Beaver dam: Difference between revisions
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{{about}} |
{{about}} |
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{{Short description|Dam constructed by beavers}} |
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[[File:AlgonquinBeaverDam.JPG|thumb|Beaver dam at [[Algonquin Park]] in [[Ontario|Ontario, Canada]]]] |
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[[File:American Beaver.jpg|thumb|[[North American beaver]] (''Castor canadensis''), one of two species of beaver that build dams]] |
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⚫ | [[File:AlgonquinBeaverDam.JPG|thumb|Beaver dam at [[Algonquin Park]] in [[Ontario|Ontario, Canada]]]]A '''beaver dam''' or '''beaver impoundment''' is a [[dam]] built by [[beaver]]s; it creates a [[pond]] which protects against predators such as coyotes, wolves and bears, and holds their food during winter. These structures modify the natural environment in such a way that the overall ecosystem builds upon the change, making beavers a [[keystone species]] and [[ecosystem engineer]]s. They build prolifically at night, carrying mud with their forepaws and timber between their teeth. |
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==Construction== |
==Construction== |
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[[File:American Beaver.jpg|thumb|[[North American beaver]] (''Castor canadensis''), one of two species of beaver|left]]A minimum water level of {{convert|0.6|to|0.9|m|ft}} is required to keep the underwater entrance to [[beaver lodge]]s from being blocked by ice during the winter.{{Citation needed|date=May 2019}} In lakes, rivers and large streams with deep enough water, beavers may not build dams, and live in bank burrows and lodges.<ref name=":0">{{Cite journal|last1=Baker|first1=B.W.|last2=Hill|first2=E.P.|date=2003|editor-last=Feldhammer|editor2-last=Thompson|editor3-last=Chapman|title=Beaver (Castor canadensis)|url=https://www.aphis.usda.gov/wildlife_damage/beaver_damage/downloads/Baker%20and%20Hill%20Beaver%20Chapter.pdf|journal=Wild Mammals of North America: Biology, Management, and Conservation|edition=Second|location=Baltimore, Maryland|publisher=The Johns Hopkins University Press|pages=288–310|via=USDA APHIS|access-date=2020-03-08|archive-date=2020-01-23|archive-url=https://web.archive.org/web/20200123132507/https://www.aphis.usda.gov/wildlife_damage/beaver_damage/downloads/Baker%20and%20Hill%20Beaver%20Chapter.pdf|url-status=dead}}</ref> |
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[[File:BeaverDam 8409.jpg|thumb|Beaver dam in [[Lassen Volcanic National Park]]]] |
[[File:BeaverDam 8409.jpg|thumb|Beaver dam in [[Lassen Volcanic National Park]]]] |
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⚫ | Beavers start construction by diverting the stream to lessen the water's flow pressure. Branches and logs are then driven into the mud of the stream bed to form a base.<ref name=":0" /> Then sticks, bark (from [[deciduous]] trees), rocks, mud, grass, leaves, masses of plants, and anything else available are used to build the superstructure.<!-- The average height of a dam is about {{convert|1.8|m|ft}} with an average depth of water behind the dam of {{convert|1.2|to|1.8|metre}}{{Citation needed|date=May 2019}}. The thickness of the dam is often around {{convert|1|m}} or more. The length depends on the stream width, but averages about {{convert|4.5|m}} long{{Citation needed|date=May 2019}}. |
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A minimum water level of {{convert|0.6|to|0.9|m|ft}} is required to keep the underwater entrance to beaver lodges from being blocked by ice during the winter. In lakes, rivers and large streams with deep enough water, beavers may not build dams and instead live in bank burrows and lodges. If the water is not deep enough to keep beavers safe from predators and their lodge entrances ice-free, beavers build dams. |
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⚫ | Beavers vary the type of dam built and how they build it, according to the speed of water on the stream. In slow-moving water, they build a straight dam, whereas in fast-moving water they tend to be curved{{Citation needed|date=May 2019}}. Spillways and passageways are built into the dam to allow excess water to drain off without damaging it{{Citation needed|date=May 2019}}. Dams are generally built wider at the base and the top is usually tilted upstream to resist the force of the current{{Citation needed|date=May 2019}}. << Please supply sources. --> Beavers can transport their own weight in material;{{Citation needed|date=May 2019}} they drag logs along mudslides and float them through canals to get them in place.<ref>{{Cite web|url=http://neurosciencelibrary.org/specimens/rodentia/beaver/index.html|title=Comparative Mammalian Brain Collections: North American beaver (Castor canadensis)|website=neurosciencelibrary.org|access-date=2020-03-08|archive-date=2016-10-05|archive-url=https://web.archive.org/web/20161005020635/http://neurosciencelibrary.org/Specimens/rodentia/beaver/index.html|url-status=dead}}</ref> Once the dam has flooded enough area to the proper depth to form a protective moat for the lodge (often covering many acres), beavers begin construction of the lodge.<ref name="beaver facts">{{cite web|url=http://fohn.net/beaver-pictures-facts/beaver-dams.html|title=Beaver pictures & facts|author=Fall, S.|year=2007|access-date=September 2, 2013|url-status=live|archive-url=https://web.archive.org/web/20130811083239/http://fohn.net/beaver-pictures-facts/beaver-dams.html|archive-date=August 11, 2013}}</ref> |
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⚫ | Beavers start construction by diverting the stream to lessen the water's flow pressure. Branches and logs are then driven into the mud of the stream bed to form a base. Then sticks, bark (from [[deciduous]] trees), rocks, mud, grass, leaves, masses of plants, and anything else available |
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⚫ | Trees approaching a diameter of {{convert|90|cm|ft}} may be used to construct a dam, although the average is {{convert|10|to|30|cm}}. Log length depends on the diameter of the tree and the size of the beaver. There are recorded cases of beavers felling trees of {{convert|45|m}} tall and {{convert|115|cm}} in diameter. Logs of this size are not intended to be used as structural members of the dam; rather, the bark is used for food, and sometimes to get to upper branches. It takes a beaver about 20 minutes to cut down a {{convert|15|cm|adj=on}} wide aspen, by gnawing a groove around the trunk in an hourglass shape. A beaver's jaws are powerful enough to cut a {{convert|1.5|cm|adj=on}} sapling in one bite.<ref name="beaver facts" /> |
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⚫ | Beavers vary the type of dam built and how they build it, according to the speed of water on the stream. In slow-moving water, they build a straight dam, whereas in fast-moving water they tend to be curved |
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If beavers are considered [[Central place foraging|central place foragers]], their canals may be considered an extension of their "central place" far beyond the lodge, according to a 2004–2012 study that mapped beaver ponds and cut stumps.<ref>{{Cite journal|last1=Abbott|first1=Matthew|last2=Fultz|first2=Brandon|last3=Wilson|first3=Jon|last4=Nicholson|first4=Jody|last5=Black|first5=Matt|last6=Thomas|first6=Adam|last7=Kot|first7=Amanda|last8=Burrows|first8=Mallory|last9=Schaffer|first9=Benton|last10=Benson|first10=David|date=2012-01-01|title=Beaver-Dredged Canals and their Spacial Relationship to Beaver-Cut Stumps|url=https://mushare.marian.edu/fp_db/4|journal=Proceedings of the Indiana Academy of Science|volume=121|issue=2|access-date=2020-03-08|archive-date=2020-05-30|archive-url=https://web.archive.org/web/20200530012230/https://mushare.marian.edu/fp_db/4/|url-status=dead}}</ref> |
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⚫ | Trees approaching a diameter of {{convert|90|cm|ft}} may be used to construct a dam, although the average is 10 |
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Some people consider{{like whom|date=May 2021}} that by building dams beavers are expressing tool use behaviour.<ref>{{cite web|url=http://www.mnn.com/earth-matters/wilderness-resources/photos/15-remarkable-animals-that-use-tools/beavers|title=15 remarkable animals that use tools|publisher=Mother Nature Network|access-date=September 3, 2013|url-status=live|archive-url=https://web.archive.org/web/20130911191732/http://www.mnn.com/earth-matters/wilderness-resources/photos/15-remarkable-animals-that-use-tools/beavers|archive-date=September 11, 2013}}</ref> |
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==={{anchor|WoodBuffaloNP}}Extreme size=== |
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Beaver dams typically range in length from a few meters to about {{convert|100|m|ft}}. The largest beaver dam known to exist is in [[Wood Buffalo National Park]] in [[Alberta]], [[Canada]] and measures {{convert|2790|ft|m|}} in length.<ref name="strege">{{cite web | last = Strege | first = David | title = Explorer first to reach world’s largest beaver dam | url= http://www.grindtv.com/outdoor/excursions/post/explorer-first-reach-worlds-largest-beaver-dam/ | accessdate = 24 September 2014}}</ref><ref>[http://news.discovery.com/animals/zoo-animals/beaver-dam-canada-space.htm http://news.discovery.com/animals/zoo-animals/beaver-dam-canada-space.htm]</ref> Satellite photos provided by [[NASA World Wind]] show the dam did not exist in 1975 but it appeared in subsequent images. It has two or more lodges and is a combination of two original dams. Google Earth images show new dams being built which could ultimately join the main dam and increase the overall length by another 50 to 100 m during the next decade.<ref>{{cite web|url=http://www.geostrategis.com/p_beavers-longestdam.htm|title=Exploring beaver habitat and distribution with Google Earth: The longest beaver dam in the World|author=Thie, J.|accessdate=September 2, 2013}}</ref> Coordinates: 58°16'15"N 112°15'6"W |
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===Size=== |
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Another large beaver dam measuring {{convert|2139|ft|m|-1}} long,<ref name="strege"/> {{convert|14|ft|m}} high and {{convert|23|ft|m}} thick at the base was found in [[Three Forks, Montana]].<ref name="beaver facts" /> |
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Beaver dams typically range in length from a few meters to about {{convert|100|m|ft}}.<ref>{{Cite web|url=https://phys.org/news/2010-05-world-biggest-beaver-northern-canada.html|title=World's biggest beaver dam discovered in northern Canada|website=phys.org|language=en-us|access-date=2020-03-08}}</ref> Canals can be over {{convert|0.5|km|ft|abbr=}} long.<ref>{{Cite journal|last1=Grudzinski|first1=Bartosz P.|last2=Cummins|first2=Hays|last3=Vang|first3=Teng Keng|date=2019-09-15|title=Beaver canals and their environmental effects|journal=Progress in Physical Geography: Earth and Environment|volume=44|issue=2|language=en|pages=189–211|doi=10.1177/0309133319873116|s2cid=204257682|issn=0309-1333}}</ref> The largest known beaver dam is in [[Wood Buffalo National Park]] in [[Alberta]], Canada, and is {{convert|775|m}} long.<ref name="parks-canada-ca">{{ cite web | title = World's Largest Beaver Dam | url = https://parks.canada.ca/pn-np/nt/woodbuffalo/nature/beaver_gallery | access-date = 8 October 2023 | url-status = live | archive-url = https://web.archive.org/web/20230505094501/https://parks.canada.ca/pn-np/nt/woodbuffalo/nature/beaver_gallery | archive-date = 5 May 2023 }}</ref> Satellite photos provided by [[NASA WorldWind]] show the dam did not exist in 1975, but it appeared in subsequent images. It has two or more lodges and is a combination of two original dams. Google Earth images show new dams being built which could ultimately join the main dam and increase the overall length by another {{convert|50|to|100|metre}} during the next decade.<ref>{{cite web|url=http://www.geostrategis.com/p_beavers-longestdam.htm|title=Exploring beaver habitat and distribution with Google Earth: The longest beaver dam in the World|author=Thie, J.|access-date=September 2, 2013|url-status=live|archive-url=https://web.archive.org/web/20130916180026/http://geostrategis.com/p_beavers-longestdam.htm|archive-date=September 16, 2013}}</ref> Coordinates: {{coord|58|16|15|N|112|15|6|W|region:CA|display=inline}}. |
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Another large beaver dam measuring {{convert|2139|ft|m|-1}} long,<ref name="strege">{{cite web | last = Strege | first = David | title = Explorer first to reach world's largest beaver dam | url = http://www.grindtv.com/outdoor/excursions/post/explorer-first-reach-worlds-largest-beaver-dam/ | access-date = 24 September 2014 | url-status = dead | archive-url = https://web.archive.org/web/20140924141056/http://www.grindtv.com/outdoor/excursions/post/explorer-first-reach-worlds-largest-beaver-dam/ | archive-date = 24 September 2014 }}</ref> {{convert|14|ft|m}} high and {{convert|23|ft|m}} thick at the base was found in [[Three Forks, Montana]].<ref name="beaver facts" /> |
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===As tool use=== |
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It is claimed by some that by building dams, beavers are expressing tool use behaviour.<ref>{{cite web|url=http://www.mnn.com/earth-matters/wilderness-resources/photos/15-remarkable-animals-that-use-tools/beavers|title=15 remarkable animals that use tools|publisher=Mother Nature Network|accessdate=September 3, 2013}}</ref> However, the definition of [[tool use by animals]] is highly contentious. For example, it has been argued that like bird nests, beaver dams are too large to be picked up by the animal and therefore cannot be classified as a tool. However, the cutting down of trees, preparing them for use, transporting them to the dam and then inserting them into the structure is consistent with other definitions of tool use. |
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==Effects== |
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[[File:Beaver dam in a cold winter day.jpg|thumb |
[[File:Beaver dam in a cold winter day.jpg|thumb|Beaver dam in winter in [[Mont Mégantic]]|left]] |
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Dam building can |
Dam building can help to restore damaged wetlands. Wetland benefits include [[flood control]] downstream, biodiversity (by providing habitat for different species), and water cleansing, both by the breakdown of toxins such as pesticides and the retention of silt by beaver dams. Beaver dams reduce erosion and decrease the turbidity that can be a limiting factor for some aquatic life. The benefits may be long-term and largely unnoticed unless a catchment is monitored closely. Almost half of endangered and threatened species in North America rely upon wetlands.<ref>{{cite web|url=http://www.beaversww.org/beavers-and-wetlands/about-beavers/|title=About beavers|publisher=Beavers: Wetlands and Wildlife|access-date=September 3, 2013|url-status=live|archive-url=https://web.archive.org/web/20130813132124/http://beaversww.org/beavers-and-wetlands/about-beavers/|archive-date=August 13, 2013}}</ref> |
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In 2012, a systematic review was conducted on the impacts of beaver dams on fishes and fish habitat (biased to North America (88%)). The most frequently cited benefits of beaver dams were increased habitat heterogeneity, rearing and overwintering habitat |
In 2012, a systematic review was conducted on the impacts of beaver dams on fishes and fish habitat (biased to North America (88%)). The most frequently cited benefits of beaver dams were increased [[habitat heterogeneity]], rearing and overwintering habitat, flow refuge, and invertebrate production. Impeded fish movement because of dams, siltation of spawning habitat and low oxygen levels in ponds were the most often cited negative impacts. Benefits (184) were cited more frequently than costs (119).<ref>Kemp, P.S., Worthington, T.A., Langford, E.L., Tree, A.R.J. and Gaywood, M.J., (2012). "Qualitative and quantitative effects of reintroduced beavers on stream fish". ''Fish and Fisheries'', 13(2): 158–181 {{doi|10.1111/j.1467-2979.2011.00421.x}}</ref> |
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{{Clear}} |
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===Flood control=== |
===Flood control=== |
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{{See|Flood#Control}}[[file:The Beaver Dam.png|thumb|Beaver dam as depicted in [[Frances Fuller Victor]]'s 1887 book [[wikisource:en:Eleven years in the Rocky Mountains and a life on the frontier|Eleven years in the Rocky Mountains and a life on the frontier]].]]A beaver dam may have a [[Freeboard (nautical)|freeboard]] above the water level. When heavy rains occur, the river or lake fills up. Afterward the dam gradually releases the extra stored water, thus somewhat reducing the height of the flood wave moving down the river.<ref>{{Cite web|date=2020-02-17|title=Beavers cut flooding and pollution and boost wildlife populations|url=http://www.theguardian.com/environment/2020/feb/17/beavers-cut-flooding-and-pollution-and-boost-wildlife-populations|access-date=2021-05-31|website=The Guardian|language=en}}</ref><ref>{{Cite web|title=River Otter Beaver Trial {{!}} University of Exeter|url=https://www.exeter.ac.uk/creww/research/beavertrial/|access-date=2021-05-31|website=www.exeter.ac.uk}}</ref> |
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⚫ | The surface of any stream intersects the surrounding water table. By raising the stream level, the gradient of the surface of the water table above the beaver dam is reduced, and water near the beaver dam flows more slowly into the stream. This may also help in reducing flood waves, and increasing water flow when there is no rain. In other words, beaver dams smooth out water flow by increasing the area wetted by the stream. This allows more water to seep into the ground where its flow is slowed. This water eventually finds its way back to the stream. Rivers with beaver dams in their head waters have lower high water and higher low water levels. |
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By raising the [[water table]] in wetlands such as [[Fen|peatlands]], they can stabilize a fluctuating water table, which influences the levels of both carbon and water. In a 2017 study of beaver dam hydrology, monitored beaver dams in a Rocky Mountain peatland were found to increase [[groundwater]] storage and regional [[water balance]], which can be beneficial for preventing drought. The study also suggested potential to improve [[carbon sequestration]].<ref>{{Cite journal|last1=Karran|first1=Daniel J.|last2=Westbrook|first2=Cherie J.|last3=Bedard‐Haughn|first3=Angela|date=2018|title=Beaver-mediated water table dynamics in a Rocky Mountain fen|journal=Ecohydrology|language=en|volume=11|issue=2|pages=e1923|doi=10.1002/eco.1923|s2cid=133775598|issn=1936-0592}}</ref> |
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===Excess nutrient removal=== |
===Excess nutrient removal=== |
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[[File:Biberfrass.jpg|thumb|upright|Beavers that work on top of heavy snowfall make cuts that are high above ground]] |
[[File:Biberfrass.jpg|thumb|upright|Beavers that work on top of heavy snowfall make cuts that are high above ground]] |
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Beaver ponds can cause the removal of nutrients from the stream flow. Farming along the banks of rivers often increases the loads of phosphates, nitrates and other nutrients, which can cause [[eutrophication]] and may contaminate drinking water. Besides silt, the beaver dam collects twigs and branches from the beavers' activity |
Beaver ponds can cause the removal of nutrients from the stream flow. Farming along the banks of rivers often increases the loads of phosphates, nitrates and other nutrients, which can cause [[eutrophication]] and may contaminate drinking water. Besides silt, the beaver dam collects twigs and branches from the beavers' activity as well as leaves, notably in the [[autumn]]. The main component of this material is [[cellulose]], a [[polymer]] of [[β-glucose]] [[monomers]]. (This creates a more crystalline structure than is found in [[starch]], which is composed of [[α-glucose]] monomers. Cellulose is a type of [[polysaccharide]].) Many bacteria produce [[cellulase]] which can split off the [[glucose]] and use it for energy. Just as algae receive energy from sunlight, these bacteria derive energy from cellulose, and form the base of a very similar food chain. |
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Additionally, bacterial populations absorb nitrogen and phosphorus compounds as they pass by in the water stream and keep these and other nutrients in the beaver pond and the surrounding ecology. <ref>{{cite news |url=https://www.sciencedaily.com/releases/2018/05/180509121552.htm |title=Beavers do 'dam' good work cleaning water |work=Science Daily |publisher=University of Exeter |date=2018-05-19 |accessdate=2021-07-09 }}</ref> |
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===Pesticide and herbicide removal=== |
===Pesticide and herbicide removal=== |
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Agriculture |
Agriculture introduces [[herbicide]]s and [[pesticide]]s into streams. Some of these [[wikt:toxicant|toxicant]]s are metabolized and decomposed by the bacteria in the cellulose-rich bottom of a beaver dam. |
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===Denitrification=== |
===Denitrification=== |
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Some scientists believe that the nitrogen cascade, the production of |
Some scientists believe that the [[Human impact on the nitrogen cycle|nitrogen cascade]], the production of more fixed nitrogen than the natural cycles can turn back into nitrogen gas, may be as much of a problem to Earth's ecology as carbon dioxide production. <ref>{{cite journal |title=Global Nitrogen: Cycling out of Control |journal=Environmental Health Perspectives |year=2004 |last=Fields |first=Scott |volume=112 |issue=10 |pages=A556–A563 |doi=10.1289/ehp.112-a556 |pmid=15238298 |pmc=1247398 }}</ref> Studies have shown that beaver dams along a stream contribute to [[denitrification]] (the conversion of nitrogen compounds back into nitrogen). Bacteria in the dirt and the plant debris, which collects at the dams, turns nitrates into nitrogen gas. The gas bubbles to the surface and mixes with the atmosphere once more.<ref>{{cite journal|last1=Lazar|first1=Julia|last2=Addy|first2=Kelly|last3=Gold|first3=Arthur|last4=Groffman|first4=Peter|last5=McKinney|first5=Richard|last6=Kellogg|first6=Dorothy|title=Beaver Ponds: Resurgent Nitrogen Sinks for Rural Watersheds in the Northeastern United States|journal=Journal of Environmental Quality|date=16 September 2015|volume=44|issue=5|pages=1684–1693|doi=10.2134/jeq2014.12.0540|pmid=26436285|doi-access=free}}</ref> |
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===Salmon and trout=== |
===Salmon and trout=== |
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[[File:Beaver dam Jämtland.JPG|right|thumb|Large European beaver dam near Olden, [[Jämtland]], [[Sweden]]]] |
[[File:Beaver dam Jämtland.JPG|right|thumb|Large European beaver dam near Olden, [[Jämtland]], [[Sweden]]]] |
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[[File:Pipe through beaver dam.jpg|thumb|An experimental pipe through a beaver dam, placed to allow migratory fish to cross through the dam during their spawning season|left]] |
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Beaver dams and the associated ponds can provide nurseries for [[salmon]] and [[trout]].<ref>{{cite web|url=http://www.beaverdam.info/|title=Beaver dam information site|author=Grannes, S.G.|year=2008| |
Beaver dams and the associated ponds can provide nurseries for [[salmon]] and [[trout]].<ref>{{cite web|url=http://www.beaverdam.info/ |title=Beaver dam information site|author=Grannes, S.G.|year=2008|access-date=August 30, 2013|url-status=live|archive-url=https://web.archive.org/web/20130912211703/http://www.beaverdam.info/ |archive-date=September 12, 2013}}</ref> An early indication of this was seen following the 1818 agreement between the British government of Canada and the government of America allowing Americans access to the Columbia watershed. The [[Hudson's Bay Company]], in a fit of pique<!-- ? -->, instructed its trappers to extirpate the fur-bearing animals in the area. The beaver was the first to be made locally extinct. Salmon runs fell precipitously in the following years, even though none of the factors associated with the decline of salmon runs were extant at that time.<ref>{{cite web|url=http://www.nwcouncil.org/history/Beavers.asp |title=Beavers|website=nwcouncil.org|access-date=2 May 2018|url-status=live|archive-url= https://web.archive.org/web/20121114074511/http://www.nwcouncil.org/history/Beavers.asp |archive-date=14 November 2012}}</ref> |
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There are several reasons why beaver dams increase salmon runs. They produce ponds that are deep enough for juvenile salmon to hide from [[Wader|predatory wading birds]]. They trap [[nutrients]] in their ecology and notably the |
There are several reasons why beaver dams increase salmon runs.<ref>{{cite web |title=When Beaver met Salmon: how fish-friendly flow devices keep their relationship moving forward |url=https://wwf.ca/stories/when-beaver-met-salmon-how-fish-friendly-flow-devices-keep-their-relationship-moving-forward/ |website=WWF.CA |date=7 April 2021}}</ref><ref>{{cite journal |last1=Smith |first1=Joseph M. |last2=Mather |first2=Martha E. |title=Beaver dams maintain fish biodiversity by increasing habitat heterogeneity throughout a low-gradient stream network |journal=Freshwater Biology |date=July 2013 |volume=58 |issue=7 |pages=1523–1538 |doi=10.1111/fwb.12153}}</ref><ref>{{cite web |title=Riverscapes Restoration Design Manual |url=https://lowtechpbr.restoration.usu.edu/resources/Topics/02_Science/fish.html |website=lowtechpbr.restoration.usu.edu}}</ref> They produce ponds that are deep enough for juvenile salmon to hide from [[Wader|predatory wading birds]]. They trap [[nutrients]] in their ecology and notably the nutrient pulse represented by the migration of the adult salmon upstream. These nutrients help feed the juveniles after the [[yolk sac]] has been digested. The dams provide calm water which means that the young salmon can use energy for growth rather than for navigating currents; larger [[Juvenile fish|smolt]]s with a food reserve have a better rate of survival when they reach the sea. Finally, beaver dams keep the water clear which favours all [[Salmonidae|salmonoids]]. |
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{{Clear}} |
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[[File:Twenty-Two Creek Beaver Dam.jpg|thumb|Smaller beaver dam on a creek near [[Fife Lake, Michigan]]]] |
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===Frogs=== |
===Frogs=== |
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Beaver dams have been shown to be beneficial to frog and toad populations, likely because they provide protected areas for larvae to mature in |
Beaver dams have been shown to be beneficial to [[frog]] and [[toad]] populations, likely because they provide protected areas for larvae to mature in warmer, well-oxygenated water.<ref>{{cite web|url=http://www.wildlifeextra.com/go/news/beaver-boost.html&template=news_archive_item#cr|title=wildlifeextra.com – wildlifeextra Resources and Information.|website=www.wildlifeextra.com|access-date=2 May 2018|url-status=dead|archive-url=https://web.archive.org/web/20161220065739/http://www.wildlifeextra.com/go/news/beaver-boost.html%26template%3Dnews_archive_item#cr|archive-date=20 December 2016}}</ref> A study in Alberta, Canada, showed that "Pitfall traps on beaver ponds captured 5.7 times more newly metamorphosed wood frogs, 29 times more western toads and 24 times more boreal chorus frogs than on nearby free-flowing streams."<ref>{{Cite news |url=https://www.sciencedaily.com/releases/2007/01/070110180828.htm |title=Beavers Helping Frogs And Toads Survive |date=January 11, 2007 |work=[[Science Daily]] <!-- original source: http://www.eurekalert.org/pub_releases/2007-01/uoa-bhf011007.php --> |url-status=live |archive-url=https://web.archive.org/web/20150610142058/http://www.sciencedaily.com/releases/2007/01/070110180828.htm |archive-date=June 10, 2015 }}</ref> |
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===Birds=== |
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Beaver dams help migrating songbirds. By stimulating the growth of species of plants that are critical to populations of songbirds in decline, beaver dams help create food and habitat |
Beaver dams help migrating [[songbirds]]. By stimulating the growth of species of plants that are critical to populations of songbirds in decline, beaver dams help create food and habitat. The presence of beaver dams has been shown to be associated with an increased diversity of songbirds.<ref>{{Cite news |url=https://www.sciencedaily.com/releases/2008/10/081008151316.htm |title=Beavers: Dam Good For Songbirds |date=October 9, 2008 |work=[[Science Daily]] <!-- original source: http://www.eurekalert.org/pub_releases/2008-10/wcs-bdg100808.php --> |url-status=live |archive-url=https://web.archive.org/web/20170530154211/https://www.sciencedaily.com/releases/2008/10/081008151316.htm |archive-date=May 30, 2017 }}</ref> They can also have positive effects on local waterfowl, such as ducks, that are in need of standing water habitats.<ref>{{Cite journal|last1=Nummi|first1=Petri|last2=Suontakanen|first2=Eeva-Maria|last3=Holopainen|first3=Sari|last4=Väänänen|first4=Veli-Matti|date=2019|title=The effect of beaver facilitation on Common Teal: pairs and broods respond differently at the patch and landscape scales|journal=Ibis|language=en|volume=161|issue=2|pages=301–309|doi=10.1111/ibi.12626|issn=1474-919X|hdl=10138/302629|s2cid=90166178|hdl-access=free}}</ref> |
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⚫ | |||
⚫ | |||
Beaver dams can be disruptive; the flooding can cause extensive property damage, and, when the flooding occurs next to a railroad roadbed, it can cause [[derailment]]s by washing out the tracks. When a beaver dam [[Dam failure|bursts]], the resulting [[flash flood]] may overwhelm a [[culvert]]. |
|||
⚫ | Traditional solutions to beaver problems have been focused on the trapping and removal of all the beavers in the area. While this is sometimes necessary, it is typically a short-lived solution, as beaver populations have made a remarkable comeback in the United States (after near extirpation in the nineteenth century) and are likely to continually recolonize suitable habitat.<ref name=Pollock>{{cite journal |title=Hydrologic and geomorphic effects of beaver dams and their influence on fishes |first1=Michael M. |last1=Pollock |author2=Morgan Heim |author3=Danielle Werner |journal=American Fisheries Society Symposium |volume=37 |year=2003 |pages=213–233 |url=http://etal.usu.edu/ICRRR/Beaver/Beaver%20dam%20effects%20paper%20final.pdf |access-date=Feb 20, 2016 }}</ref> Modern solutions include relatively cost-effective and low maintenance [[flow devices]]. |
||
Introduced to an area without its natural predators, as in [[Tierra del Fuego]], beavers have flooded thousands of acres of land and are considered a plague. One notable difference in Tierra del Fuego from most of North America is that the trees in Tierra del Fuego cannot be [[coppicing|coppiced]]{{Citation needed|date=July 2008}} as can willows, poplars, aspens, and other North American trees. Thus the damage by the beavers seems more severe. The beaver's disruption is not limited to human geography; beavers can destroy nesting habitat for endangered species. |
|||
Warming temperatures in the Arctic allow [[beaver]]s to extend their habitat further north, where their [[Beaver_dam#Effects|dams impair]] boat travel, impact access to food, affect water quality, and endanger downstream fish populations.<ref name=Guardian_20220104/> Pools formed by the dams store heat, thus changing local [[hydrology]] and causing localized [[Permafrost#Thaw|thawing of permafrost]] that [[Climate change feedback|in turn]] contributes to global warming.<ref name=Guardian_20220104>{{cite news |last1=Milman |first1=Oliver |title=Dam it: beavers head north to the Arctic as tundra continues to heat up |url=https://www.theguardian.com/world/2022/jan/04/beavers-arctic-north-climate-crisis |newspaper=The Guardian |date=January 4, 2022 |archive-url=https://web.archive.org/web/20220104220623/https://www.theguardian.com/world/2022/jan/04/beavers-arctic-north-climate-crisis |archive-date=January 4, 2022 |url-status=live }}</ref> |
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==Stream life cycle== |
==Stream life cycle== |
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[[File:Remnants of beaver pond in New York's Adirondack Park.jpg|thumb|Drained beaver pond in [[Adirondack State Park]]]] |
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[[File:Broken Beaver Dam.jpg|thumb|Drained beaver pond in [[Allegany State Park]]]] |
[[File:Broken Beaver Dam.jpg|thumb|Drained beaver pond in [[Allegany State Park]]]] |
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===Wetland creation=== |
===Wetland creation=== |
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If a |
If a beaver pond becomes too shallow due to sediment accumulation, or the tree supply is depleted, beavers will abandon the site. Eventually the dam will be breached and the water will drain out. The rich thick layer of silt, branches, and dead leaves behind the old dam is an ideal habitat for some wetland species. |
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===Meadow creation=== |
===Meadow creation=== |
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As the wetland fills up with plant debris and dries out, pasture species colonize it and the wetland |
As the wetland fills up with plant debris and dries out, pasture species colonize it and the wetland may eventually become a meadow suitable for grazing in a previously forested area. This provides a valuable niche for many animals which otherwise would be excluded. Beaver dam creation also increases the plants the dams were made from (such as willows) to reproduce by [[Cutting (plant) |cutting]], encouraging the growth of adventitious roots. |
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===Riverine forest=== |
===Riverine forest=== |
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⚫ | |||
Finally the meadow will be colonized by riverine trees, typically aspens, willows and such species which are favoured by the beaver. Beavers are then likely to recolonize the area, and the cycle begins again. |
Finally the meadow will be colonized by riverine trees, typically aspens, willows and such species which are favoured by the beaver. Beavers are then likely to recolonize the area, and the cycle begins again. |
||
=== |
===Bottomland=== |
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Each time the stream life cycle repeats itself another layer of |
Each time the stream life cycle repeats itself another layer of organic soil is added to the bottom of the valley. The valley slowly fills and the flat area at the bottom widens. Research is sparse, but it seems likely that parts of the [[Upland and lowland|bottomland]] in North America was created, or at least added to, by the efforts of the generations of beavers that lived there. <ref>{{cite journal |title=Beaver-generated disturbance extends beyond active dam sites to enhance stream morphodynamics and riparian plant recruitment |journal=Scientific Reports |date=2019-05-31 |last1=Levine |first1=Rebekah |last2=Meyer |first2=Grant |volume=9 |issue=1 |page=8124 |doi=10.1038/s41598-019-44381-2 |pmid=31148552 |pmc=6544642 |bibcode=2019NatSR...9.8124L }}</ref> |
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==Analogs== |
|||
Humans sometimes build structures similar to beaver dams in streams, either to get the benefits of beaver dams in places without beavers, or to encourage beavers to settle in a particular area.<ref name="Backhouse">{{cite book |last1=Backhouse |first1=Frances |title=Beavers - Radical Rodents and Ecosystem Engineers |date=2021 |publisher=Orca Book Publishers |isbn=9781459824690 |pages=81;100–103}}</ref> These are often called "beaver dam analogs" (BDA) although other names are also used.<ref>{{cite book |last1=Goldfarb |first1=Ben |title=Eager: The Surprising, Secret Life of Beavers and Why They Matter |date=2018 |publisher=Chelsea Green Publishing |isbn=9781603587396 |edition=1}}</ref> When the goal is to attract beavers, sometimes the site is unsuitable in its present condition, such as being too eroded for beavers to build a dam in their usual way. BDA builders may use construction techniques beyond the beaver's capabilities, such as driving wooden posts into the stream bed to brace horizontal branches that would otherwise be washed away. The hope is that beavers who wander by or are [[Species reintroduction|brought in]] will choose to live there and take over construction and maintenance of the dam. |
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⚫ | |||
⚫ | |||
⚫ | |||
==See also== |
==See also== |
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* [[ |
* [[Environmental impacts of beavers]] |
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* [[Logjam]], an accumulation of wood debris on a river or stream |
* [[Logjam]], an accumulation of wood debris on a river or stream |
||
==References== |
==References== |
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{{ |
{{Reflist|30em}} |
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==External links== |
==External links== |
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{{ |
{{Commons category|Beaver dams}} |
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* [http://www.martinezbeavers.org "Worth a Dam" (beaver information and educational site)] |
* [http://www.martinezbeavers.org "Worth a Dam" (beaver information and educational site)] |
||
* {{Cite Collier's|wstitle=Beaver dam |short=x}} |
* {{Cite Collier's|wstitle=Beaver dam |short=x}} |
Latest revision as of 01:49, 14 December 2024
This article's lead section may be too short to adequately summarize the key points. (November 2021) |
A beaver dam or beaver impoundment is a dam built by beavers; it creates a pond which protects against predators such as coyotes, wolves and bears, and holds their food during winter. These structures modify the natural environment in such a way that the overall ecosystem builds upon the change, making beavers a keystone species and ecosystem engineers. They build prolifically at night, carrying mud with their forepaws and timber between their teeth.
Construction
[edit]A minimum water level of 0.6 to 0.9 metres (2.0 to 3.0 ft) is required to keep the underwater entrance to beaver lodges from being blocked by ice during the winter.[citation needed] In lakes, rivers and large streams with deep enough water, beavers may not build dams, and live in bank burrows and lodges.[1]
Beavers start construction by diverting the stream to lessen the water's flow pressure. Branches and logs are then driven into the mud of the stream bed to form a base.[1] Then sticks, bark (from deciduous trees), rocks, mud, grass, leaves, masses of plants, and anything else available are used to build the superstructure. Beavers can transport their own weight in material;[citation needed] they drag logs along mudslides and float them through canals to get them in place.[2] Once the dam has flooded enough area to the proper depth to form a protective moat for the lodge (often covering many acres), beavers begin construction of the lodge.[3]
Trees approaching a diameter of 90 centimetres (3.0 ft) may be used to construct a dam, although the average is 10 to 30 centimetres (3.9 to 11.8 in). Log length depends on the diameter of the tree and the size of the beaver. There are recorded cases of beavers felling trees of 45 metres (148 ft) tall and 115 centimetres (45 in) in diameter. Logs of this size are not intended to be used as structural members of the dam; rather, the bark is used for food, and sometimes to get to upper branches. It takes a beaver about 20 minutes to cut down a 15-centimetre (5.9 in) wide aspen, by gnawing a groove around the trunk in an hourglass shape. A beaver's jaws are powerful enough to cut a 1.5-centimetre (0.59 in) sapling in one bite.[3]
Maintenance work on the dam and lodges is often done in autumn.[citation needed]
If beavers are considered central place foragers, their canals may be considered an extension of their "central place" far beyond the lodge, according to a 2004–2012 study that mapped beaver ponds and cut stumps.[4]
Some people consider[like whom?] that by building dams beavers are expressing tool use behaviour.[5]
Size
[edit]Beaver dams typically range in length from a few meters to about 100 metres (330 ft).[6] Canals can be over 0.5 kilometres (1,600 ft) long.[7] The largest known beaver dam is in Wood Buffalo National Park in Alberta, Canada, and is 775 metres (2,543 ft) long.[8] Satellite photos provided by NASA WorldWind show the dam did not exist in 1975, but it appeared in subsequent images. It has two or more lodges and is a combination of two original dams. Google Earth images show new dams being built which could ultimately join the main dam and increase the overall length by another 50 to 100 metres (160 to 330 ft) during the next decade.[9] Coordinates: 58°16′15″N 112°15′6″W / 58.27083°N 112.25167°W.
Another large beaver dam measuring 2,139 feet (650 m) long,[10] 14 feet (4.3 m) high and 23 feet (7.0 m) thick at the base was found in Three Forks, Montana.[3]
Effects
[edit]Dam building can help to restore damaged wetlands. Wetland benefits include flood control downstream, biodiversity (by providing habitat for different species), and water cleansing, both by the breakdown of toxins such as pesticides and the retention of silt by beaver dams. Beaver dams reduce erosion and decrease the turbidity that can be a limiting factor for some aquatic life. The benefits may be long-term and largely unnoticed unless a catchment is monitored closely. Almost half of endangered and threatened species in North America rely upon wetlands.[11]
In 2012, a systematic review was conducted on the impacts of beaver dams on fishes and fish habitat (biased to North America (88%)). The most frequently cited benefits of beaver dams were increased habitat heterogeneity, rearing and overwintering habitat, flow refuge, and invertebrate production. Impeded fish movement because of dams, siltation of spawning habitat and low oxygen levels in ponds were the most often cited negative impacts. Benefits (184) were cited more frequently than costs (119).[12]
Flood control
[edit]A beaver dam may have a freeboard above the water level. When heavy rains occur, the river or lake fills up. Afterward the dam gradually releases the extra stored water, thus somewhat reducing the height of the flood wave moving down the river.[13][14]
The surface of any stream intersects the surrounding water table. By raising the stream level, the gradient of the surface of the water table above the beaver dam is reduced, and water near the beaver dam flows more slowly into the stream. This may also help in reducing flood waves, and increasing water flow when there is no rain. In other words, beaver dams smooth out water flow by increasing the area wetted by the stream. This allows more water to seep into the ground where its flow is slowed. This water eventually finds its way back to the stream. Rivers with beaver dams in their head waters have lower high water and higher low water levels.
By raising the water table in wetlands such as peatlands, they can stabilize a fluctuating water table, which influences the levels of both carbon and water. In a 2017 study of beaver dam hydrology, monitored beaver dams in a Rocky Mountain peatland were found to increase groundwater storage and regional water balance, which can be beneficial for preventing drought. The study also suggested potential to improve carbon sequestration.[15]
Excess nutrient removal
[edit]Beaver ponds can cause the removal of nutrients from the stream flow. Farming along the banks of rivers often increases the loads of phosphates, nitrates and other nutrients, which can cause eutrophication and may contaminate drinking water. Besides silt, the beaver dam collects twigs and branches from the beavers' activity as well as leaves, notably in the autumn. The main component of this material is cellulose, a polymer of β-glucose monomers. (This creates a more crystalline structure than is found in starch, which is composed of α-glucose monomers. Cellulose is a type of polysaccharide.) Many bacteria produce cellulase which can split off the glucose and use it for energy. Just as algae receive energy from sunlight, these bacteria derive energy from cellulose, and form the base of a very similar food chain.
Additionally, bacterial populations absorb nitrogen and phosphorus compounds as they pass by in the water stream and keep these and other nutrients in the beaver pond and the surrounding ecology. [16]
Pesticide and herbicide removal
[edit]Agriculture introduces herbicides and pesticides into streams. Some of these toxicants are metabolized and decomposed by the bacteria in the cellulose-rich bottom of a beaver dam.
Denitrification
[edit]Some scientists believe that the nitrogen cascade, the production of more fixed nitrogen than the natural cycles can turn back into nitrogen gas, may be as much of a problem to Earth's ecology as carbon dioxide production. [17] Studies have shown that beaver dams along a stream contribute to denitrification (the conversion of nitrogen compounds back into nitrogen). Bacteria in the dirt and the plant debris, which collects at the dams, turns nitrates into nitrogen gas. The gas bubbles to the surface and mixes with the atmosphere once more.[18]
Salmon and trout
[edit]Beaver dams and the associated ponds can provide nurseries for salmon and trout.[19] An early indication of this was seen following the 1818 agreement between the British government of Canada and the government of America allowing Americans access to the Columbia watershed. The Hudson's Bay Company, in a fit of pique, instructed its trappers to extirpate the fur-bearing animals in the area. The beaver was the first to be made locally extinct. Salmon runs fell precipitously in the following years, even though none of the factors associated with the decline of salmon runs were extant at that time.[20]
There are several reasons why beaver dams increase salmon runs.[21][22][23] They produce ponds that are deep enough for juvenile salmon to hide from predatory wading birds. They trap nutrients in their ecology and notably the nutrient pulse represented by the migration of the adult salmon upstream. These nutrients help feed the juveniles after the yolk sac has been digested. The dams provide calm water which means that the young salmon can use energy for growth rather than for navigating currents; larger smolts with a food reserve have a better rate of survival when they reach the sea. Finally, beaver dams keep the water clear which favours all salmonoids.
Frogs
[edit]Beaver dams have been shown to be beneficial to frog and toad populations, likely because they provide protected areas for larvae to mature in warmer, well-oxygenated water.[24] A study in Alberta, Canada, showed that "Pitfall traps on beaver ponds captured 5.7 times more newly metamorphosed wood frogs, 29 times more western toads and 24 times more boreal chorus frogs than on nearby free-flowing streams."[25]
Birds
[edit]Beaver dams help migrating songbirds. By stimulating the growth of species of plants that are critical to populations of songbirds in decline, beaver dams help create food and habitat. The presence of beaver dams has been shown to be associated with an increased diversity of songbirds.[26] They can also have positive effects on local waterfowl, such as ducks, that are in need of standing water habitats.[27]
Disruption
[edit]Beaver dams can be disruptive; the flooding can cause extensive property damage, and, when the flooding occurs next to a railroad roadbed, it can cause derailments by washing out the tracks. When a beaver dam bursts, the resulting flash flood may overwhelm a culvert.
Traditional solutions to beaver problems have been focused on the trapping and removal of all the beavers in the area. While this is sometimes necessary, it is typically a short-lived solution, as beaver populations have made a remarkable comeback in the United States (after near extirpation in the nineteenth century) and are likely to continually recolonize suitable habitat.[28] Modern solutions include relatively cost-effective and low maintenance flow devices.
Introduced to an area without its natural predators, as in Tierra del Fuego, beavers have flooded thousands of acres of land and are considered a plague. One notable difference in Tierra del Fuego from most of North America is that the trees in Tierra del Fuego cannot be coppiced[citation needed] as can willows, poplars, aspens, and other North American trees. Thus the damage by the beavers seems more severe. The beaver's disruption is not limited to human geography; beavers can destroy nesting habitat for endangered species.
Warming temperatures in the Arctic allow beavers to extend their habitat further north, where their dams impair boat travel, impact access to food, affect water quality, and endanger downstream fish populations.[29] Pools formed by the dams store heat, thus changing local hydrology and causing localized thawing of permafrost that in turn contributes to global warming.[29]
Stream life cycle
[edit]Wetland creation
[edit]If a beaver pond becomes too shallow due to sediment accumulation, or the tree supply is depleted, beavers will abandon the site. Eventually the dam will be breached and the water will drain out. The rich thick layer of silt, branches, and dead leaves behind the old dam is an ideal habitat for some wetland species.
Meadow creation
[edit]As the wetland fills up with plant debris and dries out, pasture species colonize it and the wetland may eventually become a meadow suitable for grazing in a previously forested area. This provides a valuable niche for many animals which otherwise would be excluded. Beaver dam creation also increases the plants the dams were made from (such as willows) to reproduce by cutting, encouraging the growth of adventitious roots.
Riverine forest
[edit]Finally the meadow will be colonized by riverine trees, typically aspens, willows and such species which are favoured by the beaver. Beavers are then likely to recolonize the area, and the cycle begins again.
Bottomland
[edit]Each time the stream life cycle repeats itself another layer of organic soil is added to the bottom of the valley. The valley slowly fills and the flat area at the bottom widens. Research is sparse, but it seems likely that parts of the bottomland in North America was created, or at least added to, by the efforts of the generations of beavers that lived there. [30]
Analogs
[edit]Humans sometimes build structures similar to beaver dams in streams, either to get the benefits of beaver dams in places without beavers, or to encourage beavers to settle in a particular area.[31] These are often called "beaver dam analogs" (BDA) although other names are also used.[32] When the goal is to attract beavers, sometimes the site is unsuitable in its present condition, such as being too eroded for beavers to build a dam in their usual way. BDA builders may use construction techniques beyond the beaver's capabilities, such as driving wooden posts into the stream bed to brace horizontal branches that would otherwise be washed away. The hope is that beavers who wander by or are brought in will choose to live there and take over construction and maintenance of the dam.
See also
[edit]- Environmental impacts of beavers
- Logjam, an accumulation of wood debris on a river or stream
References
[edit]- ^ a b Baker, B.W.; Hill, E.P. (2003). Feldhammer; Thompson; Chapman (eds.). "Beaver (Castor canadensis)" (PDF). Wild Mammals of North America: Biology, Management, and Conservation (Second ed.). Baltimore, Maryland: The Johns Hopkins University Press: 288–310. Archived from the original (PDF) on 2020-01-23. Retrieved 2020-03-08 – via USDA APHIS.
- ^ "Comparative Mammalian Brain Collections: North American beaver (Castor canadensis)". neurosciencelibrary.org. Archived from the original on 2016-10-05. Retrieved 2020-03-08.
- ^ a b c Fall, S. (2007). "Beaver pictures & facts". Archived from the original on August 11, 2013. Retrieved September 2, 2013.
- ^ Abbott, Matthew; Fultz, Brandon; Wilson, Jon; Nicholson, Jody; Black, Matt; Thomas, Adam; Kot, Amanda; Burrows, Mallory; Schaffer, Benton; Benson, David (2012-01-01). "Beaver-Dredged Canals and their Spacial Relationship to Beaver-Cut Stumps". Proceedings of the Indiana Academy of Science. 121 (2). Archived from the original on 2020-05-30. Retrieved 2020-03-08.
- ^ "15 remarkable animals that use tools". Mother Nature Network. Archived from the original on September 11, 2013. Retrieved September 3, 2013.
- ^ "World's biggest beaver dam discovered in northern Canada". phys.org. Retrieved 2020-03-08.
- ^ Grudzinski, Bartosz P.; Cummins, Hays; Vang, Teng Keng (2019-09-15). "Beaver canals and their environmental effects". Progress in Physical Geography: Earth and Environment. 44 (2): 189–211. doi:10.1177/0309133319873116. ISSN 0309-1333. S2CID 204257682.
- ^ "World's Largest Beaver Dam". Archived from the original on 5 May 2023. Retrieved 8 October 2023.
- ^ Thie, J. "Exploring beaver habitat and distribution with Google Earth: The longest beaver dam in the World". Archived from the original on September 16, 2013. Retrieved September 2, 2013.
- ^ Strege, David. "Explorer first to reach world's largest beaver dam". Archived from the original on 24 September 2014. Retrieved 24 September 2014.
- ^ "About beavers". Beavers: Wetlands and Wildlife. Archived from the original on August 13, 2013. Retrieved September 3, 2013.
- ^ Kemp, P.S., Worthington, T.A., Langford, E.L., Tree, A.R.J. and Gaywood, M.J., (2012). "Qualitative and quantitative effects of reintroduced beavers on stream fish". Fish and Fisheries, 13(2): 158–181 doi:10.1111/j.1467-2979.2011.00421.x
- ^ "Beavers cut flooding and pollution and boost wildlife populations". The Guardian. 2020-02-17. Retrieved 2021-05-31.
- ^ "River Otter Beaver Trial | University of Exeter". www.exeter.ac.uk. Retrieved 2021-05-31.
- ^ Karran, Daniel J.; Westbrook, Cherie J.; Bedard‐Haughn, Angela (2018). "Beaver-mediated water table dynamics in a Rocky Mountain fen". Ecohydrology. 11 (2): e1923. doi:10.1002/eco.1923. ISSN 1936-0592. S2CID 133775598.
- ^ "Beavers do 'dam' good work cleaning water". Science Daily. University of Exeter. 2018-05-19. Retrieved 2021-07-09.
- ^ Fields, Scott (2004). "Global Nitrogen: Cycling out of Control". Environmental Health Perspectives. 112 (10): A556–A563. doi:10.1289/ehp.112-a556. PMC 1247398. PMID 15238298.
- ^ Lazar, Julia; Addy, Kelly; Gold, Arthur; Groffman, Peter; McKinney, Richard; Kellogg, Dorothy (16 September 2015). "Beaver Ponds: Resurgent Nitrogen Sinks for Rural Watersheds in the Northeastern United States". Journal of Environmental Quality. 44 (5): 1684–1693. doi:10.2134/jeq2014.12.0540. PMID 26436285.
- ^ Grannes, S.G. (2008). "Beaver dam information site". Archived from the original on September 12, 2013. Retrieved August 30, 2013.
- ^ "Beavers". nwcouncil.org. Archived from the original on 14 November 2012. Retrieved 2 May 2018.
- ^ "When Beaver met Salmon: how fish-friendly flow devices keep their relationship moving forward". WWF.CA. 7 April 2021.
- ^ Smith, Joseph M.; Mather, Martha E. (July 2013). "Beaver dams maintain fish biodiversity by increasing habitat heterogeneity throughout a low-gradient stream network". Freshwater Biology. 58 (7): 1523–1538. doi:10.1111/fwb.12153.
- ^ "Riverscapes Restoration Design Manual". lowtechpbr.restoration.usu.edu.
- ^ "wildlifeextra.com – wildlifeextra Resources and Information". www.wildlifeextra.com. Archived from the original on 20 December 2016. Retrieved 2 May 2018.
- ^ "Beavers Helping Frogs And Toads Survive". Science Daily. January 11, 2007. Archived from the original on June 10, 2015.
- ^ "Beavers: Dam Good For Songbirds". Science Daily. October 9, 2008. Archived from the original on May 30, 2017.
- ^ Nummi, Petri; Suontakanen, Eeva-Maria; Holopainen, Sari; Väänänen, Veli-Matti (2019). "The effect of beaver facilitation on Common Teal: pairs and broods respond differently at the patch and landscape scales". Ibis. 161 (2): 301–309. doi:10.1111/ibi.12626. hdl:10138/302629. ISSN 1474-919X. S2CID 90166178.
- ^ Pollock, Michael M.; Morgan Heim; Danielle Werner (2003). "Hydrologic and geomorphic effects of beaver dams and their influence on fishes" (PDF). American Fisheries Society Symposium. 37: 213–233. Retrieved Feb 20, 2016.
- ^ a b Milman, Oliver (January 4, 2022). "Dam it: beavers head north to the Arctic as tundra continues to heat up". The Guardian. Archived from the original on January 4, 2022.
- ^ Levine, Rebekah; Meyer, Grant (2019-05-31). "Beaver-generated disturbance extends beyond active dam sites to enhance stream morphodynamics and riparian plant recruitment". Scientific Reports. 9 (1): 8124. Bibcode:2019NatSR...9.8124L. doi:10.1038/s41598-019-44381-2. PMC 6544642. PMID 31148552.
- ^ Backhouse, Frances (2021). Beavers - Radical Rodents and Ecosystem Engineers. Orca Book Publishers. pp. 81, 100–103. ISBN 9781459824690.
- ^ Goldfarb, Ben (2018). Eager: The Surprising, Secret Life of Beavers and Why They Matter (1 ed.). Chelsea Green Publishing. ISBN 9781603587396.