Arnica

Arnica
Chamisso Arnica (Arnica chamissonis)
Scientific classification
Kingdom:	Plantae
Division:	Magnoliophyta
Class:	Magnoliopsida
Order:	Asterales
Family:	Asteraceae
Subfamily:	Asteroideae
Tribe:	Heliantheae
Subtribe:	Madiinae
Genus:	Arnica L.
Species
See text.

Arnica is a genus with about 30 perennial , herbaceous species, belonging to the sunflower family (Asteraceae). The name Arnica means lamb's skin referring to the soft, hairy leaves.

This circumboreal and montane genus occurs mostly in the temperate regions of western North America, while two are native to Eurasia (A. angustifolia and A. montana).

Arnica used to be included in the tribe Senecioneae, because it has a pappus of fine bristles. This was soon questioned and Nordenstam (1977) placed it tentatively in tribe Heliantheae s.l. This arrangement became also uncertain because of the sesquiterpene lactone chemistry in certain species. Lately Arnica was placed in an unresolved clade together with Madiinae, Eupatorieae, Heliantheae s.s. and Pectidinae.

Several species, such as Arnica montana and Arnica chamissonis contain helenalin, a sesquiterpene lactone that is a major ingredient in anti-inflammatory preparations (mostly against bruises).

Arnica species are used as food plants by the larvae of some Lepidoptera species including Bucculatrix arnicella. In folk and holistic medicine, Arnica has been used as a salve for bruises and sprains [1], and sometimes as a tincture [2], for the same purpose.

Characteristics

They have a deep-rooted, erect stem, that is usually unbranched. Their downy, opposite leaves are borne towards the apex of the stem. The ovoid, leathery, basal leaves are arranged in a rosette.

They show large yellow or orange flowers, 6-8 cm wide with 10-15 long ray florets and numerous disc florets. The phyllaries (= a bract under the flowerhead) has long spreading hairs Each phyllary is associated with a ray floret. Species of Arnica, with an involucre (= a circle of bracts arranged surrounding the flower head) arranged in two rows, have only their outer phyllaries associated with ray florets. The flowers have a slight aromatic smell.

The seed-like fruit has a pappus of plumose, white or pale tan bristles.

Species

• Arnica acaulis (Walt.) B.S.P. -- Common Leopardbane
• Arnica alpina (L.) Olin -- Alpine Arnica (synonym of Arnica angustifolia subsp. alpina)
• Arnica amplexicaulis Nutt. -- Clasping Arnica, Streambank Arnica (synonym of Arnica lanceolata subsp. amplexicaulis)
• Arnica angustifolia Vahl -- Narrowleaf Arnica
  • Arnica angustifolia subsp. alpina (L.) I. K. Ferguson
  • Arnica angustifolia subsp. tomentosa Downie & Denford
• Arnica cernua T.J. Howell -- Serpentine Arnica
• Arnica chamissonis Less. -- Chamisso Arnica
  • Arnica chamissonis subsp. foliosa (Nutt.) Maguire
• Arnica cordifolia Hook. -- Heart-leaf Leopardbane, Heartleaf Arnica
• Arnica dealbata Baldwin (formerly Whitneya dealbata)
• Arnica discoidea Benth. -- Rayless Arnica
• Arnica X diversifolia Greene (pro sp.) -- Curtis Churchmouse Threeawn, Rayless Arnica, Sticky Arnica
• Arnica frigida C.A. Mey. ex Iljin -- Snow Arnica (synonym of Arnica griscomii subsp. frigida)
• Arnica fulgens Pursh -- Foothill Arnica, Orange Arnica, Shining Leopardbane
• Arnica x gracilis Rydb. -- Smallhead Arnica (a natural hybrid between A. latifolia and A. cordifolia)
• Arnica griscomii Fernald
  • Arnica griscomii subsp. frigida (C. A. Mey. ex Iljin) S. J. Wolf
  • Arnica griscomii subsp. griscomii Downie & Denford
• Arnica lanceolata Nutt. -- Arnica, Lanceleaf Arnica
  • Arnica lanceolata subsp. amplexicaulis (Nutt.) Gruezo & Denford
  • Arnica lanceolata subsp. lanceolata Gruezo & Denford
• Arnica latifolia Bong. -- Broadleaf Arnica
• Arnica lessingii (Torr. & Gray) Greene -- Nodding Arnica
  • Arnica lessingii subsp. lessengii
  • Arnica lessingii subsp. norbergii Hult. & Maguire
• Arnica lonchophylla Greene -- Longleaf Arnica
  • Arnica lonchophylla subsp. arnoglossa (Greene) Maguire
  • Arnica lonchophylla subsp. lonchophylla
• Arnica longifolia D.C. Eat. -- Longleaf Arnica, Spearleaf Arnica
• Arnica louiseana Farr -- Lake Louise Arnica
• Arnica mallotopus (formerly Mallotopus japonicus)
• Arnica mollis Hook. -- hairy arnica, wooly arnica
• Arnica montana L. -- Mountain Arnica
• Arnica nevadensis Gray -- Nevada Arnica
• Arnica ovata Greene
• Arnica parryi Gray -- Nodding Arnica, Parry's Arnica
• Arnica rydbergii Greene -- Rydberg Arnica, Rydberg's Arnica, Subalpine Arnica
• Arnica sachalinensis (Regel) A. Gray
• Arnica sororia Greene -- Twin Arnica
• Arnica spathulata Greene -- Klamath Arnica
• Arnica unalaschcensis Less. -- Alaska Arnica
• Arnica venosa Hall -- Shasta County Arnica
• Arnica viscosa Gray -- Mt. Shasta Arnica

Reference

• Maguire, B. 1943 A monograph of the genus Arnica (Senecioneae, Compositae). Brittonia 4: 386–510
• Wolf, S.J. & K.E. Denford. 1984. Taxonomy of Arnica (Compositae) subgenus Austromontana. Rhodora Journal of the New England Botanical Club 86(847): 239 - 309.
• Nordenstam, B. 1977 Senecioneae and Liabeae—systematic review. In V. H. Heywood, J. B. Harborne, and B. L. Turner [eds.], The biology and chemistry of the Compositae, vol. II, 799–830. Academic Press, London, UK
• Baldwin, B. G. 1999 New combinations in Californian Arnica and Monolopia. Novon 9: 460–461
• Lyss, G., T. J. Schmidt, H. L. Pahl, and I. Merfort. 1999 Anti-inflammatory activity of Arnica tincture (DAB 1998) using the transcription factor NF-kappaB as molecular target. Pharmaceutical and Pharmacological Letters 9: 5–8
• Wolf, S. J., and K. E. Denford. 1984 Taxonomy of Arnica (Compositae) subgenus Austromontana. Rhodora 86: 239–309

External links

• Arnica species
• UVSC Herbarium - Arnica

Gallery

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  Mountain Arnica (Arnica montana)
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  Mountain Arnica (Arnica montana)
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  Longleaf Arnica (Arnica longifolia)