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'''Harris's hawk''' ('''''Parabuteo unicinctus'''''), formerly known as the '''bay-winged hawk''', '''dusky hawk''', and sometimes a '''wolf hawk''', and known in Latin America as '''peuco''', is a medium-large [[bird of prey]] that breeds from the [[southwestern United States]] south to [[Chile]], central [[Argentina]], and [[Brazil]]. This bird is sometimes reported to be at large in [[Western Europe]], especially [[Great Britain|Britain]], but it is a popular species in [[falconry]] and these records almost invariably all refer to escapes from [[captivity (animal)|captivity]].
'''Harris's hawk''' ('''''Parabuteo unicinctus'''''), formerly known as the '''bay-winged hawk''' or '''dusky hawk''', and known in Latin America as the '''peuco''', is a medium-large [[bird of prey]] that breeds from the [[southwestern United States]] south to [[Chile]], central [[Argentina]], and [[Brazil]]. This bird is sometimes reported to be at large in [[Western Europe]], especially [[Great Britain|Britain]], but it is a popular species in [[falconry]] and these records almost invariably all refer to escapes from [[captivity (animal)|captivity]].


The name is derived from the [[Greek language|Greek]] ''para'', meaning beside, near or like, and the [[Latin]] ''buteo'', referring to a kind of buzzard; ''uni'' meaning once; and ''cinctus'' meaning girdled, referring to the white band at the tip of the tail.<ref>{{cite book |last=Jobling |first=James A. |title=A Dictionary of Scientific Bird Names |year=1991 |isbn=978-0-19-854634-4}}</ref> [[John James Audubon]] gave this bird its English name in honor of his ornithological companion, financial supporter, and friend [[Edward Harris (ornithologist)|Edward Harris]].<ref>{{cite web |author=National Audubon Society |url=http://www.audubon.org/bird/BoA/F2_G2a.html |title=Audubon |publisher=Audubon |access-date=2013-04-29 |archive-url=https://web.archive.org/web/20080409153815/http://www.audubon.org/bird/boa/F2_G2a.html |archive-date=2008-04-09 |url-status=dead }}</ref>
The name is derived from the [[Greek language|Greek]] ''para'', meaning beside, near or like, and the [[Latin]] ''buteo'', referring to a kind of buzzard; ''uni'' meaning once; and ''cinctus'' meaning girdled, referring to the white band at the tip of the tail.<ref>{{cite book |last=Jobling |first=James A. |title=A Dictionary of Scientific Bird Names |year=1991 |publisher=Oxford University Press |isbn=978-0-19-854634-4}}</ref> [[John James Audubon]] gave this bird its English name in honor of his ornithological companion, financial supporter, and friend [[Edward Harris (ornithologist)|Edward Harris]].<ref>{{cite web |author=National Audubon Society |url=http://www.audubon.org/bird/BoA/F2_G2a.html |title=Audubon |publisher=Audubon |access-date=2013-04-29 |archive-url=https://web.archive.org/web/20080409153815/http://www.audubon.org/bird/boa/F2_G2a.html |archive-date=2008-04-09 |url-status=dead }}</ref>


Harris's hawk is notable for its behavior of hunting cooperatively in [[Pack hunter|packs]] consisting of tolerant groups, while other raptors often hunt alone. Harris hawks' social nature has been attributed to their intelligence, which makes them easy to train and has made them a popular bird for use in falconry.<ref name=beebe>Beebe, Frank (1984). ''A Falconry Manual.'' Hancock House Publishers, {{ISBN|0-88839-978-2}}, page 81.</ref>
Harris's hawk is notable for its behavior of hunting cooperatively in [[Pack hunter|packs]] consisting of tolerant groups, while other raptors often hunt alone. Harris's hawks' social nature has been attributed to their intelligence, which makes them easy to train and has made them a popular bird for use in falconry.<ref name=beebe>Beebe, Frank (1984). ''A Falconry Manual.'' Hancock House Publishers, {{ISBN|0-88839-978-2}}, page 81.</ref>


==Description==
==Description==
[[File:Harris' Hawk plumage.jpg|thumb|left|The distinctive plumage and tail feathers clearly confirm that this is a Harris's hawk]]
[[File:Harris' Hawk plumage.jpg|thumb|left|The distinctive plumage and tail feathers clearly confirm that this is a Harris's hawk]]


This medium-large hawk is roughly intermediate in size between a [[peregrine falcon]] (''Falco peregrinus'') and a [[red-tailed hawk]] (''Buteo jamaicensis''). Harris's hawks range in length from {{convert|46|to|59|cm|in|abbr=on}} and generally have a wingspan of about {{convert|103|to|120|cm|in|abbr=on}}.<ref name=Mikos>{{cite book |last=Udvardy |first=Miklos D. F. |title=National Audubon Society Field Guide to North American Birds 'Western Region' |year=2001 |isbn=978-0-679-42851-0}}</ref><ref>Clark, W. S. and B. K. Wheeler (1987). ''A Field Guide to Hawks of North America''. Houghton Mifflin Co. Boston.</ref> These hawks have a brownish plumage, reddish shoulders, and tail feathers with a white base and white tip.<ref>{{cite web |title=Harris's Hawk - Appearance |year=2015 |doi=10.2173/bow.hrshaw.01 |s2cid=216204185 |url=https://birdsna.org/Species-Account/bna/species/hrshaw/introduction |url-access=subscription |publisher=|access-date=October 22, 2018}}</ref>
This medium-large [[hawk]] is roughly intermediate in size between a [[peregrine falcon]] (''Falco peregrinus'') and a [[red-tailed hawk]] (''Buteo jamaicensis''). Harris's hawks range in length from {{convert|46|to|59|cm|in|abbr=on}} and generally have a wingspan of about {{convert|103|to|120|cm|in|abbr=on}}.<ref name=Mikos>{{cite book |last=Udvardy |first=Miklos D. F. |title=National Audubon Society Field Guide to North American Birds 'Western Region' |year=2001 |publisher=Knopf Doubleday Publishing |isbn=978-0-679-42851-0}}</ref><ref>Clark, W. S. and B. K. Wheeler (1987). ''A Field Guide to Hawks of North America''. Houghton Mifflin Co. Boston.</ref> These hawks have a brownish plumage, reddish shoulders, and tail feathers with a white base and white tip.<ref>{{cite journal |title=Harris's Hawk - Appearance |journal=Birds of the World |year=2015 |doi=10.2173/bow.hrshaw.01 |s2cid=216204185 |url=https://birdsna.org/Species-Account/bna/species/hrshaw/introduction |url-access=subscription |access-date=October 22, 2018 |last1=Dwyer |first1=James F. |last2=Bednarz |first2=James C. }}</ref>


They exhibit [[sexual dimorphism]] with the females being larger by about 35%. In the United States, the average weight for adult males is about {{convert|701|g|lb|abbr=on}}, with a range of {{convert|546|to|850|g|lb|abbr=on}}, while the adult female average is {{convert|1029|g|lb|abbr=on}}, with a range of {{convert|766|to|1633|g|lb|abbr=on}}.<ref>{{cite book |last=Dunning |first=John B. Jr. |title=CRC Handbook of Avian Masses |year=1993 |isbn=978-0-8493-4258-5}}</ref><ref name= Hamerstrom>Hamerstrom, F. (1978). "External sex characters of Harris' Hawks in winter". ''Raptor Res.'' 12:1–14.</ref> They have dark brown [[plumage]] with chestnut shoulders, wing linings, and thighs,<ref>{{cite book |author=National Geographic Society |title=Birds of North America |year=1983 |isbn=978-0-87044-472-2}}</ref> white on the base and tip of the tail,<ref>{{cite book |last=Sibley |first=David Allen |title=National Audubon Society: The Sibley Guide to Birds |year=2000 |isbn=978-0-679-45122-8 |url=https://archive.org/details/sibleyguidetobir00sibl_0 }}</ref> long, yellow legs, and a yellow [[beak|cere]].<ref name=rappole>{{cite book |last=Rappole |first=John H. |title=Birds of the Southwest |year=2000 |isbn=978-0-89096-958-8}}</ref> The vocalizations of Harris's hawk are very harsh sounds.<ref name= Mikos/>
They exhibit [[sexual dimorphism]] with the females being larger by about 35%. In the United States, the average weight for adult males is about {{convert|701|g|lb|abbr=on}}, with a range of {{convert|546|to|850|g|lb|abbr=on}}, while the adult female average is {{convert|1029|g|lb|abbr=on}}, with a range of {{convert|766|to|1633|g|lb|abbr=on}}.<ref>{{cite book |last=Dunning |first=John B. Jr. |title=CRC Handbook of Avian Masses |year=1993 |publisher=CRC Press |isbn=978-0-8493-4258-5}}</ref><ref name= Hamerstrom>Hamerstrom, F. (1978). "External sex characters of Harris' Hawks in winter". ''Raptor Res.'' 12:1–14.</ref> They have dark brown [[plumage]] with chestnut shoulders, wing linings, and thighs,<ref>{{cite book |author=National Geographic Society |title=Birds of North America |year=1983 |publisher=National Geographic Society |isbn=978-0-87044-472-2}}</ref> white on the base and tip of the tail,<ref>{{cite book |last=Sibley |first=David Allen |title=National Audubon Society: The Sibley Guide to Birds |year=2000 |publisher=Knopf |isbn=978-0-679-45122-8 |url=https://archive.org/details/sibleyguidetobir00sibl_0 }}</ref> long, yellow legs, and a yellow [[beak|cere]].<ref name=rappole>{{cite book |last=Rappole |first=John H. |title=Birds of the Southwest |year=2000 |publisher=Texas A&M University Press |isbn=978-0-89096-958-8}}</ref> The vocalizations of Harris's hawk are very harsh sounds.<ref name= Mikos/>


The lifespan of Harris's Hawk is 10–12 years in the wild and 20–25 years in captivity.<ref>{{cite web |url=https://www.oregonzoo.org/discover/animals/harris-hawk |publisher=Oregon Zoo |access-date=22 November 2021|title=Harris hawk }}</ref>
The lifespan of Harris's Hawk is 10–12 years in the wild and 20–25 years in captivity.<ref>{{cite web |url=https://www.oregonzoo.org/discover/animals/harris-hawk |publisher=Oregon Zoo |access-date=22 November 2021 |title=Harris hawk |archive-date=21 June 2022 |archive-url=https://web.archive.org/web/20220621084032/https://www.oregonzoo.org/discover/animals/harris-hawk |url-status=dead }}</ref>


===Juvenile===
===Juvenile===
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The juvenile Harris's hawk is mostly streaked with [[Buff (colour)|buff]] and appears much lighter than the dark adults. When in flight, the undersides of the juveniles' wings are buff-colored with brown streaking. They can look unlike adults at first glance, but the identical chestnut [[plumage]] is an aid for identification.<ref name=rappole/>{{Verify source|date=May 2010}}<!-- If Rappole has no detailed description, "Raptors of the World" ("Helm Guide") or something else that does should be added as source. HBW is not very comprehensive on this species. -->
The juvenile Harris's hawk is mostly streaked with [[Buff (colour)|buff]] and appears much lighter than the dark adults. When in flight, the undersides of the juveniles' wings are buff-colored with brown streaking. They can look unlike adults at first glance, but the identical chestnut [[plumage]] is an aid for identification.<ref name=rappole/>{{Verify source|date=May 2010}}<!-- If Rappole has no detailed description, "Raptors of the World" ("Helm Guide") or something else that does should be added as source. HBW is not very comprehensive on this species. -->
[[File:Parabuteo unicinctus unicinctus (recorte).jpg|thumb|'''''Unicinctus''''' subspecies (Bay-winged hawk), in Argentina.]]
There are three subspecies of Harris's hawk:



[[File:Parabuteo unicinctus unicinctus.jpg|thumb|'''''Unicinctus''''' subspecies (Bay-winged hawk), in Argentina.]]
There are three subspecies of Harris's hawk:
===Subspecies===
===Subspecies===
{| class="wikitable "
* ''P. u. superior'': found in [[Baja California]], [[Arizona]], [[Sonora]], and [[Sinaloa]]. ''P. u. superior'' was believed to have longer tails and wings and to be more blackish than ''P. u. harrisi''. However, the sample size of the original study was quite small, with only five males and six females. Later research has concluded that there is not as strong a physical difference as was originally assumed.<ref name=Bednarz>{{cite conference |last=Bednarz |first=J. C. |title=Harris' hawk subspecies: is ''superior'' larger or different than ''harrisi''? |pages=294–300 |year=1988 |book-title=in ''Proceedings of the southwest raptor management symposium and workshop'' |place= Washington, D.C.}}</ref> Other ecological differences, such as [[latitudinal cline]] were also brought up as arguments against the validity of the subspecies segmentation.<ref name=cornell>{{cite web |last=Bednarz |first=James C. |title=Harris's Hawk (''Parabuteo unicinctus'') |year=1995 |url=http://bna.birds.cornell.edu/bna/species/146}}</ref>
|-
* ''P. u. harrisi'': found in [[Texas]], eastern [[Mexico]], and much of [[Central America]].<ref name=cornell/>
! Image !! Subspecies !! Description !!Distribution
* ''P. u. unicinctus'': found exclusively in [[South America]]. It is smaller than the North American subspecies and the adult's dark brown ventrum is streaked or flecked with white or whitish. <ref> Blake, Ferguson-Lees and Christie cited in ''Birds of the world "Parabuteo unicinctrus"'' https://birdsoftheworld.org/bow/species/hrshaw/cur/systematics#subsp</ref>
|-
|[[File:Harris's Hawk Arizona.jpg|120px]]||''P. u. superior'' || ''P. u. superior'' was believed to have longer tails and wings and to be more blackish than ''P. u. harrisi''. However, the sample size of the original study was quite small, with only five males and six females. Later research has concluded that there is not as strong a physical difference as was originally assumed.<ref name=Bednarz>{{cite conference |last=Bednarz |first=J. C. |title=Harris' hawk subspecies: is ''superior'' larger or different than ''harrisi''? |pages=294–300 |year=1988 |book-title=in ''Proceedings of the southwest raptor management symposium and workshop'' |place= Washington, D.C.}}</ref> Other ecological differences, such as [[latitudinal cline]], were also brought up as arguments against the validity of the subspecies segmentation.<ref name=cornell>{{cite web |last=Bednarz |first=James C. |title=Harris's Hawk (''Parabuteo unicinctus'') |year=1995 |url=http://bna.birds.cornell.edu/bna/species/146}}</ref> ||[[Baja California]], [[Arizona]], [[Sonora]], and [[Sinaloa]].
|-
|[[File:Harris' Hawk (Parabuteo unicinctus) (10684271643).jpg|120px]]||''P. u. harrisi'' ||||[[Texas]], eastern [[Mexico]], and much of [[Central America]].<ref name=cornell/>
|-
|[[File:Gavilán mixto en barrio Savoia, City Bell, La Plata, Argentina IMG 6649.jpg|120px]]||''P. u. unicinctus'' ||It is smaller than the North American subspecies, tail and wings are proportionally longer, and the adult's dark brown ventrum is streaked or flecked with white or whitish. <ref> Blake, Ferguson-Lees and Christie cited in ''Birds of the world "Parabuteo unicinctrus"'' https://birdsoftheworld.org/bow/species/hrshaw/cur/systematics#subsp</ref>|| [[South America]].
|-
|}


==Taxonomy==
==Taxonomy==
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[[File:Aa babyharrishawks 00.jpg|left|thumb|At one week of age]]
[[File:Aa babyharrishawks 00.jpg|left|thumb|At one week of age]]


They nest in small trees, shrubby growth, or cacti. The [[bird nest|nests]] are often compact, made of sticks, plant roots, and stems and are often lined with leaves, [[moss]], bark, and plant roots. They are built mainly by the female. There are usually two to four white to blueish-white [[bird egg|eggs]] sometimes with a speckling of pale [[brown]] or gray. The nestlings start light buff, but in five to six days turn a rich brown.<ref name=baicich1997>{{cite book |last1=Baicich |first1=Paul J. |last2=Harrison |first2=Colin J. O. |title=Nests, Eggs, and Nestlings of North American Birds |year=1997 |isbn=978-0-691-12295-3}}</ref>
They nest in small trees, shrubby growth, or cacti. The [[bird nest|nests]] are often compact, made of sticks, plant roots, and stems and are often lined with leaves, [[moss]], bark, and plant roots. They are built mainly by the female. There are usually two to four white to blueish-white [[bird egg|eggs]] sometimes with a speckling of pale [[brown]] or gray. The nestlings start light buff, but in five to six days turn a rich brown.<ref name=baicich1997>{{cite book |last1=Baicich |first1=Paul J. |last2=Harrison |first2=Colin J. O. |title=Nests, Eggs, and Nestlings of North American Birds |year=1997 |publisher=Princeton University Press |isbn=978-0-691-12295-3}}</ref>


Very often, there will be three hawks attending one nest: two males and one female.<ref name=Kenn>{{cite book |last=Kaufmann |first=Kenn |title=Lives of North American Birds |year=1996 |isbn=978-0-395-77017-7 |url=https://archive.org/details/livesofnorthamer00kauf }}</ref> Whether or not this is [[polyandry]] is debated, as it may be confused with backstanding (one bird standing on another's back).<ref>{{cite book |last=Ligon |first=J. David |title=The Evolution of Avian Breeding Systems |volume=10 |series=Oxford Ornithology Series |year=1999 |isbn=978-0198549130 |publisher=Oxford University Press |url=http://ukcatalogue.oup.com/product/9780198549130.do |access-date=4 August 2013}}</ref> The female does most of the [[egg incubation|incubation]]. The eggs hatch in 31 to 36 days. The young begin to explore outside the nest at 38 days, and [[fledge]], or start to fly, at 45 to 50 days. The female sometimes breeds two or three times in a year.<ref name=baicich1997/> Young may stay with their parents for up to three years, helping to raise later broods. Nests are known to be predated by [[coyote]]s (''Canis latrans''), [[golden eagle]]s (''Aquila chrysaetos''), [[red-tailed hawk]]s (''Buteo jamaicensis''), [[great horned owl]]s (''Bubo virginianus''), and flocks of [[common raven]]s (''Corvus corax''), predators possibly too formidable to be fully displaced by Harris's hawk's cooperative nest defenses. No accounts show predation on adults in the United States and Harris's hawk may be considered an [[apex predator]], although presumably predators like eagles and great horned owls would be capable of killing them.<ref>Dawson, J. W. and R. W. Mannan. (1991). ''Dominance hierarchies and helper contributions in Harris' Hawks''. ''Auk'' 108:649–660.</ref> In [[Chile]], [[black-chested buzzard-eagle]]s (''Geranoaetus melanoleucus'') are likely predators.<ref>Jiménez, J. E., & Jaksić, F. M. (1989). Behavioral ecology of grey eagle-buzzards, Geranoaetus melanoleucus, in central Chile. ''Condor'' 913–921.</ref>
Very often, there will be three hawks attending one nest: two males and one female.<ref name=Kenn>{{cite book |last=Kaufmann |first=Kenn |title=Lives of North American Birds |year=1996 |publisher=Houghton Mifflin Company |isbn=978-0-395-77017-7 |url=https://archive.org/details/livesofnorthamer00kauf }}</ref> Whether or not this is [[Polyandry in animals|polyandry]] is debated, as it may be confused with backstanding (one bird standing on another's back).<ref>{{cite book |last=Ligon |first=J. David |title=The Evolution of Avian Breeding Systems |volume=10 |series=Oxford Ornithology Series |year=1999 |isbn=978-0198549130 |publisher=Oxford University Press |url=http://ukcatalogue.oup.com/product/9780198549130.do |access-date=4 August 2013}}</ref> The female does most of the [[egg incubation|incubation]]. The eggs hatch in 31 to 36 days. The young begin to explore outside the nest at 38 days, and [[fledge]], or start to fly, at 45 to 50 days. The female sometimes breeds two or three times in a year.<ref name=baicich1997/> Young may stay with their parents for up to three years, helping to raise later broods. Nests are known to be predated by [[coyote]]s (''Canis latrans''), [[golden eagle]]s (''Aquila chrysaetos''), [[red-tailed hawk]]s (''Buteo jamaicensis''), [[great horned owl]]s (''Bubo virginianus''), and flocks of [[common raven]]s (''Corvus corax''), predators possibly too formidable to be fully displaced by Harris's hawk's cooperative nest defenses. No accounts show predation on adults in the United States and Harris's hawk may be considered an [[apex predator]], although presumably predators like eagles and great horned owls would be capable of killing them.<ref>Dawson, J. W. and R. W. Mannan. (1991). ''Dominance hierarchies and helper contributions in Harris' Hawks''. ''Auk'' 108:649–660.</ref> In [[Chile]], [[black-chested buzzard-eagle]]s (''Geranoaetus melanoleucus'') are likely predators.<ref>Jiménez, J. E., & Jaksić, F. M. (1989). Behavioral ecology of grey eagle-buzzards, Geranoaetus melanoleucus, in central Chile. ''Condor'' 913–921.</ref>


===Feeding===
===Feeding===
[[File:Harriss Hawk being with chick leg (7913337978).jpg|thumb|Eating a chick's leg]]
[[File:Harriss Hawk being with chick leg (7913337978).jpg|thumb|Eating a chick's leg]]


The majority of Harris hawk's prey are mammals, including [[ground squirrels]], [[rabbit]]s, and larger [[black-tailed jackrabbit]]s (''Lepus californicus'').<ref name = world>Del Hoyo, Josep, et al. Handbook of the birds of the world. Vol. 1. No. 8. Lynx Ed., 1992.</ref> [[Bird]]s from the size of small passerines such as [[diuca finch]] (''Diuca diuca'') to adult [[great egret]] (''Ardea alba'') and half-grown [[wild turkey]] (''Meleagris gallapavo'') can be taken.<ref name =raptor>Ferguson-Lees, James, and David A. Christie. Raptors of the world. Houghton Mifflin Harcourt, 2001.</ref><ref>Santander, Francisco J., et al. "PREY OF HARRIS'HAWKS (PARABUTEO UNICINCTUS) DURING AUTUMN AND WINTER IN A COASTAL AREA OF CENTRAL CHILE." The Southwestern Naturalist (2011): 417-422.</ref><ref>Houcke, H.H. (1971). Predation By a White-Tailed Hawk and a Harris' Hawk on a Wild Turkey Poult ''Condor'' 4: 475.</ref> In one instance, the lone Harris hawk successfully killed a subadult [[great blue heron]] (''Ardea herodias'').<ref>Woodward, H.D. (2003). Lone Harris' Hawk Kills Great Blue Heron. The Raptor Research Foundation 1:85–86.</ref> Reptiles such as [[lizard]]s and [[snake]]s are additionally taken as well as large [[insect]]s.<ref name = world/><ref name =raptor/>
The majority of Harris's hawks' prey are mammals, including [[ground squirrels]], [[rabbit]]s, and larger [[black-tailed jackrabbit]]s (''Lepus californicus'').<ref name = world>Del Hoyo, Josep, et al. Handbook of the birds of the world. Vol. 1. No. 8. Lynx Ed., 1992.</ref> [[Bird]]s from the size of small passerines such as [[diuca finch]] (''Diuca diuca'') to adult [[great egret]] (''Ardea alba'') and half-grown [[wild turkey]] (''Meleagris gallopavo'') can be taken.<ref name =raptor>Ferguson-Lees, James, and David A. Christie. Raptors of the world. Houghton Mifflin Harcourt, 2001.</ref><ref>Santander, Francisco J., et al. "PREY OF HARRIS'HAWKS (PARABUTEO UNICINCTUS) DURING AUTUMN AND WINTER IN A COASTAL AREA OF CENTRAL CHILE." The Southwestern Naturalist (2011): 417-422.</ref><ref>Houcke, H.H. (1971). Predation By a White-Tailed Hawk and a Harris' Hawk on a Wild Turkey Poult ''Condor'' 4: 475.</ref> In one instance, a lone Harris's hawk successfully killed a subadult [[great blue heron]] (''Ardea herodias'').<ref>Woodward, H.D. (2003). Lone Harris' Hawk Kills Great Blue Heron. The Raptor Research Foundation 1:85–86.</ref> Reptiles such as [[lizard]]s and [[snake]]s are additionally taken as well as large [[insect]]s.<ref name = world/><ref name =raptor/>


When [[Pack hunter|hunts in groups]], Harris's hawk can take large prey effectively, such as [[desert cottontail]] (''Syvilagus auduboni''), the leading prey species in the north of Harris's hawk's range, usually weighs {{convert|800|g|lb|abbr=on}} or less.<ref>Bednarz, J. C. (1988). A comparative study of the breeding ecology of Harris's and Swainson's hawks in southeastern New Mexico. ''Condor'' 90:311–323.</ref><ref>Bednarz, J. C., J. W. Dawson, and W. H. Whaley. (1988). Harris' Hawk. Pages 71–82 in Proceedings of the southwest raptor management symposium and workshop. (Glinski, R. L., B. G. Pendleton, M. B. Moss, M. N. LeFranc, Jr., B. A. Millsap, and S. W. Hoffman, Eds.) Natl. Wildl. Fed. Washington, D.C.</ref> Even adult [[black-tailed jackrabbits]] weighing more than {{convert|2000|g|lb|abbr=on}} can be successfully taken by a pack of harris hawks.<ref name=Kenn/><ref>Coulson, Jennifer O., and Thomas D. Coulson. "Reexamining cooperative hunting in Harris's Hawk (Parabuteo unicinctus): large prey or challenging habitats?." The Auk 130.3 (2013): 548-552.</ref>
When [[Pack hunter|hunting in groups]], Harris's hawk can take large prey effectively, such as [[desert cottontail]] (''Sylvilagus auduboni''), the leading prey species in the north of Harris's hawk's range, usually weighs {{convert|800|g|lb|abbr=on}} or less.<ref>Bednarz, J. C. (1988). A comparative study of the breeding ecology of Harris's and Swainson's hawks in southeastern New Mexico. ''Condor'' 90:311–323.</ref><ref>Bednarz, J. C., J. W. Dawson, and W. H. Whaley. (1988). Harris' Hawk. Pages 71–82 in Proceedings of the southwest raptor management symposium and workshop. (Glinski, R. L., B. G. Pendleton, M. B. Moss, M. N. LeFranc, Jr., B. A. Millsap, and S. W. Hoffman, Eds.) Natl. Wildl. Fed. Washington, D.C.</ref> Even adult [[black-tailed jackrabbits]] weighing more than {{convert|2000|g|lb|abbr=on}} can be successfully taken by a pack of Harris's hawks.<ref name=Kenn/><ref>Coulson, Jennifer O., and Thomas D. Coulson. "Reexamining cooperative hunting in Harris's Hawk (Parabuteo unicinctus): large prey or challenging habitats?." The Auk 130.3 (2013): 548-552.</ref>


Undoubtedly because it pursues large prey often, this hawk has larger and stronger feet, with long talons, and a larger, more prominent hooked beak than most other raptors around its size.<ref name= Hamerstrom/> Locally, other buteonine hawks, including the [[ferruginous hawk]], the [[red-tailed hawk]], and the [[white-tailed hawk]] also hunt primarily cottontails and jackrabbits, but each is bigger, weighing about {{convert|500|g|oz|abbr=on}}, {{convert|300|g|oz|abbr=on}} and {{convert|200|g|oz|abbr=on}}, respectively, more on average than a Harris's hawk.<ref>Smith, D. G. and J. R. Murphy. (1978). Biology of the Ferruginous Hawk in central Utah. Sociobiology 3:79–98.</ref><ref>Thurow, T. L., C. M. White, R. P. Howard, and J. F. Sullivan. (1980). Raptor ecology of Raft River valley, Idaho. EG&G Idaho, Inc. Idaho Falls.</ref><ref>Smith, D. G. and J. R. Murphy. (1973). Breeding ecology of raptors in the East Great Basin Desert of Utah. Brigham Young Univ. Sci. Bull., Biol. Ser. Vol. 18:1–76.</ref><ref>Farquhar, C. C. (1986). Ecology and breeding behavior of the White-tailed Hawk on the northern coastal prairies of Texas. PhD. diss. Texas A & M Univ. College Station.</ref><ref>Dunning Jr., John B. (Editor). (1992). ''CRC Handbook of Avian Body Masses.'' CRC Press. {{ISBN|978-0-8493-4258-5}}.</ref>
Undoubtedly because it pursues large prey often, this hawk has larger and stronger feet, with long talons, and a larger, more prominent hooked beak than most other raptors around its size.<ref name= Hamerstrom/> Locally, other buteonine hawks, including the [[ferruginous hawk]], the [[red-tailed hawk]], and the [[white-tailed hawk]] also hunt primarily cottontails and jackrabbits, but each is bigger, weighing about {{convert|500|g|oz|abbr=on}}, {{convert|300|g|oz|abbr=on}} and {{convert|200|g|oz|abbr=on}}, respectively, more on average than a Harris's hawk.<ref>Smith, D. G. and J. R. Murphy. (1978). Biology of the Ferruginous Hawk in central Utah. Sociobiology 3:79–98.</ref><ref>Thurow, T. L., C. M. White, R. P. Howard, and J. F. Sullivan. (1980). Raptor ecology of Raft River valley, Idaho. EG&G Idaho, Inc. Idaho Falls.</ref><ref>Smith, D. G. and J. R. Murphy. (1973). Breeding ecology of raptors in the East Great Basin Desert of Utah. Brigham Young Univ. Sci. Bull., Biol. Ser. Vol. 18:1–76.</ref><ref>Farquhar, C. C. (1986). Ecology and breeding behavior of the White-tailed Hawk on the northern coastal prairies of Texas. PhD. diss. Texas A & M Univ. College Station.</ref><ref>Dunning Jr., John B. (Editor). (1992). ''CRC Handbook of Avian Body Masses.'' CRC Press. {{ISBN|978-0-8493-4258-5}}.</ref>


In the Southwestern United States, the most common prey species (in descending order of prevalence) are [[desert cottontail]] (''Syvilagus auduboni''), [[eastern cottontail]] (''Syvilagus floridanus''), [[black-tailed jackrabbit]] (''Lepus californicus''), [[ground squirrel]]s (''Ammopsermophilus'' spp. and ''Spermophilus'' spp.), [[Pack rat|woodrats]] (''Neotoma'' spp.), [[kangaroo rat]]s (''Dipodomys'' spp.), [[pocket gopher]]s (''Geomys'' and ''Thomomys'' spp.), [[Gambel's quail]] (''Callipepla gambelii''), [[scaled quail]] (''C. squamata''), [[northern bobwhite]] (''Colinus virginianus''), [[cactus wren]] (''Campylorhynchus brunneicapillus''), [[northern mockingbird]] (''Mimus polyglottos''), [[Sceloporus magister|desert spiny lizards]] (''Sceloporus magister''), and [[skink]]s (''Eumeces'' spp.)<ref>Mader, W. J. (1975). Biology of the Harris' hawk in southern Arizona. Living Bird 14:59–85.</ref><ref>Brannon, J. D. (1980). ''The reproductive ecology of a Texas Harris's hawk (Parabuteo unicinctus harrisi) population''. Master's Thesis. Univ. of Texas, Austin.</ref> In the tropics, Harris's hawks have adapted to taking prey of several varieties, including those like [[chicken]]s and [[European rabbit]]s introduced by man.<ref>Nutting, C. C. (1883). On a collection of birds from the Hacienda "La Palma," Gulf of Nicoya, Costa Rica, with critical notes by Robert Ridgway. Proc. U.S. Natl. Mus. 1982(5):382–409.</ref><ref>Johnson, A. W. (1965). ''The birds of Chile and adjacent regions of Argentina, Bolivia, and Peru.'' Platt Establecimientos Graficos, Buenos Aires.</ref> In Chile, the [[common degu]] (''Octodon degus'') makes up 67.5% of the prey.<ref>Jaksic, F. M., J. L. Yanez, and R. P. Schlatter. (1980). Prey of the Harris' hawk in central Chile. ''Auk'' 97:196–198.</ref>
In the Southwestern United States, the most common prey species (in descending order of prevalence) are [[desert cottontail]] (''Sylvilagus auduboni''), [[eastern cottontail]] (''Sylvilagus floridanus''), [[black-tailed jackrabbit]] (''Lepus californicus''), [[ground squirrel]]s (''Ammospermophilus'' spp. and ''Spermophilus'' spp.), [[Pack rat|woodrats]] (''Neotoma'' spp.), [[kangaroo rat]]s (''Dipodomys'' spp.), [[pocket gopher]]s (''Geomys'' and ''Thomomys'' spp.), [[Gambel's quail]] (''Callipepla gambelii''), [[scaled quail]] (''C. squamata''), [[northern bobwhite]] (''Colinus virginianus''), [[cactus wren]] (''Campylorhynchus brunneicapillus''), [[northern mockingbird]] (''Mimus polyglottos''), [[Sceloporus magister|desert spiny lizards]] (''Sceloporus magister''), and [[skink]]s (''Eumeces'' spp.)<ref>Mader, W. J. (1975). Biology of the Harris' hawk in southern Arizona. Living Bird 14:59–85.</ref><ref>Brannon, J. D. (1980). ''The reproductive ecology of a Texas Harris's hawk (Parabuteo unicinctus harrisi) population''. Master's Thesis. Univ. of Texas, Austin.</ref> In the tropics, Harris's hawks have adapted to taking prey of several varieties, including those like [[chicken]]s and [[European rabbit]]s introduced by man.<ref>Nutting, C. C. (1883). On a collection of birds from the Hacienda "La Palma," Gulf of Nicoya, Costa Rica, with critical notes by Robert Ridgway. Proc. U.S. Natl. Mus. 1982(5):382–409.</ref><ref>Johnson, A. W. (1965). ''The birds of Chile and adjacent regions of Argentina, Bolivia, and Peru.'' Platt Establecimientos Graficos, Buenos Aires.</ref> In Chile, the [[common degu]] (''Octodon degus'') makes up 67.5% of the prey.<ref>Jaksic, F. M., J. L. Yanez, and R. P. Schlatter. (1980). Prey of the Harris' hawk in central Chile. ''Auk'' 97:196–198.</ref>


===Hunting===
===Hunting===
While most raptors are solitary, only coming together for breeding and migration, Harris's hawks will [[Pack hunter|hunt in cooperative groups]] of two to six. This is believed to be an adaptation to the lack of prey in the desert climate in which they live. In one hunting technique, a small group flies ahead and scouts, then another group member flies ahead and scouts, and this continues until prey is bagged and shared. In another, all the hawks spread around the prey and one bird flushes it out.<ref>{{cite book|author=Cook, William E.|title=''Avian Desert Predators''|year=1997 | isbn=978-3-540-59262-4 }}{{page needed|date=April 2018}}</ref> Harris's hawks will often chase prey on foot, and are quite fast on the ground and their long, yellow legs are adapted for this, as most hawks don't spend as much time on the ground. Groups of Harris's hawks tend to be more successful at capturing prey than lone hawks, with groups of two to four individuals having ~10% higher success rates per extra individual.<ref>{{cite thesis |last1=Dawson |first1=James |year=1988 |title=The cooperative breeding system of the Harris' Hawk in Arizona |url=http://arizona.openrepository.com/arizona/handle/10150/276864 |type=M.S. |publisher=The University of Arizona |access-date=17 November 2017}}</ref>
While most raptors are solitary, only coming together for breeding and migration, Harris's hawks will [[Pack hunter|hunt in cooperative groups]] of two to six. This is believed to be an adaptation to the lack of prey in the desert climate in which they live. In one hunting technique, a small group flies ahead and scouts, then another group member flies ahead and scouts, and this continues until prey is bagged and shared. In another, all the hawks spread around the prey and one bird flushes it out.<ref>{{cite book|author=Cook, William E.|title=''Avian Desert Predators''|year=1997 |publisher=Springer | isbn=978-3-540-59262-4 }}{{page needed|date=April 2018}}</ref> Harris's hawks will often chase prey on foot and are quite fast on the ground; their long legs are adapted for this, whereas most other hawks do not spend as much time on the ground. Groups of Harris's hawks tend to be more successful at capturing prey than lone hawks, with groups of two to four individuals having ~10% higher success rates per extra individual.<ref>{{cite thesis |last1=Dawson |first1=James |year=1988 |title=The cooperative breeding system of the Harris' Hawk in Arizona |url=http://arizona.openrepository.com/arizona/handle/10150/276864 |type=M.S. |publisher=The University of Arizona |access-date=17 November 2017}}</ref>


==Relationship with humans==
==Relationship with humans==

Latest revision as of 21:20, 29 November 2024

Harris's hawk
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Accipitriformes
Family: Accipitridae
Genus: Parabuteo
Species:
P. unicinctus
Binomial name
Parabuteo unicinctus
(Temminck, 1824)
Subspecies

P. u. harrisi
P. u. superior
P. u. unicinctus

Distribution map of Parabuteo unicinctus
Synonyms[2]

Falco harrisii Audubon 1839
Buteo harrisii Bonaparte 1839, Audubon 1840
Craxirex harrisii Coues 1870
Falco unicinctus Temminck 1824
Morphnus unicinctus Lesson 1828
Nisus unicinctus Cuvier 1829
Buteo unicinctus Gray 1844
Hypomorphnus unicinctus Cab. & Tschudi 1844
Spizageranus unicinctus Kaup 1845
Craxirex unicinctus Bonaparte 1854
Asturina unicincta Burm. 1855
Urubitinga unicincta Sclater 1871
Buteo unicinctus var. harrisii Coues 1872
Erythrocnema unicinctus Sharpe 1874
Polyborus tænurius Tschudi 1844

Harris's hawk (Parabuteo unicinctus), formerly known as the bay-winged hawk or dusky hawk, and known in Latin America as the peuco, is a medium-large bird of prey that breeds from the southwestern United States south to Chile, central Argentina, and Brazil. This bird is sometimes reported to be at large in Western Europe, especially Britain, but it is a popular species in falconry and these records almost invariably all refer to escapes from captivity.

The name is derived from the Greek para, meaning beside, near or like, and the Latin buteo, referring to a kind of buzzard; uni meaning once; and cinctus meaning girdled, referring to the white band at the tip of the tail.[3] John James Audubon gave this bird its English name in honor of his ornithological companion, financial supporter, and friend Edward Harris.[4]

Harris's hawk is notable for its behavior of hunting cooperatively in packs consisting of tolerant groups, while other raptors often hunt alone. Harris's hawks' social nature has been attributed to their intelligence, which makes them easy to train and has made them a popular bird for use in falconry.[5]

Description

[edit]
The distinctive plumage and tail feathers clearly confirm that this is a Harris's hawk

This medium-large hawk is roughly intermediate in size between a peregrine falcon (Falco peregrinus) and a red-tailed hawk (Buteo jamaicensis). Harris's hawks range in length from 46 to 59 cm (18 to 23 in) and generally have a wingspan of about 103 to 120 cm (41 to 47 in).[6][7] These hawks have a brownish plumage, reddish shoulders, and tail feathers with a white base and white tip.[8]

They exhibit sexual dimorphism with the females being larger by about 35%. In the United States, the average weight for adult males is about 701 g (1.545 lb), with a range of 546 to 850 g (1.204 to 1.874 lb), while the adult female average is 1,029 g (2.269 lb), with a range of 766 to 1,633 g (1.689 to 3.600 lb).[9][10] They have dark brown plumage with chestnut shoulders, wing linings, and thighs,[11] white on the base and tip of the tail,[12] long, yellow legs, and a yellow cere.[13] The vocalizations of Harris's hawk are very harsh sounds.[6]

The lifespan of Harris's Hawk is 10–12 years in the wild and 20–25 years in captivity.[14]

Juvenile

[edit]
Juvenile in a falconry demonstration

The juvenile Harris's hawk is mostly streaked with buff and appears much lighter than the dark adults. When in flight, the undersides of the juveniles' wings are buff-colored with brown streaking. They can look unlike adults at first glance, but the identical chestnut plumage is an aid for identification.[13][verification needed]

Unicinctus subspecies (Bay-winged hawk), in Argentina.

There are three subspecies of Harris's hawk:

Subspecies

[edit]
Image Subspecies Description Distribution
P. u. superior P. u. superior was believed to have longer tails and wings and to be more blackish than P. u. harrisi. However, the sample size of the original study was quite small, with only five males and six females. Later research has concluded that there is not as strong a physical difference as was originally assumed.[15] Other ecological differences, such as latitudinal cline, were also brought up as arguments against the validity of the subspecies segmentation.[16] Baja California, Arizona, Sonora, and Sinaloa.
P. u. harrisi Texas, eastern Mexico, and much of Central America.[16]
P. u. unicinctus It is smaller than the North American subspecies, tail and wings are proportionally longer, and the adult's dark brown ventrum is streaked or flecked with white or whitish. [17] South America.

Taxonomy

[edit]

Robert Ridgway placed Harris' Hawk in its own new subgenus Urubitinga (Antenor) in 1873,[18] and introduced the generic name Parabuteo[19][20] in 1874. Richard Bowdler Sharpe also separated Harris' Hawk to a monotypic genus, Erythrocnema, in 1874. In his Catalogue of Birds in the British Museum, Sharpe gives an extensive synonymy, with various authors having earlier placed harrisii in three genera and unicinctus in eleven.[2]

Distribution and habitat

[edit]

Harris's hawks live in sparse woodland and semi-desert, as well as marshes (with some trees) in some parts of their range (Howell and Webb 1995), including mangrove swamps, as in parts of their South American range.[21] Harris's hawks are permanent residents and do not migrate.[16] Important perches and nest supports are provided by scattered larger trees or other features (e.g., power poles, woodland edges, standing dead trees, live trees, boulders, and saguaros).[22]

The wild Harris's hawk population is declining due to habitat loss; however, under some circumstances, they have been known to move into developed areas.[23]

Behaviour

[edit]

This species occurs in relatively stable groups. A dominance hierarchy occurs in Harris's hawks, wherein the mature female is the dominant bird, followed by the adult male and then the young of previous years. Groups typically include from two to seven birds. Not only do birds cooperate in hunting, but they also assist in the nesting process.[24] No other bird of prey is known to hunt in groups as routinely as this species.[25]

Breeding

[edit]
At one week of age

They nest in small trees, shrubby growth, or cacti. The nests are often compact, made of sticks, plant roots, and stems and are often lined with leaves, moss, bark, and plant roots. They are built mainly by the female. There are usually two to four white to blueish-white eggs sometimes with a speckling of pale brown or gray. The nestlings start light buff, but in five to six days turn a rich brown.[26]

Very often, there will be three hawks attending one nest: two males and one female.[27] Whether or not this is polyandry is debated, as it may be confused with backstanding (one bird standing on another's back).[28] The female does most of the incubation. The eggs hatch in 31 to 36 days. The young begin to explore outside the nest at 38 days, and fledge, or start to fly, at 45 to 50 days. The female sometimes breeds two or three times in a year.[26] Young may stay with their parents for up to three years, helping to raise later broods. Nests are known to be predated by coyotes (Canis latrans), golden eagles (Aquila chrysaetos), red-tailed hawks (Buteo jamaicensis), great horned owls (Bubo virginianus), and flocks of common ravens (Corvus corax), predators possibly too formidable to be fully displaced by Harris's hawk's cooperative nest defenses. No accounts show predation on adults in the United States and Harris's hawk may be considered an apex predator, although presumably predators like eagles and great horned owls would be capable of killing them.[29] In Chile, black-chested buzzard-eagles (Geranoaetus melanoleucus) are likely predators.[30]

Feeding

[edit]
Eating a chick's leg

The majority of Harris's hawks' prey are mammals, including ground squirrels, rabbits, and larger black-tailed jackrabbits (Lepus californicus).[31] Birds from the size of small passerines such as diuca finch (Diuca diuca) to adult great egret (Ardea alba) and half-grown wild turkey (Meleagris gallopavo) can be taken.[32][33][34] In one instance, a lone Harris's hawk successfully killed a subadult great blue heron (Ardea herodias).[35] Reptiles such as lizards and snakes are additionally taken as well as large insects.[31][32]

When hunting in groups, Harris's hawk can take large prey effectively, such as desert cottontail (Sylvilagus auduboni), the leading prey species in the north of Harris's hawk's range, usually weighs 800 g (1.8 lb) or less.[36][37] Even adult black-tailed jackrabbits weighing more than 2,000 g (4.4 lb) can be successfully taken by a pack of Harris's hawks.[27][38]

Undoubtedly because it pursues large prey often, this hawk has larger and stronger feet, with long talons, and a larger, more prominent hooked beak than most other raptors around its size.[10] Locally, other buteonine hawks, including the ferruginous hawk, the red-tailed hawk, and the white-tailed hawk also hunt primarily cottontails and jackrabbits, but each is bigger, weighing about 500 g (18 oz), 300 g (11 oz) and 200 g (7.1 oz), respectively, more on average than a Harris's hawk.[39][40][41][42][43]

In the Southwestern United States, the most common prey species (in descending order of prevalence) are desert cottontail (Sylvilagus auduboni), eastern cottontail (Sylvilagus floridanus), black-tailed jackrabbit (Lepus californicus), ground squirrels (Ammospermophilus spp. and Spermophilus spp.), woodrats (Neotoma spp.), kangaroo rats (Dipodomys spp.), pocket gophers (Geomys and Thomomys spp.), Gambel's quail (Callipepla gambelii), scaled quail (C. squamata), northern bobwhite (Colinus virginianus), cactus wren (Campylorhynchus brunneicapillus), northern mockingbird (Mimus polyglottos), desert spiny lizards (Sceloporus magister), and skinks (Eumeces spp.)[44][45] In the tropics, Harris's hawks have adapted to taking prey of several varieties, including those like chickens and European rabbits introduced by man.[46][47] In Chile, the common degu (Octodon degus) makes up 67.5% of the prey.[48]

Hunting

[edit]

While most raptors are solitary, only coming together for breeding and migration, Harris's hawks will hunt in cooperative groups of two to six. This is believed to be an adaptation to the lack of prey in the desert climate in which they live. In one hunting technique, a small group flies ahead and scouts, then another group member flies ahead and scouts, and this continues until prey is bagged and shared. In another, all the hawks spread around the prey and one bird flushes it out.[49] Harris's hawks will often chase prey on foot and are quite fast on the ground; their long legs are adapted for this, whereas most other hawks do not spend as much time on the ground. Groups of Harris's hawks tend to be more successful at capturing prey than lone hawks, with groups of two to four individuals having ~10% higher success rates per extra individual.[50]

Relationship with humans

[edit]

Falconry

[edit]

Since about 1980, Harris's hawks have been increasingly used in falconry and are now the most popular hawks in the West (outside of Asia) for that purpose, as they are one of the easiest to train and the most social.[51]

Trained Harris's hawks have been used to remove an unwanted pigeon population from London's Trafalgar Square,[citation needed] and from the tennis courts at Wimbledon.[52]

Trained Harris's hawks have been used for bird abatement by falconry experts in Canada and the United States at various locations including airports, resorts, landfill sites, and industrial sites.[53]

In art

[edit]

John James Audubon illustrated Harris's hawk in The Birds of America (published in London, 1827–38) as Plate 392 with the title "Louisiana Hawk -Buteo harrisi". The image was engraved and colored by the Robert Havell, London workshops in 1837. The original watercolor by Audubon was purchased by the New York History Society where it remains to this day (January 2009).[54]

References

[edit]
  1. ^ BirdLife International (2016). "Parabuteo unicinctus". IUCN Red List of Threatened Species. 2016: e.T22695838A93529685. doi:10.2305/IUCN.UK.2016-3.RLTS.T22695838A93529685.en. Retrieved 19 November 2021.
  2. ^ a b Sharpe, R. Bowdler (1874). "Genus 20. Erythrocnema". Catalogue of the Birds in the British Museum: Volume 1, Catalogue of the Accipitres or diurnal birds of prey in the collection of the British Museum. London. pp. 84–86.{{cite book}}: CS1 maint: location missing publisher (link)
  3. ^ Jobling, James A. (1991). A Dictionary of Scientific Bird Names. Oxford University Press. ISBN 978-0-19-854634-4.
  4. ^ National Audubon Society. "Audubon". Audubon. Archived from the original on 2008-04-09. Retrieved 2013-04-29.
  5. ^ Beebe, Frank (1984). A Falconry Manual. Hancock House Publishers, ISBN 0-88839-978-2, page 81.
  6. ^ a b Udvardy, Miklos D. F. (2001). National Audubon Society Field Guide to North American Birds 'Western Region'. Knopf Doubleday Publishing. ISBN 978-0-679-42851-0.
  7. ^ Clark, W. S. and B. K. Wheeler (1987). A Field Guide to Hawks of North America. Houghton Mifflin Co. Boston.
  8. ^ Dwyer, James F.; Bednarz, James C. (2015). "Harris's Hawk - Appearance". Birds of the World. doi:10.2173/bow.hrshaw.01. S2CID 216204185. Retrieved October 22, 2018.
  9. ^ Dunning, John B. Jr. (1993). CRC Handbook of Avian Masses. CRC Press. ISBN 978-0-8493-4258-5.
  10. ^ a b Hamerstrom, F. (1978). "External sex characters of Harris' Hawks in winter". Raptor Res. 12:1–14.
  11. ^ National Geographic Society (1983). Birds of North America. National Geographic Society. ISBN 978-0-87044-472-2.
  12. ^ Sibley, David Allen (2000). National Audubon Society: The Sibley Guide to Birds. Knopf. ISBN 978-0-679-45122-8.
  13. ^ a b Rappole, John H. (2000). Birds of the Southwest. Texas A&M University Press. ISBN 978-0-89096-958-8.
  14. ^ "Harris hawk". Oregon Zoo. Archived from the original on 21 June 2022. Retrieved 22 November 2021.
  15. ^ Bednarz, J. C. (1988). "Harris' hawk subspecies: is superior larger or different than harrisi?". in Proceedings of the southwest raptor management symposium and workshop. Washington, D.C. pp. 294–300.
  16. ^ a b c Bednarz, James C. (1995). "Harris's Hawk (Parabuteo unicinctus)".
  17. ^ Blake, Ferguson-Lees and Christie cited in Birds of the world "Parabuteo unicinctrus" https://birdsoftheworld.org/bow/species/hrshaw/cur/systematics#subsp
  18. ^ Ridgway, Robert (1873). "Catalogue of the ornithological collection of the Boston Society of Natural History. Part II. Falconidae". Proceedings of the Boston Society of Natural History (Proc. Boston Soc. Nat. Hist.). 16: 43–72. Intermediate between Buteo and Urubitinga, … Type Falco unicincta Temm. Antenor Ridgeway Nov. subgen. is on page 63.
  19. ^ Baird, Spencer Fullerton; Brewer, T. M.; Ridgway, Robert (1874). A history of North American birds. Land birds. Vol. 3. Little, Brown. pp. 248–252. … a new generic name should be instituted for the present species, since it so well merits separation to that rank. On page 248, the heading is "Genus Antenor Ridgway", but the subspecies heading is "Parabuteo unicinctus var. harisii (Ridgway)".
  20. ^ "IRMNG - Parabuteo Baird, Brewer & Ridgway, 1874". www.irmng.org. Retrieved 30 October 2023.
  21. ^ (in Spanish) Olmos Fábio & Robson Silva e Silva (2003). Guará-Ambiente, Flora e Fauna dos Manguezais de Santos-Cubatão Empresa das Artes, ISBN 85-89138-06-2
  22. ^ Bednarz, J. C. and J. D. Ligon. (1988). A study of the ecological bases of cooperative breeding in the Harris' hawk. Ecology 69:1176–1187.
  23. ^ Discoll, James T. "Harris' Hawk". Archived from the original on 2007-11-23. Retrieved 2007-11-19.
  24. ^ Dawson, J. W. and R. W. Mannan. (1991). The role of territoriality in the social organization of Harris' hawks. Auk 108:661–672.
  25. ^ Griffin, C. R. (1976). A preliminary comparison of Texas and Arizona Harris' Hawks (Parabuteo unicinctus) populations. Raptor Res. 10:50–54.
  26. ^ a b Baicich, Paul J.; Harrison, Colin J. O. (1997). Nests, Eggs, and Nestlings of North American Birds. Princeton University Press. ISBN 978-0-691-12295-3.
  27. ^ a b Kaufmann, Kenn (1996). Lives of North American Birds. Houghton Mifflin Company. ISBN 978-0-395-77017-7.
  28. ^ Ligon, J. David (1999). The Evolution of Avian Breeding Systems. Oxford Ornithology Series. Vol. 10. Oxford University Press. ISBN 978-0198549130. Retrieved 4 August 2013.
  29. ^ Dawson, J. W. and R. W. Mannan. (1991). Dominance hierarchies and helper contributions in Harris' Hawks. Auk 108:649–660.
  30. ^ Jiménez, J. E., & Jaksić, F. M. (1989). Behavioral ecology of grey eagle-buzzards, Geranoaetus melanoleucus, in central Chile. Condor 913–921.
  31. ^ a b Del Hoyo, Josep, et al. Handbook of the birds of the world. Vol. 1. No. 8. Lynx Ed., 1992.
  32. ^ a b Ferguson-Lees, James, and David A. Christie. Raptors of the world. Houghton Mifflin Harcourt, 2001.
  33. ^ Santander, Francisco J., et al. "PREY OF HARRIS'HAWKS (PARABUTEO UNICINCTUS) DURING AUTUMN AND WINTER IN A COASTAL AREA OF CENTRAL CHILE." The Southwestern Naturalist (2011): 417-422.
  34. ^ Houcke, H.H. (1971). Predation By a White-Tailed Hawk and a Harris' Hawk on a Wild Turkey Poult Condor 4: 475.
  35. ^ Woodward, H.D. (2003). Lone Harris' Hawk Kills Great Blue Heron. The Raptor Research Foundation 1:85–86.
  36. ^ Bednarz, J. C. (1988). A comparative study of the breeding ecology of Harris's and Swainson's hawks in southeastern New Mexico. Condor 90:311–323.
  37. ^ Bednarz, J. C., J. W. Dawson, and W. H. Whaley. (1988). Harris' Hawk. Pages 71–82 in Proceedings of the southwest raptor management symposium and workshop. (Glinski, R. L., B. G. Pendleton, M. B. Moss, M. N. LeFranc, Jr., B. A. Millsap, and S. W. Hoffman, Eds.) Natl. Wildl. Fed. Washington, D.C.
  38. ^ Coulson, Jennifer O., and Thomas D. Coulson. "Reexamining cooperative hunting in Harris's Hawk (Parabuteo unicinctus): large prey or challenging habitats?." The Auk 130.3 (2013): 548-552.
  39. ^ Smith, D. G. and J. R. Murphy. (1978). Biology of the Ferruginous Hawk in central Utah. Sociobiology 3:79–98.
  40. ^ Thurow, T. L., C. M. White, R. P. Howard, and J. F. Sullivan. (1980). Raptor ecology of Raft River valley, Idaho. EG&G Idaho, Inc. Idaho Falls.
  41. ^ Smith, D. G. and J. R. Murphy. (1973). Breeding ecology of raptors in the East Great Basin Desert of Utah. Brigham Young Univ. Sci. Bull., Biol. Ser. Vol. 18:1–76.
  42. ^ Farquhar, C. C. (1986). Ecology and breeding behavior of the White-tailed Hawk on the northern coastal prairies of Texas. PhD. diss. Texas A & M Univ. College Station.
  43. ^ Dunning Jr., John B. (Editor). (1992). CRC Handbook of Avian Body Masses. CRC Press. ISBN 978-0-8493-4258-5.
  44. ^ Mader, W. J. (1975). Biology of the Harris' hawk in southern Arizona. Living Bird 14:59–85.
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  46. ^ Nutting, C. C. (1883). On a collection of birds from the Hacienda "La Palma," Gulf of Nicoya, Costa Rica, with critical notes by Robert Ridgway. Proc. U.S. Natl. Mus. 1982(5):382–409.
  47. ^ Johnson, A. W. (1965). The birds of Chile and adjacent regions of Argentina, Bolivia, and Peru. Platt Establecimientos Graficos, Buenos Aires.
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  49. ^ Cook, William E. (1997). Avian Desert Predators. Springer. ISBN 978-3-540-59262-4.[page needed]
  50. ^ Dawson, James (1988). The cooperative breeding system of the Harris' Hawk in Arizona (M.S.). The University of Arizona. Retrieved 17 November 2017.
  51. ^ "Raptors page". Users.cybercity.dk. Archived from the original on 2013-03-05. Retrieved 2013-04-29.
  52. ^ Mount, Harry (2017-07-03). "Introducing Rufus the hawk: the official bird scarer of the Wimbledon Championships". Daily Telegraph. Retrieved 2018-10-22.
  53. ^ Mraz, Tiffany. "BLOG: Phoenician resort hires hawks to keep pesky birds away from diners". AZFamily. Retrieved 2020-08-19.
  54. ^ "Audubon's Watercolors Octavo Pl. 392, Harris' Hawk". Joel Oppenheimer, Inc. Retrieved 2019-10-29.
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Historical material

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  • John James Audubon. "Louisiana Hawk", Ornithological Biography volume 5 (1839). Illustration from Birds of America octavo edition, 1840.