum = [[Animal]]ia
| phylum = [[Arthropod]]a
| classis = [[Insect]]a
| ordo = [[Lepidoptera]]
| subordo = [[Glossata]]
| infraordo = [[Heteroneura]]
| unranked_superfamilia = [[Apoditrysia]]
| superfamilia = [[Cossoidea]]
| familia = [[Cossidae]]
| subfamilia = '''Cossinae'''
| subfamilia_authority = Leach, 1830
| subdivision_ranks = [[Genera]]
| subdivision_ranks = [[Genera]]
| subdivision =
| subdivision = Numerous, see [[#Selected genera|text]]
Numerous, see [[#Selected genera|text]]
}}
}}


The '''Cossinae''' are the nominate [[subfamily]] of the [[Cossidae]] (carpenter or goat moths). The [[caterpillar]]s of several Cossinae [[species]], such as the [[Prionoxystus robiniae|Carpenterworm]] (''[[Prionoxystus robiniae]]'') and the [[Goat Moth]] (''Cossus cossus''), are significant [[Pest (organism)|pest]]s. On the other hand, in [[Chile]] the caterpillars of the [[Chilean Moth]] (''[[Chilecomadia moorei]]'') are collected on a commercial scale for sale as [[fishing bait]] and [[terrarium]] [[pet food]]; they are usually called "[[butterworm]]s" in international trade.
The '''Cossinae''' are the nominate [[subfamily]] of the [[Cossidae]] (carpenter or goat moths). The [[caterpillar]]s of several Cossinae [[species]], such as the carpenterworm (''[[Prionoxystus robiniae]]'') and the goat moth (''[[Cossus cossus]]''), are significant [[Pest (organism)|pest]]s. On the other hand, in [[Chile]] the caterpillars of the Chilean moth (''[[Chilecomadia moorei]]'') are collected on a commercial scale for sale as [[fishing bait]] and [[terrarium]] [[pet food]]; they are usually called "butterworms" in international trade.


The [[Cossulinae]] have been separated from the Cossinae in recent decades, but this was not universally accepted at first. Some misplaced genera have been moved between the subfamilies, and as it seems the Cossulinae at least now represent a [[monophyletic]] group.
The [[Cossulinae]] have been separated from the Cossinae in recent decades, but this was not universally accepted at first. Some misplaced genera have been moved between the subfamilies, and as it seems the Cossulinae at least now represent a [[monophyletic]] group.


==Selected genera==
==Systematics==
Some significant species are also listed:<!-- list species here when they have articles but no genus article yet --><ref>See references in Savela (2008)</ref>
Some significant species are also listed:<!-- list species here when they have articles but no genus article yet --><ref>See references in Savela (2008)</ref>


*Unplaced to tribe/placement unknown
* ''[[Acossus]]'' <small>Dyar, 1905</small> (tentatively placed here)
** ''[[Citharalia]]'' <small>Clench, 1957</small> (formerly in Cossulinae)
** ''[[Comadia]]'' <small>Barnes & McDunnough, 1911</small>
** ''[[Cossodes]]'' <small>White, 1841</small>
** ''[[Culama]]'' <small>Walker, 1856</small>
** ''[[Dieidapersa]]'' <small>Strand in Stichel, 1911</small>
** ''[[Dieida]]'' <small>Strand, 1911</small>
** ''[[Fania (moth)|Fania]]'' <small>Barnes & McDunnough, 1911</small> (tentatively placed here)
** ''[[Idioses]]'' <small>Turner, 1927</small>
** ''[[Macrocyttara]]'' <small>Turner, 1918</small>
** ''[[Miacora]]'' <small>Dyar, 1905</small> (tentatively placed here)
** ''[[Prionoxystus]]'' <small>Grote, 1882</small><!-- AnimBehav62:349. -->
** ''[[Psychidocossus]]'' <small>D. S. Fletcher, 1982</small>
** ''[[Surcossus]]'' <small>Heimlich, 1960</small>
** ''[[Trigena]]'' <small>Dyar, 1905</small> (formerly in Cossulinae)
** ''[[Zyganisus]]'' <small>Viette, 1951</small>
* Zeuzerocossini <small>Yakovlev, 2008</small>
** ''[[Acossus]]'' <small>Dyar, 1905</small>
** ''[[Alcterogystia]]'' <small>Schoorl, 1990</small>
** ''[[Arctiocossus]]'' <small>Felder, 1874</small>
** ''[[Assegaj]]'' <small>Yakovlev, 2006</small>
** ''[[Brachygystia]]'' <small>Schoorl, 1990</small>
** ''[[Chingizid (genus)|Chingizid]]'' <small>Yakovlev, 2011</small>
** ''[[Chinocossus]]'' <small>Yakovlev, 2006</small>
** ''[[Dervishiya]]'' <small>Yakovlev, 2006</small>
** ''[[Eogystia]]'' <small>Schoorl, 1990</small>
** ''[[Eremocossus]]'' <small>Hampson, 1893</small>
** ''[[Gobibatyr]]'' <small>Yakovlev, 2004</small>
** ''[[Kalimantanossus]]'' <small>Yakovlev, 2011</small>
** ''[[Mahommedella]]'' <small>Yakovlev, 2011</small>
** ''[[Neurocossus]]'' <small>Yakovlev, 2010</small>
** ''[[Paropta]]'' <small>Staudinger, 1900</small>
** ''[[Patoptoformis]]'' <small>Yakovlev, 2006</small>
** ''[[Planctogystia]]'' <small>Schoorl, 1990</small>
** ''[[Rethona]]'' <small>Walker, 1855</small>
** ''[[Ronaldocossus]]'' <small>Yakovlev, 2006</small>
** ''[[Wiltshirocossus]]'' <small>Yakovlev, 2007</small>
** ''[[Zeuzerocossus]]'' <small>Yakovlev, 2008</small>
* Endagriini <small>Duponchel, [1845]</small>
** ''[[Afroarabiella]]'' <small>Yakovlev, 2008</small>
** ''[[Dyspessa]]'' <small>Hübner, 1820</small>
** ''[[Groenendaelia]]'' <small>Yakovlev, 2004</small>
** ''[[Hollowiella]]'' <small>Yakovlev & Witt, 2009</small>
** ''[[Isoceras]]'' <small>Turati, 1924</small>
** ''[[Paracossulus]]'' <small>Schoorl, 1990</small>
** ''[[Stygoides]]'' <small>Bruand, 1853</small>
** ''[[Samagystia]]'' <small>Schoorl, 1990</small>
* Holcocerini <small>Yakovlev, 2006</small>
** ''[[Afrikanetz]]'' <small>Yakovlev, 2009</small>
** ''[[Aholcocerus]]'' <small>Yakovlev, 2006</small>
** ''[[Barchaniella]]'' <small>Yakovlev, 2006</small>
** ''[[Brachylia]]'' <small>Felder, 1874</small>
** ''[[Camellocossus]]'' <small>Yakovlev, 2011</small>
** ''[[Coryphodema]]'' <small>Felder, 1874</small>
** ''[[Cossulus]]'' <small>Staudinger, 1887</small>
** ''[[Cossus]]'' <small>Fabricius, 1794</small>
** ''[[Cryptoholcocerus]]'' <small>Yakovlev, 2006</small>
** ''[[Deserticossus]]'' <small>Yakovlev, 2006</small>
** ''[[Dyspessacossus]]'' <small>Daniel, 1953</small>
** ''[[Franzdanielia]]'' <small>Yakovlev, 2006</small>
** ''[[Gumilevia]]'' <small>Yakovlev, 2011</small>
** ''[[Hirtocossus]]'' <small>Schoorl, 1990</small>
** ''[[Holcocerus]]'' <small>Staudinger, 1884</small>
** ''[[Isocossus]]'' <small>Roepke, 1957</small>
** ''[[Kerzhnerocossus]]'' <small>Yakovlev, 2011</small>
** ''[[Koboldocossus]]'' <small>Yakovlev, 2011</small>
** ''[[Kotchevnik]]'' <small>Yakovlev, 2004</small>
** ''[[Mirocossus]]'' <small>Schoorl, 1990</small>
** ''[[Macrocossus]]'' <small>Aurivillius, 1900</small>
** ''[[Mormogystia]]'' <small>Schoorl, 1990</small>
** ''[[Paracossus]]'' <small>Hampson, 1904</small>
** ''[[Parahypopta]]'' <small>Daniel, 1961</small>
** ''[[Plyustchiella]]'' <small>Yakovlev, 2006</small>
** ''[[Pygmeocossus]]'' <small>Yakovlev, 2005</small>
** ''[[Rambuasalama]]'' <small>Yakovlev & Saldaitis, 2008</small>
** ''[[Reticulocossus]]'' <small>Yakovlev, 2011</small>
** ''[[Roepkiella]]'' <small>Yakovlev & Witt, 2009</small>
** ''[[Semitocossus]]'' <small>Yakovlev, 2007</small>
** ''[[Streltzoviella]]'' <small>Yakovlev, 2006</small>
** ''[[Sundacossus]]'' <small>Yakovlev, 2006</small>
** ''[[Vartiania]]'' <small>Yakovlev, 2004</small>
** ''[[Wittocossus]]'' <small>Yakovlev, 2004</small>
** ''[[Yakudza]]'' <small>Yakovlev, 2006</small>

==Former genera==
* ''[[Bifiduncus]]''
* ''[[Butaya]]'' <small>Yakovlev, 2004</small>
* ''[[Catopta]]'' <small>Staudinger, 1899</small>
* ''[[Catopta]]'' <small>Staudinger, 1899</small>
* ''[[Chiangmaiana]]'' <small>Kemal & Koçak, 2005</small> (≡''Nirvana'' <small>Yakovlev, 2004 nom. invalid.(preoc.name) non Stål, 1859 nec Kirkaldy, 1900 nec Yakovlev, 2007 nec Tsukuda & Nishiyama, 1979</small>);(≡''Nirrvanna'' <small>Yakovlev, 2007 nom. invalid.( junior object. syn.), objective replacement name (mistaken repl.n.))</small>
* ''[[Citharalia]]'' <small>Clench, 1957</small> (formerly in Cossulinae)
* ''[[Chilecomadia]]''
* ''[[Chilecomadia]]''
* ''[[Comadia]]'' <small>Barnes & McDunnough, 1911</small>
* ''[[Cossulus]]'' <small>Staudinger, 1887</small> (tentatively placed here)
* ''[[Cossus]]'' <small>Fabricius, 1794</small>
* ''[[Culama]]''
* ''[[Danielostygia]]'' <small>Reisser, 1962</small> (tentatively placed here)
* ''[[Danielostygia]]'' <small>Reisser, 1962</small> (tentatively placed here)
* ''[[Dyspessa]]'' <small>Hübner, 1820</small>
* ''[[Dyspessacossus]]'' <small>Daniel, 1953</small> (tentatively placed here)
* ''[[Fania (moth)|Fania]]'' <small>Barnes & McDunnough, 1911</small> (tentatively placed here)
* ''[[Idioses]]''
* ''[[Isoceras]]'' <small>Turati, 1924</small> (tentatively placed here)
* ''[[Lamellocossus]]'' <small>Daniel, 1956</small> (tentatively placed here)
* ''[[Lamellocossus]]'' <small>Daniel, 1956</small> (tentatively placed here)
* ''[[Toronia (moth)|Toronia]]'' <small>Barnes & McDunnough, 1911</small>
* ''[[Macrocyttara]]''
* ''[[Miacora]]'' <small>Dyar, 1905</small> (tentatively placed here)
* ''[[Yakovlevina]]'' <small>Kemal & Koçak, 2005</small> (=''Garuda'' <small>Yakovlev, 2005</small>)
* ''[[Parahypopta]]'' <small>Daniel, 1961</small>
* ''[[Patoptoformis]]'' <small>Yakovlev, 2006</small>
* ''[[Paropta]]'' <small>Staudinger, 1900</small> (tentatively placed here)
* ''[[Prionoxystus]]'' <small>Grote, 1882</small><!-- AnimBehav62:349. -->
** ''[[Prionoxystus robiniae]]'' &ndash; [[Carpenterworm]]
* ''[[Samagystia]]'' <small>Schoorl, 1990</small> (tentatively placed here)
* ''[[Schreiteriana]]'' <small>Fletcher, 1982</small> (formerly in Cossulinae)
* ''[[Stygia (genus)|Stygia]]'' <small>Latreille, 1803</small> (tentatively placed here)
* ''[[Stygia (genus)|Stygia]]'' <small>Latreille, 1803</small> (tentatively placed here)
* ''[[Stygoides]]'' <small>Bruand, 1853</small> (tentatively placed here)
* ''[[Surcossus]]''
* ''[[Trigena]]'' <small>Dyar, 1905</small> (formerly in Cossulinae)


==Footnotes==
==Footnotes==
Line 60: Line 117:


==References==
==References==
{{commons category}}
{{Commons category}}
* {{aut|Savela, Markku}} (2008): Markku Savela's Lepidoptera and some other life forms – [http://www.nic.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/cossoidea/cossidae/cossinae/index.html Cossinae]. Version of 2008-JUL-17. Retrieved 2009-FEB-27.
* {{aut|Savela, Markku}} (2008): ''Lepidoptera and Some Other Life Forms'' [http://www.nic.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/cossoidea/cossidae/cossinae/index.html Cossinae]. Version of 17 July 2008. Retrieved 27 February 2009.
* [http://eversmannia.entomology.ru/eversmannia_dop1.pdf Ревизия древоточцев рода Holcocerus Staudinger, 1884 (s. l.)]


{{Taxonbar|from=Q5174707}}
[[Category:Cossidae]]


[[Category:Cossinae| ]]
{{Cossoidea-stub}}
[[Category:Moth subfamilies]]

Latest revision as of 17:55, 10 January 2021

Cossinae
Carpenterworm moth
(Prionoxystus robiniae)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Cossidae
Subfamily: Cossinae
Leach, 1830
Genera

Numerous, see text

The Cossinae are the nominate subfamily of the Cossidae (carpenter or goat moths). The caterpillars of several Cossinae species, such as the carpenterworm (Prionoxystus robiniae) and the goat moth (Cossus cossus), are significant pests. On the other hand, in Chile the caterpillars of the Chilean moth (Chilecomadia moorei) are collected on a commercial scale for sale as fishing bait and terrarium pet food; they are usually called "butterworms" in international trade.

The Cossulinae have been separated from the Cossinae in recent decades, but this was not universally accepted at first. Some misplaced genera have been moved between the subfamilies, and as it seems the Cossulinae at least now represent a monophyletic group.

Systematics

[edit]

Some significant species are also listed:[1]

Former genera

[edit]
  • Bifiduncus
  • Butaya Yakovlev, 2004
  • Catopta Staudinger, 1899
  • Chiangmaiana Kemal & Koçak, 2005 (≡Nirvana Yakovlev, 2004 nom. invalid.(preoc.name) non Stål, 1859 nec Kirkaldy, 1900 nec Yakovlev, 2007 nec Tsukuda & Nishiyama, 1979);(≡Nirrvanna Yakovlev, 2007 nom. invalid.( junior object. syn.), objective replacement name (mistaken repl.n.))
  • Chilecomadia
  • Danielostygia Reisser, 1962 (tentatively placed here)
  • Lamellocossus Daniel, 1956 (tentatively placed here)
  • Toronia Barnes & McDunnough, 1911
  • Yakovlevina Kemal & Koçak, 2005 (=Garuda Yakovlev, 2005)
  • Stygia Latreille, 1803 (tentatively placed here)

Footnotes

[edit]
  1. ^ See references in Savela (2008)

References

[edit]