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'''''Anacamptis morio''''', the '''green-winged orchid''' or '''green-veined orchid''' ([[synonym (taxonomy)|synonym]] ''Orchis morio''), is a [[flowering plant]] of the [[orchid]] family, [[Orchidaceae]]. It usually has purple flowers, and is found in [[Europe]] and [[the Middle East]].
'''''Anacamptis morio''''', the '''green-winged orchid''' or '''green-veined orchid''' ([[synonym (taxonomy)|synonym]] ''Orchis morio''), is a [[flowering plant]] of the [[orchid]] family, [[Orchidaceae]]. It usually has purple flowers, and is found in [[Europe]] and [[the Middle East]].


== Description ==
==Description==

It flowers from late April to June in the British Isles, and as early as February in other countries, such as France. The inflorescence is of various colours, mainly purple but ranging from white, through pink, to deep purple. From 5 to 25 helmet-shaped flowers grow in a loose, linear bunch at the top of the single stalk. A pair of lateral [[sepals]] with prominent green, occasionally purple veins extend laterally like "wings", giving the orchid its name. The broad, three lobed, lower petal is pale in the center with dark spots.
It flowers from late April to June in the British Isles, and as early as February in other countries, such as France. The inflorescence is of various colours, mainly purple but ranging from white, through pink, to deep purple. From 5 to 25 helmet-shaped flowers grow in a loose, linear bunch at the top of the single stalk. A pair of lateral [[sepals]] with prominent green, occasionally purple veins extend laterally like "wings", giving the orchid its name. The broad, three lobed, lower petal is pale in the center with dark spots.


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The species was first described, as ''[[Orchis]] morio'', by [[Carl Linnaeus]], in 1753. It was transferred to the genus ''[[Anacamptis]]'' in 1997.<ref name=WCSP_8585/> ''Anacamptis'' comes from the Greek ανακάμτειν "anakamptein" which means to bend, although according to different sources it may mean to bend backward, to bend down or to bend forward.<ref>[https://books.google.co.uk/books?id=rQIsAwAAQBAJ&pg=PA164&lpg=PA164&dq=anakamptein+bend&source=bl&ots=V0OtMXp3wR&sig=ACfU3U3UqIWbpWFBaamC3IZ_DTiTWiOqzg&hl=en&sa=X&ved=2ahUKEwiapaDts_zhAhUrSxUIHcJFBk4Q6AEwBHoECAYQAQ#v=onepage&q=anakamptein%20bend&f=false Philoponus - On Aristotle On The Soul]</ref><ref>[https://books.google.co.uk/books?id=gf3uCAAAQBAJ&pg=PA9&lpg=PA9&dq=anakamptein+bend&source=bl&ots=N4X1ebW11c&sig=ACfU3U21p72AhQfciluUIJz5FHCLx3mZZQ&hl=en&sa=X&ved=2ahUKEwiapaDts_zhAhUrSxUIHcJFBk4Q6AEwDnoECAkQAQ#v=onepage&q=anakamptein%20bend&f=false Etymological Dictionary of Succulent Plant Names]</ref><ref>[https://species.nbnatlas.org/species/NBNSYS0000002347 NBN Atlas - Anacamptis pyramidalis]</ref> The name ''morio'' is Latin for "[[clown]]", which its striped and spotted flowers were held to resemble.<ref>[https://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/green-winged-orchid Plantlife - Green Winged Orchid]</ref><ref>[https://books.google.co.uk/books?id=_hVnDwAAQBAJ&pg=PA102&lpg=PA102&dq=morio+greek+fool+clown&source=bl&ots=94qhPX-c6G&sig=ACfU3U2dlMBDsj4hm7GcqsRy1cvEDenkxQ&hl=en&sa=X&ved=2ahUKEwiRy5CJsvzhAhXdURUIHQ0VDrM4ChDoATACegQIBxAB#v=onepage&q=morio%20greek%20fool%20clown&f=false The Poor, the Crippled, the Blind and the Lame: Physical and Sensory Disability in the Gospels of the New Testament]</ref>
The species was first described, as ''[[Orchis]] morio'', by [[Carl Linnaeus]], in 1753. It was transferred to the genus ''[[Anacamptis]]'' in 1997.<ref name=WCSP_8585/> ''Anacamptis'' comes from the Greek ανακάμτειν "anakamptein" which means to bend, although according to different sources it may mean to bend backward, to bend down or to bend forward.<ref>[https://books.google.co.uk/books?id=rQIsAwAAQBAJ&pg=PA164&lpg=PA164&dq=anakamptein+bend&source=bl&ots=V0OtMXp3wR&sig=ACfU3U3UqIWbpWFBaamC3IZ_DTiTWiOqzg&hl=en&sa=X&ved=2ahUKEwiapaDts_zhAhUrSxUIHcJFBk4Q6AEwBHoECAYQAQ#v=onepage&q=anakamptein%20bend&f=false Philoponus - On Aristotle On The Soul]</ref><ref>[https://books.google.co.uk/books?id=gf3uCAAAQBAJ&pg=PA9&lpg=PA9&dq=anakamptein+bend&source=bl&ots=N4X1ebW11c&sig=ACfU3U21p72AhQfciluUIJz5FHCLx3mZZQ&hl=en&sa=X&ved=2ahUKEwiapaDts_zhAhUrSxUIHcJFBk4Q6AEwDnoECAkQAQ#v=onepage&q=anakamptein%20bend&f=false Etymological Dictionary of Succulent Plant Names]</ref><ref>[https://species.nbnatlas.org/species/NBNSYS0000002347 NBN Atlas - Anacamptis pyramidalis]</ref> The name ''morio'' is Latin for "[[clown]]", which its striped and spotted flowers were held to resemble.<ref>[https://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/green-winged-orchid Plantlife - Green Winged Orchid]</ref><ref>[https://books.google.co.uk/books?id=_hVnDwAAQBAJ&pg=PA102&lpg=PA102&dq=morio+greek+fool+clown&source=bl&ots=94qhPX-c6G&sig=ACfU3U2dlMBDsj4hm7GcqsRy1cvEDenkxQ&hl=en&sa=X&ved=2ahUKEwiRy5CJsvzhAhXdURUIHQ0VDrM4ChDoATACegQIBxAB#v=onepage&q=morio%20greek%20fool%20clown&f=false The Poor, the Crippled, the Blind and the Lame: Physical and Sensory Disability in the Gospels of the New Testament]</ref>


== Subspecies ==
==Subspecies==
{{As of|2014|May}}, the [[World Checklist of Selected Plant Families]] accepts six subspecies:<ref name=WCSP_Anacamptis_morio>{{Cite web |title=Search for ''Anacamptis morio''|work=[[World Checklist of Selected Plant Families]] |publisher=[[Royal Botanic Gardens, Kew]] |url=http://apps.kew.org/wcsp/qsearch.do?plantName=Anacamptis%20morio |access-date=2014-05-17 }}</ref>
{{As of|2014|May}}, the [[World Checklist of Selected Plant Families]] accepts six subspecies:<ref name=WCSP_Anacamptis_morio>{{Cite web |title=Search for ''Anacamptis morio''|work=[[World Checklist of Selected Plant Families]] |publisher=[[Royal Botanic Gardens, Kew]] |url=http://apps.kew.org/wcsp/qsearch.do?plantName=Anacamptis%20morio |access-date=2014-05-17 }}</ref>


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*''[[Anacamptis morio subsp. syriaca|Anacamptis morio'' subsp. ''syriaca'']]
*''[[Anacamptis morio subsp. syriaca|Anacamptis morio'' subsp. ''syriaca'']]


== Distribution and habitat ==
==Distribution and habitat==

It is a native of western [[Eurasia]], ranging from [[Europe]] to [[Iran]]. In the [[British Isles]] it is found in central-southern England, Wales and Ireland.
It is a native of western [[Eurasia]], ranging from [[Europe]] to [[Iran]]. In the [[British Isles]] it is found in central-southern England, Wales and Ireland.


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The maximum altitude for this species is somewhere between 1500 and 2000m.<ref>[https://bgflora.net/families/orchidaceae/orchis/orchis_morio/orchis_morio_en.html The Bulgarian Flora Online]</ref><ref>[https://www.monaconatureencyclopedia.com/anacamptis-morio/?lang=en Monaco Nature Encyclopedia]</ref>
The maximum altitude for this species is somewhere between 1500 and 2000m.<ref>[https://bgflora.net/families/orchidaceae/orchis/orchis_morio/orchis_morio_en.html The Bulgarian Flora Online]</ref><ref>[https://www.monaconatureencyclopedia.com/anacamptis-morio/?lang=en Monaco Nature Encyclopedia]</ref>


[[Image:Orchis morio plants.jpg|thumb|upright|left|Green-winged orchid (''A. morio'') in bloom in meadow habitat]]
[[Image:Orchis morio plants.jpg|thumb|upright|left|In bloom in meadow habitat]]


== Ecology ==
==Ecology==
⚫
Pollination is by bees.<ref name=":1" /><ref>[https://europepmc.org/articles/pmc1691667/pdf/15255098.pdf The Royal Society - The effects of nectar addition on pollen removal and geitonogamy in the non-rewarding orchid Anacamptis morio]</ref>

⚫
Pollination is by [[bees]].<ref name=":1" /><ref>[https://europepmc.org/articles/pmc1691667/pdf/15255098.pdf The Royal Society - The effects of nectar addition on pollen removal and geitonogamy in the non-rewarding orchid Anacamptis morio]</ref>


The flowers do not produce nectar, but can attract pollinators with their visual appearance. This nectar deception "facilitates the mixing of pollen between different individual plants, promotes genetic diversity in the species, and has been favored evolutionarily over nectar production."<ref>[http://blogs.britannica.com/2011/03/deceptive-flowers-orchids/ Britannica Blog: The Deceptive Flowers of Orchids]</ref>
The flowers do not produce nectar, but can attract pollinators with their visual appearance. This nectar deception "facilitates the mixing of pollen between different individual plants, promotes genetic diversity in the species, and has been favored evolutionarily over nectar production."<ref>[http://blogs.britannica.com/2011/03/deceptive-flowers-orchids/ Britannica Blog: The Deceptive Flowers of Orchids]</ref>
Line 58: Line 55:
Plants cannot establish without a mycorrhizal partner. This makes them vulnerable to chemicals, particularly fungicides, but also other chemical applications, including fertilisers, which could reduce the prevalence of particular species of fungi. [[Mycorrhizal fungi]] known to grow in association with the green-winged orchid include ''[[Epulorhiza]] repens'' ([[Tulasnellaceae]]) and ''Moniliopsis solani'' ([[Ceratobasidiaceae]]).<ref name=":1" />
Plants cannot establish without a mycorrhizal partner. This makes them vulnerable to chemicals, particularly fungicides, but also other chemical applications, including fertilisers, which could reduce the prevalence of particular species of fungi. [[Mycorrhizal fungi]] known to grow in association with the green-winged orchid include ''[[Epulorhiza]] repens'' ([[Tulasnellaceae]]) and ''Moniliopsis solani'' ([[Ceratobasidiaceae]]).<ref name=":1" />


== Conservation ==
==Conservation==

This species' conservation status is vulnerable and near threatened.<ref>[https://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/green-winged-orchid Plantlife - Green-winged orchid]</ref>
This species' conservation status is vulnerable and near threatened.<ref>[https://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/green-winged-orchid Plantlife - Green-winged orchid]</ref>


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[[File:Anacamptis Fleur global.jpg|thumb|center|upright=1.6| Flower]]
[[File:Anacamptis Fleur global.jpg|thumb|center|upright=1.6| Flower]]


== References ==
==References==
{{Reflist}}
{{Reflist}}


== External links ==
==External links==
{{Commons}}
* {{Commons-inline|Anacamptis morio|''Anacamptis morio''}}
* {{Wikispecies-inline|Anacamptis morio|''Anacamptis morio''}}
* {{eol|1099026}} Images. Habitats, habitus etc. and images of ''Anacamptis morio'' subsp. ''morio'' Anacamptis ''morio'' subsp. ''longicornu'', ''Anacamptis morio'' subsp. ''picta'', ''Anacamptis morio'' subsp. ''syriaca''
* {{eol|1099026}} Images. Habitats, habitus etc. and images of ''Anacamptis morio'' subsp. ''morio'' Anacamptis ''morio'' subsp. ''longicornu'', ''Anacamptis morio'' subsp. ''picta'', ''Anacamptis morio'' subsp. ''syriaca''
* [http://linnaeus.nrm.se/flora/mono/orchida/orchi/orchmorv.jpg Den virtuella floran - Distribution]
* [http://linnaeus.nrm.se/flora/mono/orchida/orchi/orchmorv.jpg Den virtuella floran - Distribution]

Revision as of 19:13, 10 April 2021

Anacamptis morio
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Asparagales
Family: Orchidaceae
Subfamily: Orchidoideae
Genus: Anacamptis
Species:
A. morio
Binomial name
Anacamptis morio
Synonyms[2]
  • Orchis morio L.
  • Herorchis morio (L.) D.Tyteca & E.Klein

Anacamptis morio, the green-winged orchid or green-veined orchid (synonym Orchis morio), is a flowering plant of the orchid family, Orchidaceae. It usually has purple flowers, and is found in Europe and the Middle East.

Description

It flowers from late April to June in the British Isles, and as early as February in other countries, such as France. The inflorescence is of various colours, mainly purple but ranging from white, through pink, to deep purple. From 5 to 25 helmet-shaped flowers grow in a loose, linear bunch at the top of the single stalk. A pair of lateral sepals with prominent green, occasionally purple veins extend laterally like "wings", giving the orchid its name. The broad, three lobed, lower petal is pale in the center with dark spots.

Leaves are lanceolate, or sometimes ovate, and grow in a rosette around the base of the plan, with some thinner leaves clasping the stem and sheathing almost up to the flowers. Leaves are green and unspotted. Plants grow to 40 cm in height.

It is similar in appearance to the early purple orchid Orchis mascula, which flowers around the same time of year, but Anacamptis morio has green stripes on the two lateral sepals, and lacks the spots or blotches of the Early Purple's leaves.

Individual plants may flower for up to 17 years.[3]

Taxonomy

The species was first described, as Orchis morio, by Carl Linnaeus, in 1753. It was transferred to the genus Anacamptis in 1997.[2] Anacamptis comes from the Greek ανακάμτειν "anakamptein" which means to bend, although according to different sources it may mean to bend backward, to bend down or to bend forward.[4][5][6] The name morio is Latin for "clown", which its striped and spotted flowers were held to resemble.[7][8]

Subspecies

As of May 2014, the World Checklist of Selected Plant Families accepts six subspecies:[9]

Distribution and habitat

It is a native of western Eurasia, ranging from Europe to Iran. In the British Isles it is found in central-southern England, Wales and Ireland.

It grows in unimproved grassy meadows, especially on limestone-rich soil. This species thrives where grass is cut once or twice a year after flowering is complete, or where it is grazed after flowering is complete.[3] Cutting or mowing should not take place immediately after flowering but give time for seed dispersal.

It can grow in dry or wet grazed meadows. It can also be found in coastal grasslands, quarries, churchyards, as well as on roadsides and lawns.[3] On the European continent it is also found in alpine pasture and in xerothermic grassland on porphyry outcrops.[3]

The maximum altitude for this species is somewhere between 1500 and 2000m.[10][11]

In bloom in meadow habitat

Ecology

Pollination is by bees.[3][12]

The flowers do not produce nectar, but can attract pollinators with their visual appearance. This nectar deception "facilitates the mixing of pollen between different individual plants, promotes genetic diversity in the species, and has been favored evolutionarily over nectar production."[13]

Plants cannot establish without a mycorrhizal partner. This makes them vulnerable to chemicals, particularly fungicides, but also other chemical applications, including fertilisers, which could reduce the prevalence of particular species of fungi. Mycorrhizal fungi known to grow in association with the green-winged orchid include Epulorhiza repens (Tulasnellaceae) and Moniliopsis solani (Ceratobasidiaceae).[3]

Conservation

This species' conservation status is vulnerable and near threatened.[14]

It is a protected species in Northern Ireland under the Wildlife (NI) Order of 1985.[15]

In 2001 Anacamptis morio was adopted as the logo for Priory Vale, the third and final instalment in Swindon's 'Northern Expansion' project. Due to a rapid decline in the species they are protected in certain cases, although still regarded as being quite common in the Swindon area, especially Clifford Meadow, a Site of Special Scientific Interest (SSSI) off Thamesdown Drive, Swindon.

Flower

References

  1. ^ R. M. Bateman; A. M. Pridgeon; M. W. Chase (1997). "Phylogenetics of subtribe Orchidinae (Orchidoideae, Orchidaceae) based on nuclear its sequences. 2. Infrageneric relationships and reclassification to achieve monophyly of Orchis sensu stricto". Lindleyana. 12: 113–141.
  2. ^ a b "Anacamptis morio". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew. Retrieved 2014-05-17.
  3. ^ a b c d e f BSBI Species Account - Anacamptis morio
  4. ^ Philoponus - On Aristotle On The Soul
  5. ^ Etymological Dictionary of Succulent Plant Names
  6. ^ NBN Atlas - Anacamptis pyramidalis
  7. ^ Plantlife - Green Winged Orchid
  8. ^ The Poor, the Crippled, the Blind and the Lame: Physical and Sensory Disability in the Gospels of the New Testament
  9. ^ "Search for Anacamptis morio". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew. Retrieved 2014-05-17.
  10. ^ The Bulgarian Flora Online
  11. ^ Monaco Nature Encyclopedia
  12. ^ The Royal Society - The effects of nectar addition on pollen removal and geitonogamy in the non-rewarding orchid Anacamptis morio
  13. ^ Britannica Blog: The Deceptive Flowers of Orchids
  14. ^ Plantlife - Green-winged orchid
  15. ^ Orchis morio :: Flora of Northern Ireland web site
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