Horseshoe Canyon Formation: Difference between revisions
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{{Short description|Geological formation in Canada}} |
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{{Infobox rockunit |
{{Infobox rockunit |
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| name = Horseshoe Canyon Formation |
| name = Horseshoe Canyon Formation |
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| caption = Horseshoe Canyon Formation at [[Horsethief Canyon (Alberta)|Horsethief Canyon]], near [[Drumheller]]. The dark bands are coal seams. |
| caption = Horseshoe Canyon Formation at [[Horsethief Canyon (Alberta)|Horsethief Canyon]], near [[Drumheller]]. The dark bands are coal seams. |
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| type = [[Geological formation]] |
| type = [[Geological formation]] |
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| age = [[Campanian]]-[[Maastrichtian]]<br>~{{Fossil range|73.1|68}}<ref>{{Cite journal |last1=Eberth |first1=David A. |last2=Kamo |first2=Sandra L. |date=October 2020 |title=High-precision U–Pb CA–ID–TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada |url=http://dx.doi.org/10.1139/cjes-2019-0019 |journal=Canadian Journal of Earth Sciences |volume=57 |issue=10 |pages=1220–1237 |doi=10.1139/cjes-2019-0019 |bibcode=2020CaJES..57.1220E |issn=0008-4077}}</ref> |
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| age = [[Maastrichtian]]<br>~{{Fossil range|74|67}}<ref name=velox_redescription/> |
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| period = Maastrichtian |
| period = Maastrichtian |
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| prilithology = [[Sandstone]] |
| prilithology = [[Sandstone]] |
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| coordinates = {{Coord|51|25|24|N|112|53|18|W|name=Horseshoe Canyon|display=inline,title}} |
| coordinates = {{Coord|51|25|24|N|112|53|18|W|name=Horseshoe Canyon|display=inline,title}} |
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| unitof = [[Edmonton Group]] |
| unitof = [[Edmonton Group]] |
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| subunits =Strathmore Member, Drumheller Member, Horsethief Member, Morrin Member, Tolman Member, Carbon Member, Whitemud Member |
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| subunits = |
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| underlies = [[ |
| underlies = [[Battle Formation]], [[Scollard Formation]] |
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| overlies = [[Bearpaw Formation]] |
| overlies = [[Bearpaw Formation]] |
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| thickness = {{convert|227|m|ft|sp=us|abbr=on}}<ref name=lexicon>{{Cite web|url=http://cgkn1.cgkn.net/weblex/weblex_litho_detail_e.pl?00053:006702 |title=Horseshoe Canyon Formation|author=Lexicon of Canadian Geological Units|accessdate=2009-02-06}}</ref> |
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| extent = |
| extent = |
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| area = |
| area = |
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The '''Horseshoe Canyon Formation''' is a [[Stratigraphy|stratigraphic]] unit of the [[Western Canadian Sedimentary Basin|Western Canada Sedimentary Basin]] in southwestern [[Alberta]].<ref name="Map600">{{Cite web|url=http://www.ags.gov.ab.ca/publications/abstracts/MAP_600.html|title=Bedrock Geology of Alberta. Alberta Geological Survey, Map 600|author=Prior, G. J., Hathaway, B., Glombick, P.M., Pana, D.I., Banks, C.J., Hay, D.C., Schneider, C.L., Grobe, M., Elgr, R., and Weiss, J.A.|year=2013| |
The '''Horseshoe Canyon Formation''' is a [[Stratigraphy|stratigraphic]] unit of the [[Western Canadian Sedimentary Basin|Western Canada Sedimentary Basin]] in southwestern [[Alberta]].<ref name="Map600">{{Cite web|url=http://www.ags.gov.ab.ca/publications/abstracts/MAP_600.html|title=Bedrock Geology of Alberta. Alberta Geological Survey, Map 600|author=Prior, G. J., Hathaway, B., Glombick, P.M., Pana, D.I., Banks, C.J., Hay, D.C., Schneider, C.L., Grobe, M., Elgr, R., and Weiss, J.A.|year=2013|access-date=2013-08-13|url-status=dead|archive-url=https://web.archive.org/web/20130705151548/http://www.ags.gov.ab.ca/publications/abstracts/MAP_600.html|archive-date=2013-07-05}}</ref><ref name="Mossop24">{{Cite web|url=http://www.ags.gov.ab.ca/publications/wcsb_atlas/a_ch24/ch_24.html|title=The Geological Atlas of the Western Canada Sedimentary Basin, Chapter 24: Upper Cretaceous and Tertiary strata of the Western Canada Sedimentary Basin|author=Mossop, G.D. and Shetsen, I., (compilers), Canadian Society of Petroleum Geologists|year=1994|access-date=2013-08-01|url-status=dead|archive-url=https://web.archive.org/web/20130721174353/http://www.ags.gov.ab.ca/publications/wcsb_atlas/a_ch24/ch_24.html|archive-date=2013-07-21}}</ref> It takes its name from [[Horseshoe Canyon (Alberta)|Horseshoe Canyon]], an area of [[badlands]] near [[Drumheller]]. |
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The Horseshoe Canyon Formation is part of the [[Edmonton Group]] |
The Horseshoe Canyon Formation is part of the [[Edmonton Group]]. In its type section (Red Deer River Valley at [[Drumheller]]), it is ~{{Convert|250|m|ft}} thick, but further west the formation is older and thicker, exceeding {{Convert|500|m|ft}} near [[Calgary]].<ref name=":0">{{Cite journal |last1=Eberth |first1=David A. |last2=Braman |first2=Dennis R. |date=September 2012 |title=A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains |url=http://dx.doi.org/10.1139/e2012-035 |journal=Canadian Journal of Earth Sciences |volume=49 |issue=9 |pages=1053–1086 |doi=10.1139/e2012-035 |bibcode=2012CaJES..49.1053E |issn=0008-4077}}</ref> It is of Late [[Cretaceous]] age, [[Campanian]] to early [[Maastrichtian]] stage (Edmontonian [[Biochronology|Land-Mammal Age]]), and is composed of [[mudstone]], [[sandstone]], [[carbon]]aceous [[shale]]s, and [[coal]] seams. A variety of depositional environments are represented in the succession, including [[floodplain]]s, [[estuary|estuarine]] channels, and coal swamps, which have yielded a diversity of fossil material. Tidally-influenced estuarine point bar deposits are easily recognizable as Inclined Heterolithic Stratification (IHS). Brackish-water trace fossil assemblages occur within these bar deposits and demonstrate periodic incursion of marine waters into the estuaries. |
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The Horseshoe Canyon Formation crops out extensively in the area around Drumheller, as well as farther north along the [[Red Deer River]] near [[Trochu, Alberta|Trochu]] and along the [[North Saskatchewan River]] in [[Edmonton]].<ref name="Map600" /> It is overlain by the Battle |
The Horseshoe Canyon Formation crops out extensively in the area around Drumheller, as well as farther north along the [[Red Deer River]] near [[Trochu, Alberta|Trochu]] and along the [[North Saskatchewan River]] in [[Edmonton]].<ref name="Map600" /> It is overlain by the Battle and [[Scollard Formation|Scollard]] formations.<ref name=":0" /> The Drumheller Coal Zone, located in the lower part of the Horseshoe Canyon Formation, was mined for [[sub-bituminous]] coal in the Drumheller area from 1911 to 1979, and the [[Atlas Coal Mine]] in Drumheller has been preserved as a National Historic Site.<ref name="acmnhs1">{{cite web|title=Mine History |url=http://www.atlascoalmine.ab.ca/history.html |publisher=Atlas Coal Mine National Historic Site |access-date=9 June 2010 |url-status=dead |archive-url=https://web.archive.org/web/20100823212059/http://www.atlascoalmine.ab.ca/history.html |archive-date=23 August 2010 }}</ref> In more recent times, the Horseshoe Canyon Formation has become a major target for [[coalbed methane]] (CBM) production. |
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[[File:Bearpaw-Horseshoe Canyon.jpg|thumb|Contact (red arrow) between the underlying marine shales of the Bearpaw Formation and the coastal Horseshoe Canyon Formation. Coal beds (black bands) are common in the Horseshoe Canyon Formation and were formed in coastal swamps.]] |
[[File:Bearpaw-Horseshoe Canyon.jpg|thumb|upright=1.2|Contact (red arrow) between the underlying marine shales of the Bearpaw Formation and the coastal Horseshoe Canyon Formation. Coal beds (black bands) are common in the Horseshoe Canyon Formation and were formed in coastal swamps.]] |
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Dinosaurs found in the Horseshoe Canyon Formation include ''[[Albertavenator]]'', ''[[Albertosaurus]]'', ''[[Anchiceratops]]'', ''[[Anodontosaurus]]'', ''[[Arrhinoceratops]]'', ''[[Atrociraptor]]'', ''[[Epichirostenotes]]'', ''[[Edmontonia]]'', ''[[Edmontosaurus]]'', ''[[Hypacrosaurus]]'', ''[[Ornithomimus]]'', ''[[Pachyrhinosaurus]]'', ''[[Parksosaurus]]'', ''[[Saurolophus]]'', and ''[[Struthiomimus]]''. Other finds have included [[mammal]]s such as ''[[Didelphodon]] coyi'', non-dinosaur [[reptile]]s, [[amphibian]]s, [[fish]], marine and terrestrial [[invertebrate]]s and plant fossils. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers.<ref>{{cite journal | last1 = Quinney | first1 = Annie | last2 = Therrien | first2 = François | last3 = Zelenitsky | first3 = Darla K. | last4 = Eberth | first4 = David A. | year = 2013 | title = Palaeoenvironmental and palaeoclimatic reconstruction of the Upper Cretaceous (late Campanian–early Maastrichtian) Horseshoe Canyon Formation, Alberta, Canada |
Dinosaurs found in the Horseshoe Canyon Formation include ''[[Albertavenator]]'', ''[[Albertosaurus]]'', ''[[Anchiceratops]]'', ''[[Anodontosaurus]]'', ''[[Arrhinoceratops]]'', ''[[Atrociraptor]]'', ''[[Epichirostenotes]]'', ''[[Edmontonia]]'', ''[[Edmontosaurus]]'', ''[[Hypacrosaurus]]'', ''[[Ornithomimus]]'', ''[[Pachyrhinosaurus]]'', ''[[Parksosaurus]]'', ''[[Saurolophus]]'', and ''[[Struthiomimus]]''. Other finds have included [[mammal]]s such as ''[[Didelphodon]] coyi'', non-dinosaur [[reptile]]s, [[amphibian]]s, [[fish]], marine and terrestrial [[invertebrate]]s and plant fossils. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers.<ref>{{cite journal | last1 = Quinney | first1 = Annie | last2 = Therrien | first2 = François | last3 = Zelenitsky | first3 = Darla K. | last4 = Eberth | first4 = David A. | year = 2013 | title = Palaeoenvironmental and palaeoclimatic reconstruction of the Upper Cretaceous (late Campanian–early Maastrichtian) Horseshoe Canyon Formation, Alberta, Canada | journal = Palaeogeography, Palaeoclimatology, Palaeoecology | volume = 371 | pages = 26–44 | doi = 10.1016/j.palaeo.2012.12.009 | bibcode = 2013PPP...371...26Q }}</ref> |
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==Oil/gas production== |
==Oil/gas production== |
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==Biostratigraphy== |
==Biostratigraphy== |
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The timeline below follows work by David A. Eberth and Sandra L. Kamo published in 2019.<ref name=biostratigraphy>{{cite journal | url=https://www.nrcresearchpress.com/doi/abs/10.1139/cjes-2019-0019#.XuGwP-fQ_BV | title=High-precision U-Pb CA-ID-TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada | last1=Eberth | first1=David A. | last2=Kamo | first2=Sandra | journal=Canadian Journal of Earth Sciences | year=2019 | volume=57 | issue=10 | pages=1220–1237 | doi=10.1139/cjes-2019-0019| bibcode=2020CaJES..57.1220E | s2cid=210299227 }}</ref><br> |
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The timeline below follows syntheses presented by [[Victoria Arbour|Arbour]] et al. 2009, Cullen et al. 2013<ref name=cullen2013>{{cite journal | last1 = Cullen | first1 = T. M. | last2 = Ryan | first2 = M. J. | last3 = Schröder-Adams | first3 = C. | last4 = Currie | first4 = P. J. | last5 = Kobayashi | first5 = Y. | year = 2013 | title = An Ornithomimid (Dinosauria) Bonebed from the Late Cretaceous of Alberta, with Implications for the Behavior, Classification, and Stratigraphy of North American Ornithomimids | journal = PLOS ONE | volume = 8 | issue = 3| page = e58853 | doi=10.1371/journal.pone.0058853 | pmid=23554938 | pmc=3595220}}</ref> Larson et al. 2010,<ref name=larson2010>{{cite journal | last1 = Larson | first1 = D. W. | last2 = Brinkman | first2 = D. B. | last3 = Bell | first3 = P. R. | year = 2010 | title = Faunal assemblages from the upper Horseshoe Canyon Formation, an early Maastrichtian cool-climate assemblage from Alberta, with special reference to the Albertosaurus sarcophagus bonebed This article is one of a series of papers published in this Special Issue on the theme Albertosaurus | url = | journal = Canadian Journal of Earth Sciences | volume = 47 | issue = 9| pages = 1159–1181 | doi=10.1139/e10-005}}</ref> Williamson & Carr 2002, Claessens & Loewen 2015,<ref name=velox_redescription>Claessens, L. & Mark A. Loewen, M.A. (2015). A redescription of ''Ornithomimus velour'' Marsh, 1890 (Dinosauria, Theropoda). ''Journal of Vertebrate Paleontology'' (advance online publication). {{DOI|10.1080/02724634.2015.1034593}}</ref> and Funston & Currie (2016).<ref name=apatoraptor1>{{cite journal |author1=Gregory F. Funston |author2=Philip J. Currie |year=2016 |title=A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae |journal=Journal of Vertebrate Paleontology |volume=Online edition |issue= |pages=e1160910 |doi=10.1080/02724634.2016.1160910 }}</ref><br> |
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from: -68.0 till: -67.2 color:triassic text:- |
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from: -73.2 till: -68.0 color:jurassic text:Horseshoe Canyon Formation |
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from: -68.5 till: -67 color:triassic text:Unit 5 |
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from: -69.8 till: -68.5 color:jurassic text:Unit 4 |
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from: -70.2 till: -69.8 color:triassic text:Unit 3 |
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color:HER bar:NAM1 from:-72.7 till:-71.9 text:[[Pachyrhinosaurus canadensis]] |
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color:CAR bar:NAM2 from:-72.6 till:-70.9 text:[[Struthiomimus]] sp. |
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color:CAR bar:NAM3 from:-72.5 till:-69.6 text:[[Ornithomimus edmontonicus]] |
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color:ANK bar:NAM4 from:-72.2 till:-70.8 text:[[Anodontosaurus lambei]] |
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color:HER bar:NAM5 from:-72.6 till:-70.3 text:[[Anchiceratops ornatus]] |
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color:HAD bar:NAM6 from:-73.0 till:-71.5 text:[[Edmontosaurus regalis]] |
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color:CAR bar:NAM7 from:-73.0 till:-69.6 text:[[Albertosaurus sarcophagus]] |
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color:ANK bar:NAM8 from:-71.8 till:-71.5 text:[[Edmontonia longiceps]] |
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color:HER bar:NAM9 from:-72.3 till:-70.5 text:[[Arrhinoceratops brachyops]] |
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color:HAD bar:NAM10 from:-71.5 till:-69.6 text:[[Hypacrosaurus altispinus]] |
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color:HAD bar:NAM11 from:-70.9 till:-69.6 text:[[Saurolophus osborni]] |
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color:HER bar:NAM12 from:-70.8 till:-70.5 text:[[Sphaerotholus edmontonense]] |
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color:HER bar:NAM13 from:-70.9 till:-69.8 text:[[Parksosaurus warreni]] |
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color:HER bar:NAM14 from:-68.9 till:-68.6 text:[[Eotriceratops xerinsularis]] |
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color:HER bar:NAM12 from:-68 till:-67.6 text:[[Eotriceratops xerinsularis]] |
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color:HER bar:NAM17 from:-70 till:-69.8 text:[[Parksosaurus warreni]] |
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color:CAR bar:NAM14 from:-68.7 till:-68.5 text:[[Albertonykus borealis]] |
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color:CAR bar:NAM13 from:-68.7 till:-68.5 text:[[Atrociraptor marshalli]] |
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color:CAR bar:NAM19 from:-68.7 till:-68.5 text:[[Struthiomimus]] sp. |
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from: -70.9 till: -69.6 color:jurassic text:Tolman M. |
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from: -71.5 till: -70.9 color:triassic text:Morrin M. |
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from: -72.3 till: -71.5 color:jurassic text:Horsethief M. |
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==Dinosaurs== |
==Dinosaurs== |
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=== Ornithischians === |
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==== Ankylosaurs ==== |
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===Ankylosaurs=== |
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! colspan="6" align="center" |'''[[Ankylosaur]]s reported from the Horseshoe Canyon Formation''' |
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''A. lambei''<ref name=Oohkotokia/><ref name=Arbour2010/> |
''A. lambei''<ref name=Oohkotokia/><ref name=Arbour2010/> |
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Horsethief, Morrin, and lowest Tolman<ref name=Oohkotokia/><ref name=Arbour2010/> |
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|[Five] skulls, mandibles, cervical vertebra, caudal vertebrae, dorsal vertebrae, sacrocaudal vertebrae, sacrum, illium, ischium, partial pelvis, coossified ribs, femur, ?manual phalanx, tail clubs, numerous cervical half-rings and osteoderms.<ref name=Oohkotokia/><ref name=Arbour2010/> |
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An [[ankylosaurinae|ankylosaurine]] [[ankylosauridae|ankylosaurid]] also known from the middle [[Dinosaur Park Formation]] and closely related to ''[[Ankylosaurus]]''.<ref>Paul Penkalski (2018). Revised systematics of the armoured dinosaur Euoplocephalus and its allies. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 287(3): 261-306. doi: https://doi.org/10.1127/njgpa/2018/0717</ref> |
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Units 2-4 |
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|[[File:Anodontosaurus.tif|frameless]] |
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[[File:Anodontosaurus LM.png|frameless]] |
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[[File:Edmontonia_dinosaur.png|thumb|center|150px|''[[Edmontonia]]'']] |
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''[[Edmontonia]]''<ref name=Sternberg1928>{{cite journal | last1 = Sternberg | first1 = C.M. | year = 1928 | title = A new armored dinosaur from the Edmonton Formation of Alberta | journal = Transactions of the Royal Society of Canada |series=Series 3 | volume = 22 | pages = 93–106 }}</ref> |
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''[[Edmontonia]]'' |
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''E. longiceps'' |
''E. longiceps''<ref name=Sternberg1928/> |
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Unit 2 |
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Upper Horsethief<ref name=Sternberg1928/> |
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| A skull with mandible, dorsal vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, cervical ribs, dorsal ribs, humerus, radius, ulna, radius, ilia, ischia, both pubes (?), femur, tibia, fibula and osteoderms.<ref name=Sternberg1928/> |
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A [[Panoplosaurini|panoplosaurin]] [[nodosauridae|nodosaurid]] also known from the lower Dinosaur Park Formation and closely related ''[[Denversaurus]]''.<ref>{{cite journal | last1 = Russell | first1 = L.S. | year = 1940 | title = Edmontonia rugosidens (Gilmore), an armored dinosaur from the Belly River Series of Alberta | journal = University of Toronto Studies, Geology Series | volume = 43 | pages = 3–28 }}</ref> |
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|[[File:Edmontonia half.jpg|frameless]] |
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[[File:Edmontonia Scale.svg|frameless]] |
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Walter Coombs (1971) synonymised ''[[Anodontosaurus lambei]]'' with ''[[Euoplocephalus|E. tutus]]''. However, recent studies suggest that ''Anodontosaurus'' is distinct enough from ''Euoplocephalus'' to be placed in its own genus and species.<ref name=Oohkotokia>{{Cite journal | last1 = Penkalski | first1 = P. | doi = 10.4202/app.2012.0125 | title = A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA | journal = Acta Palaeontologica Polonica | year = 2013 | doi-access = free }}</ref><ref name=ScolosaurusRev>{{Cite journal | last1 = Penkalski | first1 = P. | last2 = Blows | first2 = W. T. | doi = 10.1139/cjes-2012-0098 | title = ''Scolosaurus cutleri'' (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada | journal = Canadian Journal of Earth Sciences | pages = 171–182 | year = 2013 | volume=50| issue = 2 | bibcode = 2013CaJES..50..171P }}</ref> Furthermore, all Horseshoe Canyon Formation [[ankylosaurine]] specimens were suggested to be reassigned to ''Anodontosaurus''.<ref name=Arbour2010>{{cite journal |last=Arbour |first=Victoria |year=2010 |title=A Cretaceous armoury: Multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA |journal=Journal of Vertebrate Paleontology |volume=30 |issue=Supplement 2 |pages=55A |doi=10.1080/02724634.2010.10411819}}</ref> |
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| style="background:#f3e9f3;" | |
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|[[File:Euoplocephalus TMP 1991.127.1.tif|frameless]] |
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Walter Coombs (1971) synonymised ''[[Anodontosaurus lambei]]'' with ''[[Euoplocephalus|E. tutus]]''. However, recent studies suggest that ''Anodontosaurus'' is distinct enough from ''Euoplocephalus'' to be placed in its own genus and species.<ref name=Oohkotokia>{{Cite journal | last1 = Penkalski | first1 = P. | doi = 10.4202/app.2012.0125 | title = A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA | journal = Acta Palaeontologica Polonica | year = 2013 | pmid = | pmc = }}</ref><ref name=ScolosaurusRev>{{Cite journal | last1 = Penkalski | first1 = P. | last2 = Blows | first2 = W. T. | doi = 10.1139/cjes-2012-0098 | title = ''Scolosaurus cutleri'' (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada | journal = Canadian Journal of Earth Sciences | pages = 130110052638009 | year = 2013 | pmid = | pmc = | volume=50}}</ref> Furthermore, all Horseshoe Canyon Formation [[ankylosaurine]] specimens were suggested to be reassigned to ''Anodontosaurus''.<ref name=Arbour2010>{{cite journal |last=Arbour |first=Victoria |year=2010 |title=A Cretaceous armoury: Multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA |journal=Journal of Vertebrate Paleontology |volume=30 |issue=Supplement 2 |pages=55A |doi=10.1080/02724634.2010.10411819}}</ref> |
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[[File:Euoplocephalus scale.png|frameless]] |
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=== |
==== Hadrosaurs ==== |
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{| class="wikitable" align="center" |
{| class="wikitable" align="center" width="100%" |
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|- |
|- |
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! colspan="7" align="center" |'''[[ |
! colspan="7" align="center" |'''[[Hadrosaur]]s reported from the Horseshoe Canyon Formation''' |
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! Genus |
! Genus |
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Line 212: | Line 198: | ||
! Material |
! Material |
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! Notes |
! Notes |
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! |
!Images |
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''[[Edmontosaurus]]'' |
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''[[Albertavenator]]''<ref>Evans, D.C., Cullen, T.M., Larson, D.W., and Rego, A. “A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada.” Canadian Journal of Earth Sciences Early Online: 813-826. DOI: dx.doi.org/10.1139/cjes-2017-0034</ref> |
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''[[Edmontosaurus_regalis|E. regalis]]'' |
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''A. curriei'' |
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Horsethief; likely present in Drumheller. |
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Frontals, [[type specimen]] |
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A troodontid |
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|[[File:Edmontosaurus regalis skull.jpg|frameless]] |
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''[[ |
''[[Hypacrosaurus]]'' |
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'' |
''H. altispinus'' |
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Morrin and Tolman. |
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Upper unit 4 |
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"[Five to ten] articulated skulls, some associated with postcrania, isolated skull elements, isolated postcranial elements, many individuals, embryo to adult."<ref name="table-20-1-441">"Table 20.1," in Weishampel, et al. (2004). Page 441.</ref> |
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Limb bones, [[type specimen]] |
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|[[File:Hypacrosaurus altispinus, head.jpg|frameless]] |
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An alvarezsaurid |
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| rowspan="99" | |
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[[File:Albertonykus borealis.jpg|thumb|center|150px|''[[Albertonykus]]'']] |
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[[File:Atrociraptor.jpg|thumb|center|150px|''[[Atrociraptor]]'']] |
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|- |
|- |
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''[[Saurolophus]]'' |
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''[[Apatoraptor]]'' <ref name=apatoraptor1/> |
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''S. osborni'' |
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''A. pennatus'' <ref name=apatoraptor1/> |
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Upper Morrin and Tolman. |
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Unit 1 <ref name=apatoraptor1/> |
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"Complete skull and skeleton, [two] complete skulls."<ref name="table-20-1-441"/> |
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|[[File:Saurolophus osborni.jpg|frameless]] |
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A caenagnathid.<ref name=apatoraptor1/> |
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|- |
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|} |
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==== Marginocephalians ==== |
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===== Ceratopsians ===== |
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{{paleobiota-key-compact}} |
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{| class="wikitable" align="center" width="100%" |
|||
|- |
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! colspan="7" align="center" |'''[[Ceratopsian]]s reported from the Horseshoe Canyon Formation''' |
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|- |
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! Genus |
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! Species |
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! Stratigraphic position |
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! Material |
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! Notes |
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!Images |
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|- |
|- |
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| |
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''[[Anchiceratops]]''<ref name="Brown1914">{{cite journal | last1 = Brown | first1 = B | year = 1914 | title = ''Anchiceratops'', a new genus of horned dinosaurs from the Edmonton Cretaceous of Alberta. With a discussion of the origin of the ceratopsian crest and the brain casts of ''Anchiceratops'' and ''Trachodon''". | journal = Bulletin of the American Museum of Natural History | volume = 33 | pages = 539–548 }}</ref><ref name="Mallon2012"/> |
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''[[Atrociraptor]]'' |
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| |
| |
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''A. ornatus''<ref name="Brown1914"/><ref name="Mallon2012"/> |
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''A. marshalli'' |
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| |
| |
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Horsethief, Morrin, and Tolman; may have been present in Drumheller<ref name="Mallon2012">{{cite journal | last1 = Mallon | first1 = Jordan C. | last2 = Holmes | first2 = Robert | last3 = Eberth | first3 = David A. | last4 = Ryan | first4 = Michael J. | last5 = Anderson | first5 = Jason S. | year = 2012 | title = Variation in the skull of ''Anchiceratops'' (Dinosauria, Ceratopsidae) from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta | journal = Journal of Vertebrate Paleontology | volume = 31 | issue = 5| pages = 1047–1071 | doi=10.1080/02724634.2011.601484| s2cid = 86349131 }}</ref> |
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| |
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[Two] nearly complete skulls and [seven] partial skulls.<ref name="Mallon2012"/> |
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Upper unit 4 |
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| |
| |
||
A chasmosaurine ceratopsid contemporaneous and closely related to ''[[Arrhinoceratops]]''.<ref name="Mallon2012"/> |
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Partial skull, [[type specimen]] |
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|[[File:Anchiceratops.png|frameless]] |
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|- |
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| |
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''[[Arrhinoceratops]]''<ref name="Parks1925">Parks, W.A. (1925). "''Arrhinoceratops brachyops'', a new genus and species of Ceratopsia from the Edmonton Formation of Alberta". ''University of Toronto Studies, Geology Series'' 19:1-15</ref> |
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| |
|||
''A. brachyops''<ref name="Parks1925"/> |
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| |
|||
Horsethief<ref name="Parks1925"/> |
|||
| |
|||
A nearly complete skull.<ref name="Parks1925"/> |
|||
| |
| |
||
A [[chasmosaurinae|chasmosaurine]] [[ceratopsidae|ceratopsid]] restricted to the holotype specimen as Farke (2007) assigned the referred specimen ROM 1439 to ''[[Torosaurus]]''.<ref>Farke, A.A., 2007, "Cranial osteology and phylogenetic relationships of the chasmosaurine ceratopsid ''Torosaurus latus''", pp 235-257 in: K. Carpenter (ed.), ''Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs'', Bloomington, Indiana University Press</ref> |
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A dromaeosaurid |
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|[[File:Arrhinoceratops brachyops skull, Near Drumheller, Alberta, Late Cretaceous - Royal Ontario Museum - DSC00076.JPG|frameless]] |
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|- |
|- |
||
| |
| |
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''[[Eotriceratops]]''<ref name="wuetal2007">{{cite journal |last1=Wu |first1=X-C. |last2=Brinkman |first2=D.B. |last3=Eberth |first3=D.A. |last4=Braman |first4=D.R. |year=2007 |title=A new ceratopsid dinosaur (Ornithischia) from the uppermost Horseshoe Canyon Formation (upper Maastrichtian), Alberta, Canada |journal=Canadian Journal of Earth Sciences |volume=44 |issue=9 |pages=1243–1265 |doi=10.1139/E07-011|bibcode = 2007CaJES..44.1243W }}</ref> |
|||
''[[Epichirostenotes]]'' |
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| |
| |
||
''E. xerinsularis''<ref name="wuetal2007"/> |
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''E. curriei''<ref name=caenagnathids>{{cite journal |author1=Robert M. Sullivan |author2=Steven E. Jasinski |author3=Mark P.A. Van Tomme |year=2011 |title=A new caenagnathid ''Ojoraptorsaurus boerei'', n. gen., n. sp. (Dinosauria, Oviraptorosauria), from the Upper Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico |url=http://www.robertmsullivanphd.com/uploads/169._Sullivan_et_al.__Ojoraptorsaurus__COLOR.pdf |journal=Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin |volume=53 |pages=418–428}}</ref> |
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| |
| |
||
Carbon<ref name="wuetal2007"/> |
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| |
| |
||
Premaxillae, maxillae, rostral, supraorbital horncore with lacrimal, prefrontal, frontal, postorbital, jugal, epijugal, quadratojugal, quadrate, partial parietal, squamosal frill, braincase, syncervical, cervical vertebrae, dorsal vertebrae, ribs and fragments of ossified ligaments.<ref name="wuetal2007"/> |
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Unit 1 |
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| |
| |
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A large [[chasmosaurinae|chasmosaurine]] [[ceratopsidae|ceratopsid]] that may represent a [[synonym_(taxonomy)|senior synonym]] of ''[[Ojoceratops]]'' from the [[Ojo Alamo Formation]].<ref name=Bravoceratops>{{Cite journal | last1 = Wick | first1 = S. L. | last2 = Lehman | first2 = T. M. | doi = 10.1007/s00114-013-1063-0 | title = A new ceratopsian dinosaur from the Javelina Formation (Maastrichtian) of West Texas and implications for chasmosaurine phylogeny | journal = Naturwissenschaften | volume = 100| issue = 7| pages =667–82| year = 2013 | pmid = 23728202|bibcode = 2013NW....100..667W | s2cid = 16048008 }}</ref> |
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Partial skeleton, [[type specimen]] |
|||
|[[File:Eotriceratops.jpg|frameless]] |
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|- |
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| |
| |
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''[[Montanoceratops]]''<ref name=brown1942>{{cite journal | last1 = Brown | first1 = B. | last2 = Schlaikjer | first2 = E. M. | year = 1942 | title = The skeleton of ''Leptoceratops'' with the description of a new species | journal = American Museum Novitates | issue = 1169 | pages = 1–15 }}</ref><ref name="J. Makovicky 2001"/> |
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A caenagnathid |
|||
| |
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''M. cerorhynchus''<ref name=brown1942/><ref name="J. Makovicky 2001"/> |
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| |
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Upper Tolman<ref name="J. Makovicky 2001"/> |
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| |
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An isolated braincase.<ref name="J. Makovicky 2001">P. J. Makovicky. 2001. A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta. Mesozoic Vertebrate Life, D. H. Tanke and K. Carpenter (eds.), Indiana University Press, Bloomington 243-262</ref> |
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| |
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A [[leptoceratopsidae|leptoceratopsid]] [[ceratopsia|ceratopsian]] also known from the [[St. Mary River Formation]].<ref name=brown1942/> |
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|[[File:Montanoceratops BW flipped transparent.png|frameless]] |
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|- |
|- |
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''[[Pachyrhinosaurus]]''<ref name=Sternberg1950>{{cite journal | last1 = Sternberg | first1 = C. M. | year = 1950 | title = Pachyrhinosaurus canadensis, representing a new family of the Ceratopsia, from southern Alberta | journal = National Museum of Canada Bulletin | volume = 118 | pages = 109–120 }}</ref> |
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| |
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''P. canadensis''<ref name=Sternberg1950/> |
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| |
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Drumheller and Horsethief<ref name=Sternberg1950/> |
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| |
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[Two] partial skulls.<ref name=Sternberg1950/> |
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| |
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A [[centrosaurinae|centrosaurine]] [[ceratopsidae|ceratopsid]] also known from the [[St. Mary River Formation]].<ref name=Sternberg1950/> |
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|[[File:Pachyrhinosaurus canadensis skull.jpg|frameless]] |
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|} |
|} |
||
=== |
===== Pachycephalosaurs ===== |
||
{{paleobiota-key-compact}} |
|||
{| class="wikitable" align="center" width="100%" |
{| class="wikitable" align="center" width="100%" |
||
|- |
|- |
||
! colspan="7" align="center" |'''[[ |
! colspan="7" align="center" |'''[[Pachycephalosaurid]]s reported from the Horseshoe Canyon Formation''' |
||
|- |
|- |
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! Genus |
! Genus |
||
Line 294: | Line 326: | ||
|- |
|- |
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''[[Sphaerotholus]]''<ref name=Longrich2010>{{cite journal |author1=Longrich N. R. |author2=Sankey J. T. | year = 2010 | title = ''Texacephale langstoni'', a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA | journal = Cretaceous Research | volume = 31 |issue=2 | pages = 274–284 | doi=10.1016/j.cretres.2009.12.002|bibcode=2010CrRes..31..274L |display-authors=etal}}</ref><ref name="W&C">{{cite journal |author2=Carr Thomas D. |author1=Williamson Thomas E. | year = 2002 | title = A new genus of highly derived pachycephalosaurian from western North America | doi = 10.1671/0272-4634(2002)022[0779:angodp]2.0.co;2 | journal = Journal of Vertebrate Paleontology | volume = 22 | issue = 4| pages = 779–801 |s2cid=86112901 }}</ref> |
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''[[Anchiceratops]]'' |
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| |
| |
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''S. edmontonense''<ref name="W&C"/> |
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''A. ornatus'' |
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| |
| |
||
| |
| |
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Tolman<ref name=Longrich2010/> |
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Units 1 & 2 |
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| |
| |
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A frontoparietal.<ref name=Longrich2010/><ref name="W&C"/> |
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| |
| |
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A pachycephalosaurine [[pachycephalosauridae|pachycephalosaurid]] also known from the [[Hell Creek Formation|Hell Creek]] and [[Kirtland Formation]].<ref name="W&C"/> |
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| rowspan="99" | |
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[[File: |
|[[File:Sphaerotholus.jpg|center|150px|''[[Sphaerotholus]]''|frameless]] |
||
|} |
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[[File:Arrhinoceratops BW.jpg|thumb|center|150px|''[[Arrhinoceratops]]'']] |
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[[File:Eotriceratops BW.jpg|thumb|center|150px|''[[Eotriceratops]]'']] |
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==== Thescelosaurs ==== |
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[[File:Pachyrhinosaurus BW.jpg|thumb|center|150px|''[[Pachyrhinosaurus lakustai]]'']] |
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{| class="wikitable" align="center" width="100%" |
|||
|- |
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! colspan="7" align="center" |'''[[Thescelosaurid]]s reported from the Horseshoe Canyon Formation''' |
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|- |
|||
! Genus |
|||
! Species |
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! Location |
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! Stratigraphic position |
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! Material |
|||
! Notes |
|||
! Images |
|||
|- |
|- |
||
| |
| |
||
''[[ |
''[[Parksosaurus]]'' |
||
| |
| |
||
'' |
''P. warreni'' |
||
| |
| |
||
| |
| |
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Tolman |
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Units 1 & 2 |
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| A partially articulated skeleton and partial skull. |
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| |
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| A [[Thescelosauridae|thescelosaurid]] [[neornithischian]]. |
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"Complete skull."<ref name="table-23-1-495">"Table 23.1," in Weishampel, et al. (2004). Page 495.</ref> |
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| [[File:Parksosaurus_warreni,_Near_Drumheller,_Alberta,_Canada,_Late_Cretaceous_-_Royal_Ontario_Museum_-_DSC00035.JPG|center|150px|''[[Parksosaurus]]''|frameless]] |
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|} |
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=== Theropods === |
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==== Maniraptorans ==== |
|||
{| class="wikitable" align="center" <!-- width="100%" --> |
|||
|- |
|||
! colspan="7" align="center" |'''[[Maniraptor]]s reported from the Horseshoe Canyon Formation''' |
|||
|- |
|||
! Genus |
|||
! Species |
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! Stratigraphic position |
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! Material |
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! Notes |
|||
!Images |
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|- |
|- |
||
| |
| |
||
''[[Albertavenator]]''<ref name=Evans2017>Evans, D.C., Cullen, T.M., Larson, D.W., and Rego, A. "A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada." Canadian Journal of Earth Sciences Early Online: 813-826. DOI: dx.doi.org/10.1139/cjes-2017-0034</ref> |
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''[[Eotriceratops]]'' |
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| |
| |
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''A. curriei''<ref name=Evans2017/> |
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''E. xerinsularis'' |
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| |
| |
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Horsethief<ref name=Evans2017/> |
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| |
| |
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[Two] frontals.<ref name=Evans2017/> |
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Unit 5 |
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| |
| |
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A [[troodontidae|troodontid]] [[theropoda|theropod]] that adds to the diversity of North American troodontids.<ref name=Evans2017/> |
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| |
| |
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|- |
|- |
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|style="background:#f3e9f3;" | |
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''[[Montanoceratops]]''<ref name="mcer-brain-abs-243" /> |
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|style="background:#f3e9f3;" | |
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''M. cerorhynchus''<ref name="mcer-brain-abs-243" /> |
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|style="background:#f3e9f3;" | |
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|style="background:#f3e9f3;" | |
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|style="background:#f3e9f3;" | |
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Isolated braincase AMNH 5244.<ref name="mcer-brain-abs-243" /> |
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|style="background:#f3e9f3;" | |
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AMNH 5244 was probably left by an indeterminate leptoceratopsid. |
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|- |
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|rowspan=2| |
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''[[Pachyrhinosaurus]]'' |
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| |
| |
||
''[[Albertonykus]]''<ref name=LC08>{{cite journal |last=Longrich |first=Nicholas R. |author2=Currie, Philip J. |year=2009 |title=''Albertonykus borealis'', a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae |journal=Cretaceous Research |volume=30 |issue=1 |pages= 239–252|doi=10.1016/j.cretres.2008.07.005 |bibcode=2009CrRes..30..239L }}</ref> |
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''P. canadensis'' |
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| |
| |
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''A. borealis''<ref name=LC08/> |
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| |
| |
||
Upper |
Upper Tolman<ref name=LC08/> |
||
| |
| |
||
Ulna, tibiae, metatarsals, manual ungual, pedal phalanxes and unidentified pedal phalanges.<ref name=LC08/> |
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|rowspan=2| |
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| |
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Ceratopsids |
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An mononykin [[alvarezsauridae|alvarezsaurid]] specialized for digging insect nests.<ref name=LC08/> |
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|[[File:Albertonykus borealis life reconstruction.png|frameless]] |
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|- |
|- |
||
| |
| |
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''[[Apatoraptor]]'' <ref name=apatoraptor1>{{cite journal |author1=Gregory F. Funston |author2=Philip J. Currie |year=2016 |title=A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae |journal=Journal of Vertebrate Paleontology |volume=36 |issue=4 |pages=e1160910 |doi=10.1080/02724634.2016.1160910 |bibcode=2016JVPal..36E0910F |s2cid=131090028 |url=https://figshare.com/articles/dataset/A_new_caenagnathid_Dinosauria_Oviraptorosauria_from_the_Horseshoe_Canyon_Formation_of_Alberta_Canada_and_a_reevaluation_of_the_relationships_of_Caenagnathidae/3172573 }}</ref> |
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''P. lakustai'' |
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| |
| |
||
''A. pennatus''<ref name=apatoraptor1/> |
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| |
| |
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Horsethief<ref name=apatoraptor1/> |
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Lower unit 1 |
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| |
| |
||
A palatine, mandibles, ceratobranchials, axis, cervical vertebrae, dorsal vertebrae, pectoral girdles, sternal plate, forelimb, ilium, femur, tibia and fibula.<ref name=apatoraptor1/> |
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| |
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A [[caenagnathidae|caenagnathid]] [[oviraptorosauria|oviraptorosaur]] recovered as sister taxon to ''[[Elmisaurus]]'' .<ref name=apatoraptor1/> |
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|[[File:Apatoraptor NT small.jpg|frameless]] |
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|- |
|- |
||
| |
| |
||
''[[Atrociraptor]]''<ref name="Currie2004">Currie, P. J. and D. J. Varricchio (2004). "A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada". Pp. 112–132 in P. J. Currie, E. B. Koppelhus, M. A. Shugar and J. L. Wright. (eds.), ''Feathered Dragons''. Indianapolis: Indiana University Press. [http://lesdinos.free.fr/003%202004.pdf]</ref> |
|||
?''[[Sphaerotholus]]'' |
|||
| |
| |
||
''A. marshalli''<ref name=Currie2004/> |
|||
''S. edmontonense'' |
|||
| |
| |
||
Lower Horsethief<ref name=Currie2004/> |
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| |
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Premaxillae, maxilla, dentaries, teeth and numerous bone fragments.<ref name=Currie2004/> |
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Unit 1 |
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A [[Dromaeosauridae|dromaeosaurid]]; teeth indicate it may have been present across all members.<ref name=Currie2004/> |
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|[[File:Atrociraptor royal tyrrell.jpg|frameless]] |
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?''[[Prenocephale]]''. |
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''[[Epichirostenotes]]''<ref name=Sullivan2011/> |
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"Almond Formation" ceratopsid |
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''E. curriei''<ref name=Sullivan2011>{{cite journal |author1=Robert M. Sullivan |author2=Steven E. Jasinski |author3=Mark P.A. Van Tomme |year=2011 |title=A new caenagnathid ''Ojoraptorsaurus boerei'', n. gen., n. sp. (Dinosauria, Oviraptorosauria), from the Upper Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico |url=http://www.robertmsullivanphd.com/uploads/169._Sullivan_et_al.__Ojoraptorsaurus__COLOR.pdf |journal=Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin |volume=53 |pages=418–428}}</ref> |
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Unnamed |
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Horsethief, Morrin, and Tolman<ref name=Sullivan2011/> |
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A maxilla, probable palatine, partial braincase, cervical vertebrae, dorsal vertebrae, caudal vertebrae, synsacrum, an anterior and a posterior cervical rib, a mid-dorsal rib, fragments of ilia, ischium, both pubes, partial tibia, and unidentifiable bones.<ref name=Sullivan2011/> |
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Upper Unit 1, Horseshoe Canyon Formation, 72.2-71Ma ago).<ref name=ABS09>{{cite journal |last=Arbour |first=V. M. |author2=Burns, M. E.|author3=Sissons, R. L. |year=2009 |title=A redescription of the ankylosaurid dinosaur ''Dyoplosaurus acutosquameus'' Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus |journal=Journal of Vertebrate Paleontology |volume=29 |issue=4 |pages=1117–1135 |doi=10.1671/039.029.0405}}</ref> |
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A caenagnathid oviraptorosaur known from material previously assigned to ''[[Chirostenotes]]''.<ref name=Sullivan2011/> |
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|[[File:Epichirostenotes.jpg|frameless]] |
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Misidentified as ''[[Anchiceratops]]'', it is actually a new species, probably the same as a new ''[[Pentaceratops]]''-like form from the [[Almond Formation]] of [[Wyoming]] <ref>Farke, A. A. "Cranial osteology and phylogenetic relationships of the chasmosaurine ceratopsid ''Torosaurus latus''", pp. 235-257. In K. Carpenter (ed.). ''Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs''. Indiana Univ. Press (Bloomington), 2006.</ref> |
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===Ornithomimids=== |
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==== Ornithomimids ==== |
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{{paleobiota-key-compact}} |
{{paleobiota-key-compact}} |
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Several specimens. [[type specimen]] |
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An [[ornithomimid]] |
An [[Ornithomimidae|ornithomimid]] |
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|[[File:Ornithomimid.jpg|frameless]] |
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[[File:"Ornithomimus" sp. by Tom Parker.png|thumb|center|150px|''[[Ornithomimus]]'']] |
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[[File:Struthiomimus BW.jpg|thumb|center|150px|''[[Struthiomimus]]'']] |
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Junior synonym of ''O. edmontonicus'' |
Junior synonym of ''O. edmontonicus'' |
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Drumheller, Horsethief, Morrin, and Tolman |
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Units 1-4 |
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Several specimens, [[type specimen]] |
Several specimens, [[type specimen]] |
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An ornithomimid |
An ornithomimid |
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|[[File:Ornithomimus edmontonicus.jpg|frameless]] |
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''[[Struthiomimus]]'' |
''[[Struthiomimus]]'' |
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''S. altus'' |
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Unnamed |
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Drumheller, Horsethief, and Morrin |
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Unit 4 |
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An ornithomimid |
An [[ornithomimid]] |
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|[[File:Struthiomimus ROM.jpg|frameless]] |
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===Ornithopods=== |
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{| class="wikitable" align="center" width="100%" |
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! colspan="7" align="center" |'''[[Ornithopod]]s reported from the Horseshoe Canyon Formation''' |
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! Genus |
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! Species |
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! Location |
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! Stratigraphic position |
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! Material |
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! Notes |
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''[[Edmontosaurus]]'' |
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''E. regalis'' |
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Units 1-2 |
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[[File:Edmontosaurus BW.jpg|thumb|center|150px|''[[Edmontosaurus]]'']] |
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[[File:Hypacrosaurus-v2.jpg|thumb|center|150px|''[[Hypacrosaurus]]'']] |
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[[File:Parksosaurus Steveoc86.jpg|thumb|center|150px|''[[Parksosaurus]]'']] |
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[[File:Saurolophus scalation.png|thumb|center|150px|''[[Saurolophus]]'']] |
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''[[Hypacrosaurus]]'' |
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''H. altispinus'' |
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Units 4-5 |
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"[Five to ten] articulated skulls, some associated with postcrania, isolated skull elements, isolated postcranial elements, many individuals, embryo to adult."<ref name="table-20-1-441">"Table 20.1," in Weishampel, et al. (2004). Page 441.</ref> |
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''[[Parksosaurus]]'' |
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''P. warreni'' |
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Unit 3 |
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''[[Saurolophus]]'' |
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''S. osborni'' |
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Unit 4 |
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"Complete skull and skeleton, [two] complete skulls."<ref name="table-20-1-441"/> |
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===Tyrannosaurs=== |
==== Tyrannosaurs ==== |
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{{paleobiota-key-compact}} |
{{paleobiota-key-compact}} |
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! colspan="7" align="center" |'''[[ |
! colspan="7" align="center" |'''[[Tyrannosaurids]] reported from the Horseshoe Canyon Formation''' |
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! Genus |
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Junior synonym of ''A. sarcophagus'' |
Junior synonym of ''A. sarcophagus'' |
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[[File:Albertosaurus |
[[File:Albertosaurus skull cast.jpg|thumb|center|150px|''[[Albertosaurus]]'']] |
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Horsethief, Morrin, and Tolman; likely present in Drumheller and Carbon. |
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Units 2-5 |
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Several skeletons and partial skeletons, [[type specimen]] |
Several skeletons and partial skeletons, [[type specimen]] |
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A [[Tyrannosauridae|tyrannosaurid]] which was the most common large carnivore in the area.<ref>{{cite journal |last1=Coulton |first1=Richard L. |title=The Theropod Dinosaurs of the Edmonton Formation, Alberta, Canada |journal=Rocks & Minerals |date=1977 |volume=52 |issue=4 |pages=176–181 |doi=10.1080/00357529.1977.11761949 |bibcode=1977RoMin..52..176C |url=https://www.tandfonline.com/doi/abs/10.1080/00357529.1977.11761949?journalCode=vram20 |access-date=21 March 2021}}</ref><ref>{{cite journal |last1=Tanke |first1=Darren H. |last2=Currie |first2=Phillip J. |title=A history of Albertosaurus discoveries in Alberta, Canada |journal=Canadian Journal of Earth Sciences |date=7 September 2010 |volume=47 |issue=9 |pages=1197–1211 |doi=10.1139/e10-057 |bibcode=2010CaJES..47.1197T |url=https://cdnsciencepub.com/doi/abs/10.1139/e10-057 |access-date=21 March 2021}}</ref> |
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A tyrannosaurid |
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|style="background:#f3e9f3;" | |
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<small>''[[Daspletosaurus]]''</small> |
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<small>''[[Daspletosaurus]]''</small><ref name=SciDaily07>{{cite web|url=https://www.sciencedaily.com/releases/2007/06/070629091349.htm|title=City Site Was Dinosaur Dining Room|date=2007-07-03|work=ScienceDaily|publisher=ScienceDaily|accessdate=2008-12-07}}</ref> |
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|style="background:#f3e9f3;" | |
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''D''. sp. |
''D''. sp. |
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|style="background:#f3e9f3;" | |
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|style="background:#f3e9f3;" | |
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Bonebed, Toothmarks on ''[[Saurolophus]]'' and ''[[Edmontosaurus]]'' |
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Suggested from the skeleton of an immature tyrannosaurid (CMN 11315), thorough analysis of this specimen supports a referral to ''A. sarcophagus''.<ref>{{cite journal |last1=Mallon |first1=J.C. |last2=Bura |first2=J.R. |last3=Currie |first3=P.J. |title=A Problematic Tyrannosaurid (Dinosauria: Theropoda) Skeleton and Its Implications for Tyrannosaurid Diversity in the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta |year=2019 |journal=[[The Anatomical Record]] |volume=303 |issue=4 |pages=673–690 |doi=10.1002/ar.24199|doi-access=free |pmid=31254458 |pmc=7079176 }}</ref> An isolated [[maxilla]] and teeth from an ''Edmontosaurus'' bonebed were also mistakenly referred to ''Daspletosaurus'', however all the tyrannosaurid material in the bonebed was confirmed to belong to ''A. sarcophagus''.<ref>{{cite journal|last1=Torices |first1=A. |last2=Reichel |first2=M. |last3=Currie |first3=P.J. |title=Multivariate analysis of isolated tyrannosaurid teeth from the Danek Bonebed, Horseshoe Canyon Formation, Alberta, Canada. |year=2014 |journal=[[Canadian Journal of Earth Sciences]] |volume=51 |issue=11 |pages=1045–1051 |doi=10.1139/cjes-2014-0072|bibcode=2014CaJES..51.1045T }}</ref> |
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A tyrannosaurid. A specimen of ''[[Edmontosaurus regalis]]'' and ''[[Saurolophus osborni]]'' have toothmarks from a species of ''Daspletosaurus''.<ref name=SciDaily07/> |
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==Other animals== |
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==Other Animals== |
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===Mammals=== |
===Mammals=== |
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! Images |
! Images |
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''[[Didelphodon]]''<ref>R. C. Fox and B. G. Naylor. 1986. A new species of Didelphodon Marsh (Marsupialia) from the Upper Cretaceous of Alberta, Canada: paleobiology and phylogeny. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 172(3):357-380 |
''[[Didelphodon]]''<ref>R. C. Fox and B. G. Naylor. 1986. A new species of Didelphodon Marsh (Marsupialia) from the Upper Cretaceous of Alberta, Canada: paleobiology and phylogeny. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 172(3):357-380</ref> |
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''D. coyi'' |
''D. coyi'' |
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a [[stagodontid]] [[metatherian]] |
a [[stagodontid]] [[metatherian]] |
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|[[File:Didelphodon NT small.jpg|center|150px|''[[Didelphodon]]''|frameless]] |
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===Other |
===Other reptiles=== |
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{{paleobiota-key-compact}} |
{{paleobiota-key-compact}} |
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''[[Stangerochampsa]]''<ref>X.-C. Wu, D. B. Brinkman, and A. P. Russell. 1996. A new alligator from the Upper Cretaceous of Canada and the relationships of early eusuchians. Palaeontology 39(2):351-375 |
''[[Stangerochampsa]]''<ref>X.-C. Wu, D. B. Brinkman, and A. P. Russell. 1996. A new alligator from the Upper Cretaceous of Canada and the relationships of early eusuchians. Palaeontology 39(2):351-375</ref> |
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''S. mccabei'' |
''S. mccabei'' |
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''[[Champsosaurus]]''<ref>K. -Q. Gao and R. C. Fox. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, a phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124:303-353 |
''[[Champsosaurus]]''<ref>K. -Q. Gao and R. C. Fox. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, a phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124:303-353</ref> |
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''C. albertensis'' |
''C. albertensis'' |
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a [[choristodere]] |
a [[choristodere]] |
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|[[File:Champsosaurus BW.jpg|center|150px|''[[Champsosaurus]]''|frameless]] |
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''[[Leurospondylus]]''<ref>B. Brown. 1913. A new plesiosaur, Leurospondylus, from the Edmonton Cretaceous of Alberta. Bulletin of the American Museum of Natural History 32(40):605-615 |
''[[Leurospondylus]]''<ref>B. Brown. 1913. A new plesiosaur, Leurospondylus, from the Edmonton Cretaceous of Alberta. Bulletin of the American Museum of Natural History 32(40):605-615</ref> |
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''L. ultimus'' |
''L. ultimus'' |
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''[[Basilemys]]''<ref>Jordan C. |
''[[Basilemys]]''<ref>{{cite journal | last1 = Mallon | first1 = Jordan C. | last2 = Brinkman | first2 = Donald B. | year = 2018 | title = Basilemys morrinensis, a new species of nanhsiungchelyid turtle from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada | journal = Journal of Vertebrate Paleontology | volume = 38| issue = 2 | page = e1431922| doi = 10.1080/02724634.2018.1431922 | bibcode = 2018JVPal..38E1922M | doi-access = free }}</ref> |
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''B. morrinensis'' |
''B. morrinensis'' |
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! Images |
! Images |
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|''[[Boreiosturion]]''<ref>{{Cite journal |last1=Murray |first1=Alison M. |last2=Nelson |first2=Luke E. |last3=Brinkman |first3=Donald B. |date=2023-08-08 |title=A new sturgeon from the Upper Cretaceous Horseshoe Canyon Formation in central Alberta, Canada |url=https://www.tandfonline.com/doi/full/10.1080/02724634.2023.2232846 |journal=Journal of Vertebrate Paleontology |volume=43 |language=en |doi=10.1080/02724634.2023.2232846 |issn=0272-4634}}</ref> |
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|''B. labyrinthicus'' |
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|[[Edmonton]]. |
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|Partial skull. |
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''[[Horseshoeichthys]]''<ref>Canadian Journal of Earth Sciences, 2010, 47(9): 1183-1196, [[https://doi.org/10.1139/E10-041]]</ref> |
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|A [[sturgeon]]. |
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''[[Horseshoeichthys]]''<ref>{{cite journal | url=https://doi.org/10.1139/E10-041 | doi=10.1139/E10-041 | title=A new articulated freshwater fish (Clupeomorpha, Ellimmichthyiformes) from the Horseshoe Canyon Formation, Maastrichtian, of Alberta, CanadaThis article is one of a series of papers published in this Special Issue on the theme ''Albertosaurus'' | year=2010 | last1=Newbrey | first1=Michael G. | last2=Murray | first2=Alison M. | last3=Brinkman | first3=Donald B. | last4=Wilson | first4=Mark V. H. | last5=Neuman | first5=Andrew G. | journal=Canadian Journal of Earth Sciences | volume=47 | issue=9 | pages=1183–1196 }}</ref> |
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''H. armaserratus'' |
''H. armaserratus'' |
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An [[ellimmichthyiform]] |
An [[Ellimmichthyiformes|ellimmichthyiform]] |
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== See also == |
== See also == |
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* [[List of dinosaur-bearing rock formations]] |
* [[List of dinosaur-bearing rock formations]] |
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{{ |
{{Clear}} |
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== References == |
== References == |
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{{Reflist |
{{Reflist}} |
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<ref name="mcer-brain-abs-243">"Abstract," Makovicky (2001); page 243.</ref> |
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}} |
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=== Bibliography === |
=== Bibliography === |
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* Makovicky, P. J., 2001, A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta: In: Mesozoic Vertebrate |
* Makovicky, P. J., 2001, A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 243–262. |
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* Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana. |
* Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana. |
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* Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. {{ISBN|0-520-24209-2}}. |
* Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. {{ISBN|0-520-24209-2}}. |
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{{Western Canadian Sedimentary Basin|Central_Plains=yes}} |
{{Western Canadian Sedimentary Basin|Central_Plains=yes}} |
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{{Authority control}} |
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[[Category:Geologic formations of Alberta]] |
[[Category:Geologic formations of Alberta]] |
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[[Category:Cretaceous Alberta]] |
[[Category:Cretaceous Alberta]] |
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[[Category:Maastrichtian Stage of North America]] |
[[Category:Maastrichtian Stage of North America]] |
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[[Category:Late Cretaceous North America]] |
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[[Category:Upper Cretaceous Series of North America]] |
[[Category:Upper Cretaceous Series of North America]] |
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[[Category:Western Canadian Sedimentary Basin]] |
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[[Category:Sandstone formations]] |
[[Category:Sandstone formations]] |
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[[Category:Shale formations]] |
[[Category:Shale formations]] |
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[[Category:Reservoir rock formations]] |
[[Category:Reservoir rock formations]] |
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[[Category:Source rock formations]] |
[[Category:Source rock formations]] |
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[[Category:Western Canadian Sedimentary Basin]] |
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[[Category:Fossiliferous stratigraphic units of North America]] |
[[Category:Fossiliferous stratigraphic units of North America]] |
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[[Category:Paleontology in Alberta]] |
[[Category:Paleontology in Alberta]] |
Latest revision as of 01:07, 17 November 2024
Horseshoe Canyon Formation | |
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Stratigraphic range: Campanian-Maastrichtian ~[1] | |
Type | Geological formation |
Unit of | Edmonton Group |
Sub-units | Strathmore Member, Drumheller Member, Horsethief Member, Morrin Member, Tolman Member, Carbon Member, Whitemud Member |
Underlies | Battle Formation, Scollard Formation |
Overlies | Bearpaw Formation |
Lithology | |
Primary | Sandstone |
Other | Shale, coal |
Location | |
Coordinates | 51°25′24″N 112°53′18″W / 51.42333°N 112.88833°W |
Region | Alberta Western Canadian Sedimentary Basin |
Country | Canada |
Type section | |
Named for | Horseshoe Canyon |
Named by | E.J.W. Irish, 1970 |
The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta.[2][3] It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.
The Horseshoe Canyon Formation is part of the Edmonton Group. In its type section (Red Deer River Valley at Drumheller), it is ~250 metres (820 ft) thick, but further west the formation is older and thicker, exceeding 500 metres (1,600 ft) near Calgary.[4] It is of Late Cretaceous age, Campanian to early Maastrichtian stage (Edmontonian Land-Mammal Age), and is composed of mudstone, sandstone, carbonaceous shales, and coal seams. A variety of depositional environments are represented in the succession, including floodplains, estuarine channels, and coal swamps, which have yielded a diversity of fossil material. Tidally-influenced estuarine point bar deposits are easily recognizable as Inclined Heterolithic Stratification (IHS). Brackish-water trace fossil assemblages occur within these bar deposits and demonstrate periodic incursion of marine waters into the estuaries.
The Horseshoe Canyon Formation crops out extensively in the area around Drumheller, as well as farther north along the Red Deer River near Trochu and along the North Saskatchewan River in Edmonton.[2] It is overlain by the Battle and Scollard formations.[4] The Drumheller Coal Zone, located in the lower part of the Horseshoe Canyon Formation, was mined for sub-bituminous coal in the Drumheller area from 1911 to 1979, and the Atlas Coal Mine in Drumheller has been preserved as a National Historic Site.[5] In more recent times, the Horseshoe Canyon Formation has become a major target for coalbed methane (CBM) production.
Dinosaurs found in the Horseshoe Canyon Formation include Albertavenator, Albertosaurus, Anchiceratops, Anodontosaurus, Arrhinoceratops, Atrociraptor, Epichirostenotes, Edmontonia, Edmontosaurus, Hypacrosaurus, Ornithomimus, Pachyrhinosaurus, Parksosaurus, Saurolophus, and Struthiomimus. Other finds have included mammals such as Didelphodon coyi, non-dinosaur reptiles, amphibians, fish, marine and terrestrial invertebrates and plant fossils. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers.[6]
Oil/gas production
[edit]The Drumheller Coal Zone has been a primary coalbed methane target for industry. In the area between Bashaw and Rockyford, the Coal Zone lies at relatively shallow depths (about 300 metres) and is about 70 to 120 metres thick. It contains 10 to 20 metres of cumulative coal, in up to 20 or more individual thin seams interbedded with sandstone and shale, which combine to make an attractive multi-completion CBM drilling target. In total, it is estimated there are 14 trillion cubic metres (500 tcf) of gas in place in all the coal in Alberta.
Biostratigraphy
[edit]The timeline below follows work by David A. Eberth and Sandra L. Kamo published in 2019.[7]
Dinosaurs
[edit]Ornithischians
[edit]Ankylosaurs
[edit]Ankylosaurs reported from the Horseshoe Canyon Formation | |||||
---|---|---|---|---|---|
Genus | Species | Stratigraphic position | Material | Notes | Images |
[Five] skulls, mandibles, cervical vertebra, caudal vertebrae, dorsal vertebrae, sacrocaudal vertebrae, sacrum, illium, ischium, partial pelvis, coossified ribs, femur, ?manual phalanx, tail clubs, numerous cervical half-rings and osteoderms.[8][9] |
An ankylosaurine ankylosaurid also known from the middle Dinosaur Park Formation and closely related to Ankylosaurus.[10] |
||||
E. longiceps[11] |
Upper Horsethief[11] |
A skull with mandible, dorsal vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, cervical ribs, dorsal ribs, humerus, radius, ulna, radius, ilia, ischia, both pubes (?), femur, tibia, fibula and osteoderms.[11] |
A panoplosaurin nodosaurid also known from the lower Dinosaur Park Formation and closely related Denversaurus.[12] |
||
E. tutus |
Walter Coombs (1971) synonymised Anodontosaurus lambei with E. tutus. However, recent studies suggest that Anodontosaurus is distinct enough from Euoplocephalus to be placed in its own genus and species.[8][13] Furthermore, all Horseshoe Canyon Formation ankylosaurine specimens were suggested to be reassigned to Anodontosaurus.[9] |
Hadrosaurs
[edit]Hadrosaurs reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Horsethief; likely present in Drumheller. |
||||||
H. altispinus |
Morrin and Tolman. |
"[Five to ten] articulated skulls, some associated with postcrania, isolated skull elements, isolated postcranial elements, many individuals, embryo to adult."[14] |
||||
S. osborni |
Upper Morrin and Tolman. |
"Complete skull and skeleton, [two] complete skulls."[14] |
Marginocephalians
[edit]Ceratopsians
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Ceratopsians reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Stratigraphic position | Material | Notes | Images | |
Horsethief, Morrin, and Tolman; may have been present in Drumheller[16] |
[Two] nearly complete skulls and [seven] partial skulls.[16] |
A chasmosaurine ceratopsid contemporaneous and closely related to Arrhinoceratops.[16] |
||||
A. brachyops[17] |
Horsethief[17] |
A nearly complete skull.[17] |
A chasmosaurine ceratopsid restricted to the holotype specimen as Farke (2007) assigned the referred specimen ROM 1439 to Torosaurus.[18] |
|||
E. xerinsularis[19] |
Carbon[19] |
Premaxillae, maxillae, rostral, supraorbital horncore with lacrimal, prefrontal, frontal, postorbital, jugal, epijugal, quadratojugal, quadrate, partial parietal, squamosal frill, braincase, syncervical, cervical vertebrae, dorsal vertebrae, ribs and fragments of ossified ligaments.[19] |
A large chasmosaurine ceratopsid that may represent a senior synonym of Ojoceratops from the Ojo Alamo Formation.[20] |
|||
Upper Tolman[22] |
An isolated braincase.[22] |
A leptoceratopsid ceratopsian also known from the St. Mary River Formation.[21] |
||||
P. canadensis[23] |
Drumheller and Horsethief[23] |
[Two] partial skulls.[23] |
A centrosaurine ceratopsid also known from the St. Mary River Formation.[23] |
Pachycephalosaurs
[edit]Pachycephalosaurids reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
S. edmontonense[25] |
Tolman[24] |
A pachycephalosaurine pachycephalosaurid also known from the Hell Creek and Kirtland Formation.[25] |
Thescelosaurs
[edit]Thescelosaurids reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
P. warreni |
Tolman |
A partially articulated skeleton and partial skull. | A thescelosaurid neornithischian. |
Theropods
[edit]Maniraptorans
[edit]Maniraptors reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Stratigraphic position | Material | Notes | Images | |
A. curriei[26] |
Horsethief[26] |
[Two] frontals.[26] |
A troodontid theropod that adds to the diversity of North American troodontids.[26] |
|||
A. borealis[27] |
Upper Tolman[27] |
Ulna, tibiae, metatarsals, manual ungual, pedal phalanxes and unidentified pedal phalanges.[27] |
An mononykin alvarezsaurid specialized for digging insect nests.[27] |
|||
A. pennatus[28] |
Horsethief[28] |
A palatine, mandibles, ceratobranchials, axis, cervical vertebrae, dorsal vertebrae, pectoral girdles, sternal plate, forelimb, ilium, femur, tibia and fibula.[28] |
A caenagnathid oviraptorosaur recovered as sister taxon to Elmisaurus .[28] |
|||
A. marshalli[29] |
Lower Horsethief[29] |
Premaxillae, maxilla, dentaries, teeth and numerous bone fragments.[29] |
A dromaeosaurid; teeth indicate it may have been present across all members.[29] |
|||
E. curriei[30] |
Horsethief, Morrin, and Tolman[30] |
A maxilla, probable palatine, partial braincase, cervical vertebrae, dorsal vertebrae, caudal vertebrae, synsacrum, an anterior and a posterior cervical rib, a mid-dorsal rib, fragments of ilia, ischium, both pubes, partial tibia, and unidentifiable bones.[30] |
A caenagnathid oviraptorosaur known from material previously assigned to Chirostenotes.[30] |
Ornithomimids
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Ornithomimids reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
D. brevitertius |
Several specimens. type specimen |
An ornithomimid |
||||
O. currelli |
Junior synonym of O. edmontonicus |
|||||
O. edmontonicus |
Drumheller, Horsethief, Morrin, and Tolman |
Several specimens, type specimen |
An ornithomimid |
|||
S. altus |
Drumheller, Horsethief, and Morrin |
An ornithomimid |
Tyrannosaurs
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Tyrannosaurids reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
A. arctunguis |
Junior synonym of A. sarcophagus |
|||||
A. sarcophagus |
Horsethief, Morrin, and Tolman; likely present in Drumheller and Carbon. |
Several skeletons and partial skeletons, type specimen |
A tyrannosaurid which was the most common large carnivore in the area.[31][32] | |||
D. sp. |
Suggested from the skeleton of an immature tyrannosaurid (CMN 11315), thorough analysis of this specimen supports a referral to A. sarcophagus.[33] An isolated maxilla and teeth from an Edmontosaurus bonebed were also mistakenly referred to Daspletosaurus, however all the tyrannosaurid material in the bonebed was confirmed to belong to A. sarcophagus.[34] |
Other animals
[edit]Mammals
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Mammals reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
D. coyi |
Other reptiles
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Reptiles reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
S. mccabei |
"a skull, partial lower jaws, and partial postcranial skeleton" |
An alligatoroid |
||||
C. albertensis |
"partial skeleton with partial skull" |
|||||
L. ultimus |
"a partial skeleton" |
a plesiosaur of uncertain classification |
||||
B. morrinensis |
"nearly complete shell" |
Fish
[edit]Color key
|
Notes Uncertain or tentative taxa are in small text; |
Fish reported from the Horseshoe Canyon Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Boreiosturion[40] | B. labyrinthicus | Edmonton. | Partial skull. | A sturgeon. | ||
H. armaserratus |
See also
[edit]References
[edit]- ^ Eberth, David A.; Kamo, Sandra L. (October 2020). "High-precision U–Pb CA–ID–TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada". Canadian Journal of Earth Sciences. 57 (10): 1220–1237. Bibcode:2020CaJES..57.1220E. doi:10.1139/cjes-2019-0019. ISSN 0008-4077.
- ^ a b Prior, G. J., Hathaway, B., Glombick, P.M., Pana, D.I., Banks, C.J., Hay, D.C., Schneider, C.L., Grobe, M., Elgr, R., and Weiss, J.A. (2013). "Bedrock Geology of Alberta. Alberta Geological Survey, Map 600". Archived from the original on 2013-07-05. Retrieved 2013-08-13.
{{cite web}}
: CS1 maint: multiple names: authors list (link) - ^ Mossop, G.D. and Shetsen, I., (compilers), Canadian Society of Petroleum Geologists (1994). "The Geological Atlas of the Western Canada Sedimentary Basin, Chapter 24: Upper Cretaceous and Tertiary strata of the Western Canada Sedimentary Basin". Archived from the original on 2013-07-21. Retrieved 2013-08-01.
{{cite web}}
: CS1 maint: multiple names: authors list (link) - ^ a b Eberth, David A.; Braman, Dennis R. (September 2012). "A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains". Canadian Journal of Earth Sciences. 49 (9): 1053–1086. Bibcode:2012CaJES..49.1053E. doi:10.1139/e2012-035. ISSN 0008-4077.
- ^ "Mine History". Atlas Coal Mine National Historic Site. Archived from the original on 23 August 2010. Retrieved 9 June 2010.
- ^ Quinney, Annie; Therrien, François; Zelenitsky, Darla K.; Eberth, David A. (2013). "Palaeoenvironmental and palaeoclimatic reconstruction of the Upper Cretaceous (late Campanian–early Maastrichtian) Horseshoe Canyon Formation, Alberta, Canada". Palaeogeography, Palaeoclimatology, Palaeoecology. 371: 26–44. Bibcode:2013PPP...371...26Q. doi:10.1016/j.palaeo.2012.12.009.
- ^ Eberth, David A.; Kamo, Sandra (2019). "High-precision U-Pb CA-ID-TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada". Canadian Journal of Earth Sciences. 57 (10): 1220–1237. Bibcode:2020CaJES..57.1220E. doi:10.1139/cjes-2019-0019. S2CID 210299227.
- ^ a b c d e Penkalski, P. (2013). "A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA". Acta Palaeontologica Polonica. doi:10.4202/app.2012.0125.
- ^ a b c d e Arbour, Victoria (2010). "A Cretaceous armoury: Multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA". Journal of Vertebrate Paleontology. 30 (Supplement 2): 55A. doi:10.1080/02724634.2010.10411819.
- ^ Paul Penkalski (2018). Revised systematics of the armoured dinosaur Euoplocephalus and its allies. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 287(3): 261-306. doi: https://doi.org/10.1127/njgpa/2018/0717
- ^ a b c d Sternberg, C.M. (1928). "A new armored dinosaur from the Edmonton Formation of Alberta". Transactions of the Royal Society of Canada. Series 3. 22: 93–106.
- ^ Russell, L.S. (1940). "Edmontonia rugosidens (Gilmore), an armored dinosaur from the Belly River Series of Alberta". University of Toronto Studies, Geology Series. 43: 3–28.
- ^ Penkalski, P.; Blows, W. T. (2013). "Scolosaurus cutleri (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada". Canadian Journal of Earth Sciences. 50 (2): 171–182. Bibcode:2013CaJES..50..171P. doi:10.1139/cjes-2012-0098.
- ^ a b "Table 20.1," in Weishampel, et al. (2004). Page 441.
- ^ a b Brown, B (1914). "Anchiceratops, a new genus of horned dinosaurs from the Edmonton Cretaceous of Alberta. With a discussion of the origin of the ceratopsian crest and the brain casts of Anchiceratops and Trachodon"". Bulletin of the American Museum of Natural History. 33: 539–548.
- ^ a b c d e Mallon, Jordan C.; Holmes, Robert; Eberth, David A.; Ryan, Michael J.; Anderson, Jason S. (2012). "Variation in the skull of Anchiceratops (Dinosauria, Ceratopsidae) from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta". Journal of Vertebrate Paleontology. 31 (5): 1047–1071. doi:10.1080/02724634.2011.601484. S2CID 86349131.
- ^ a b c d Parks, W.A. (1925). "Arrhinoceratops brachyops, a new genus and species of Ceratopsia from the Edmonton Formation of Alberta". University of Toronto Studies, Geology Series 19:1-15
- ^ Farke, A.A., 2007, "Cranial osteology and phylogenetic relationships of the chasmosaurine ceratopsid Torosaurus latus", pp 235-257 in: K. Carpenter (ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, Bloomington, Indiana University Press
- ^ a b c d Wu, X-C.; Brinkman, D.B.; Eberth, D.A.; Braman, D.R. (2007). "A new ceratopsid dinosaur (Ornithischia) from the uppermost Horseshoe Canyon Formation (upper Maastrichtian), Alberta, Canada". Canadian Journal of Earth Sciences. 44 (9): 1243–1265. Bibcode:2007CaJES..44.1243W. doi:10.1139/E07-011.
- ^ Wick, S. L.; Lehman, T. M. (2013). "A new ceratopsian dinosaur from the Javelina Formation (Maastrichtian) of West Texas and implications for chasmosaurine phylogeny". Naturwissenschaften. 100 (7): 667–82. Bibcode:2013NW....100..667W. doi:10.1007/s00114-013-1063-0. PMID 23728202. S2CID 16048008.
- ^ a b c Brown, B.; Schlaikjer, E. M. (1942). "The skeleton of Leptoceratops with the description of a new species". American Museum Novitates (1169): 1–15.
- ^ a b c d P. J. Makovicky. 2001. A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta. Mesozoic Vertebrate Life, D. H. Tanke and K. Carpenter (eds.), Indiana University Press, Bloomington 243-262
- ^ a b c d e Sternberg, C. M. (1950). "Pachyrhinosaurus canadensis, representing a new family of the Ceratopsia, from southern Alberta". National Museum of Canada Bulletin. 118: 109–120.
- ^ a b c Longrich N. R.; Sankey J. T.; et al. (2010). "Texacephale langstoni, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA". Cretaceous Research. 31 (2): 274–284. Bibcode:2010CrRes..31..274L. doi:10.1016/j.cretres.2009.12.002.
- ^ a b c d Williamson Thomas E.; Carr Thomas D. (2002). "A new genus of highly derived pachycephalosaurian from western North America". Journal of Vertebrate Paleontology. 22 (4): 779–801. doi:10.1671/0272-4634(2002)022[0779:angodp]2.0.co;2. S2CID 86112901.
- ^ a b c d e Evans, D.C., Cullen, T.M., Larson, D.W., and Rego, A. "A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada." Canadian Journal of Earth Sciences Early Online: 813-826. DOI: dx.doi.org/10.1139/cjes-2017-0034
- ^ a b c d e Longrich, Nicholas R.; Currie, Philip J. (2009). "Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae". Cretaceous Research. 30 (1): 239–252. Bibcode:2009CrRes..30..239L. doi:10.1016/j.cretres.2008.07.005.
- ^ a b c d e Gregory F. Funston; Philip J. Currie (2016). "A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae". Journal of Vertebrate Paleontology. 36 (4): e1160910. Bibcode:2016JVPal..36E0910F. doi:10.1080/02724634.2016.1160910. S2CID 131090028.
- ^ a b c d e Currie, P. J. and D. J. Varricchio (2004). "A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada". Pp. 112–132 in P. J. Currie, E. B. Koppelhus, M. A. Shugar and J. L. Wright. (eds.), Feathered Dragons. Indianapolis: Indiana University Press. [1]
- ^ a b c d e Robert M. Sullivan; Steven E. Jasinski; Mark P.A. Van Tomme (2011). "A new caenagnathid Ojoraptorsaurus boerei, n. gen., n. sp. (Dinosauria, Oviraptorosauria), from the Upper Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico" (PDF). Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin. 53: 418–428.
- ^ Coulton, Richard L. (1977). "The Theropod Dinosaurs of the Edmonton Formation, Alberta, Canada". Rocks & Minerals. 52 (4): 176–181. Bibcode:1977RoMin..52..176C. doi:10.1080/00357529.1977.11761949. Retrieved 21 March 2021.
- ^ Tanke, Darren H.; Currie, Phillip J. (7 September 2010). "A history of Albertosaurus discoveries in Alberta, Canada". Canadian Journal of Earth Sciences. 47 (9): 1197–1211. Bibcode:2010CaJES..47.1197T. doi:10.1139/e10-057. Retrieved 21 March 2021.
- ^ Mallon, J.C.; Bura, J.R.; Currie, P.J. (2019). "A Problematic Tyrannosaurid (Dinosauria: Theropoda) Skeleton and Its Implications for Tyrannosaurid Diversity in the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta". The Anatomical Record. 303 (4): 673–690. doi:10.1002/ar.24199. PMC 7079176. PMID 31254458.
- ^ Torices, A.; Reichel, M.; Currie, P.J. (2014). "Multivariate analysis of isolated tyrannosaurid teeth from the Danek Bonebed, Horseshoe Canyon Formation, Alberta, Canada". Canadian Journal of Earth Sciences. 51 (11): 1045–1051. Bibcode:2014CaJES..51.1045T. doi:10.1139/cjes-2014-0072.
- ^ R. C. Fox and B. G. Naylor. 1986. A new species of Didelphodon Marsh (Marsupialia) from the Upper Cretaceous of Alberta, Canada: paleobiology and phylogeny. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 172(3):357-380
- ^ X.-C. Wu, D. B. Brinkman, and A. P. Russell. 1996. A new alligator from the Upper Cretaceous of Canada and the relationships of early eusuchians. Palaeontology 39(2):351-375
- ^ K. -Q. Gao and R. C. Fox. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, a phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124:303-353
- ^ B. Brown. 1913. A new plesiosaur, Leurospondylus, from the Edmonton Cretaceous of Alberta. Bulletin of the American Museum of Natural History 32(40):605-615
- ^ Mallon, Jordan C.; Brinkman, Donald B. (2018). "Basilemys morrinensis, a new species of nanhsiungchelyid turtle from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada". Journal of Vertebrate Paleontology. 38 (2): e1431922. Bibcode:2018JVPal..38E1922M. doi:10.1080/02724634.2018.1431922.
- ^ Murray, Alison M.; Nelson, Luke E.; Brinkman, Donald B. (2023-08-08). "A new sturgeon from the Upper Cretaceous Horseshoe Canyon Formation in central Alberta, Canada". Journal of Vertebrate Paleontology. 43. doi:10.1080/02724634.2023.2232846. ISSN 0272-4634.
- ^ Newbrey, Michael G.; Murray, Alison M.; Brinkman, Donald B.; Wilson, Mark V. H.; Neuman, Andrew G. (2010). "A new articulated freshwater fish (Clupeomorpha, Ellimmichthyiformes) from the Horseshoe Canyon Formation, Maastrichtian, of Alberta, CanadaThis article is one of a series of papers published in this Special Issue on the theme Albertosaurus". Canadian Journal of Earth Sciences. 47 (9): 1183–1196. doi:10.1139/E10-041.
Bibliography
[edit]- Makovicky, P. J., 2001, A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 243–262.
- Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana.
- Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN 0-520-24209-2.
- Geologic formations of Alberta
- Cretaceous Alberta
- Maastrichtian Stage of North America
- Upper Cretaceous Series of North America
- Sandstone formations
- Shale formations
- Coal formations
- Fluvial deposits
- Tidal deposits
- Reservoir rock formations
- Source rock formations
- Western Canadian Sedimentary Basin
- Fossiliferous stratigraphic units of North America
- Paleontology in Alberta