Jump to content

Mallard: Difference between revisions

From Wikipedia, the free encyclopedia
Content deleted Content added
added wikilinks
Line 84: Line 84:
During the breeding season, both male and female mallards can become aggressive, driving off competitors to themselves or their mate by charging at them.<ref>{{Cite book|url=https://books.google.co.in/books?id=c3brsyiAJ0wC&pg=PA66&dq=mallard+breeding+season+male+female+mallards+aggressive&hl=en&sa=X&ved=0ahUKEwj--vzeoLnUAhULuY8KHdOCDqYQ6AEINDAE#v=onepage&q=mallard%20breeding%20season%20male%20female%20mallards%20aggressive&f=false|title=Man and Wildfowl|last=Kear|first=Janet|date=2010-11-30|publisher=Bloomsbury Publishing|isbn=9781408137604|language=en}}</ref> Males tend to fight more than females, and attack each other by repeatedly pecking at their rival's chest, ripping out feathers and even skin on rare occasions.
During the breeding season, both male and female mallards can become aggressive, driving off competitors to themselves or their mate by charging at them.<ref>{{Cite book|url=https://books.google.co.in/books?id=c3brsyiAJ0wC&pg=PA66&dq=mallard+breeding+season+male+female+mallards+aggressive&hl=en&sa=X&ved=0ahUKEwj--vzeoLnUAhULuY8KHdOCDqYQ6AEINDAE#v=onepage&q=mallard%20breeding%20season%20male%20female%20mallards%20aggressive&f=false|title=Man and Wildfowl|last=Kear|first=Janet|date=2010-11-30|publisher=Bloomsbury Publishing|isbn=9781408137604|language=en}}</ref> Males tend to fight more than females, and attack each other by repeatedly pecking at their rival's chest, ripping out feathers and even skin on rare occasions.


The group of drakes, end up being left out, after the others are paired off with mating partners, sometimes targets an isolated female duck, even one of a different species, and proceeds to chase and peck at her until she weakens, at which point the males take turns copulating with the female.<ref name=":10">{{Cite journal|last=Moeliker|first=Cornelis|date=2001|title=The first case of homosexual necrophilia in the mallard Anas platyrhynchos (Aves:Anatidae)|url=http://www.hetnatuurhistorisch.nl/fileadmin/user_upload/documents-nmr/Persberichten/Persberichten/persberichten_2013/DSA8_243-248.pdf|journal=Deinsea 8|volume=|pages=243–248|via=}}</ref> Lebret (1961) calls this behaviour "Attempted Rape Flight" and [[Stanley Cramp]] & K.E.L. Simmons (1977) speak of "rape-intent flights".<ref name=":10" /> Male mallards also occasionally chase other male ducks of a different species, and even each other, in the same way.<ref name=":10" /> In one documented case of "homosexual necrophilia", a male mallard copulated with another male he was chasing after the chased male died upon flying into a glass window.<ref name=Moeliker/> This paper was awarded with an [[Ig Nobel Prize]] in 2003.<ref name=MacLeod/>
The group of drakes, end up being left out, after the others are paired off with mating partners, sometimes targets an isolated female duck, even one of a different species, and proceeds to chase and peck at her until she weakens, at which point the males take turns copulating with the female.<ref name=":10">{{Cite journal|last=Moeliker|first=Cornelis|date=2001|title=The first case of homosexual necrophilia in the mallard Anas platyrhynchos (Aves:Anatidae)|url=http://www.hetnatuurhistorisch.nl/fileadmin/user_upload/documents-nmr/Persberichten/Persberichten/persberichten_2013/DSA8_243-248.pdf|journal=Deinsea 8|volume=|pages=243–248|via=}}</ref> Lebret (1961) calls this behaviour "Attempted Rape Flight" and [[Stanley Cramp]] & K.E.L. Simmons (1977) speak of "rape-intent flights".<ref name=":10" /> Male mallards also occasionally chase other male ducks of a different species, and even each other, in the same way.<ref name=":10" /> In one documented case of "homosexual necrophilia", a male mallard copulated with another male he was chasing after the chased male died upon flying into a glass window.<ref name=Moeliker/> This paper was awarded an [[Ig Nobel Prize]] in 2003.<ref name=MacLeod/>


Mallards are opportunistically targeted by [[brood parasite]]s, occasionally having eggs laid in their nests by [[Redhead (duck)|redheads]], [[ruddy duck]]s, [[lesser scaup]], [[gadwall]]s, [[northern shoveler]]s, [[northern pintail]]s, [[cinnamon teal]], [[common goldeneye]]s, and other mallards.<ref>{{Cite book|url=https://books.google.co.in/books?id=wvpPMwEACAAJ&printsec=frontcover#v=snippet&q=gadwall%20brood%20mallard&f=false|title=Ducks, Geese, and Swans of North America|last=Baldassarre|first=Guy A.|date=2014|publisher=JHU Press|isbn=9781421407517|language=en}}</ref> These eggs are generally accepted when they resemble the eggs of the host mallard, although the hen may attempt to eject them or even abandon the nest if parasitism occurs during egg laying.<ref name=BNA/>
Mallards are opportunistically targeted by [[brood parasite]]s, occasionally having eggs laid in their nests by [[Redhead (duck)|redheads]], [[ruddy duck]]s, [[lesser scaup]], [[gadwall]]s, [[northern shoveler]]s, [[northern pintail]]s, [[cinnamon teal]], [[common goldeneye]]s, and other mallards.<ref>{{Cite book|url=https://books.google.co.in/books?id=wvpPMwEACAAJ&printsec=frontcover#v=snippet&q=gadwall%20brood%20mallard&f=false|title=Ducks, Geese, and Swans of North America|last=Baldassarre|first=Guy A.|date=2014|publisher=JHU Press|isbn=9781421407517|language=en}}</ref> These eggs are generally accepted when they resemble the eggs of the host mallard, although the hen may attempt to eject them or even abandon the nest if parasitism occurs during egg laying.<ref name=BNA/>

Revision as of 04:49, 17 June 2017

Mallard
Female (left) and male (right)
Female call
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Anseriformes
Family: Anatidae
Genus: Anas
Species:
A. platyrhynchos
Binomial name
Anas platyrhynchos
Linnaeus, 1758
Subspecies

A. p. platyrhynchos Linnaeus, 1758
A. p. domesticus Linnaeus, 1758
A. p. conboschas C. L. Brehm, 1831 (disputed)

Global range (native and introduced)
  Year-Round Range
  Summer Range
  Winter Range
Synonyms

Anas boschas Linnaeus, 1758

The mallard (/ˈmælɑːrd/ or /ˈmælərd/) or wild duck (Anas platyrhynchos) is a dabbling duck which breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Argentina, Chile, the Falkland Islands and South Africa.[2] This duck belongs to the subfamily Anatinae of the waterfowl family Anatidae.

Male mallard, Sweden 2016
Fledgling
Juvenile
An American black duck (top left) and a male mallard (bottom right) in eclipse plumage

The male birds (drakes) have a glossy green head and are grey on wings and belly, while the females (hens or ducks) have mainly brown-speckled plumage. Both sexes have an area of white-bordered black speculum feathers which commonly also include iridescent blue feathers especially among males. Mallards live in wetlands, eat water plants and small animals, and are social animals preferring to congregate in groups or flocks of varying sizes. This species is the main ancestor of most breeds of domesticated ducks.[3]

Taxonomy and evolution

Plate 221 of the Birds of America by John James Audubon.

The mallard was one of the many bird species originally described by Carl Linnaeus in his 18th-century work Systema Naturae, and still bears its original binomial name,[4] and in 1758, he had given it the scientific name Anas boschas.[5] The scientific name is from Latin Anas, "duck" and Ancient Greek platyrhynchus, "broad-billed" ( from platus, "broad" and rhunkhos, "bill").[6]

Mallard originally referred to any wild drake and it is sometimes still used this way.[7] It was derived from the Old French malart or mallart for "wild drake", although its true derivation is unclear.[8] It may be related to (or at least influenced by) an Old High German masculine proper name Madelhart, clues lying in the alternate English forms "maudelard" or "mawdelard".[9] Masle (male) has also been proposed as an influence.[10]

Mallards frequently interbreed with their closest relatives in the genus Anas, such as the American black duck, and also with species more distantly related, such as the northern pintail, leading to various hybrids that may be fully fertile.[11] This is quite unusual among such different species, and apparently is because the mallard evolved very rapidly and recently, during the Late Pleistocene.[12] The distinct lineages of this radiation are usually kept separate due to non-overlapping ranges and behavioural cues, but are still not fully genetically incompatible.[13] Mallards and their domesticated conspecifics are also fully interfertile.[14]

The genome of Anas platyrhynchos was sequenced in 2013.[15]

Certain mallards, by their divergent haplotype analysis, appear to be closer to their Indo-Pacific relatives, and certain others, to their American ones.[16] Considering mitochondrial DNA D-loop sequence data, they may have evolved in the general area of Siberia; mallard bones rather abruptly appear in food remains of ancient humans and other deposits of fossil bones in Europe, without a good candidate for a local predecessor species.[17] The large ice age palaeosubspecies which made up at least the European and west Asian populations during the Pleistocene has been named Anas platyrhynchos palaeoboschas.[18]

In their mitochondrial DNA, mallards are differentiated between North America and Eurasia;[19] however, in the nuclear genome there is a particular lack of genetic structure.[20] Haplotypes typical of American mallard relatives and spotbills can be found in mallards around the Bering Sea.[21] The Aleutian Islands hold a population of mallards that appear to be evolving towards a subspecies, as gene flow with other populations is very limited.[17]

The size of the mallard varies clinally, and birds from Greenland, although larger than birds further south, have smaller bills, paler plumage and are stockier.[22] They are sometimes separated as subspecies, the Greenland mallard (A. p. conboschas).[22]

Description

Calls
Iridescent speculum feathers of the male
Owing to their highly 'malleable' genetic code, mallards can display a large amount of variation,[23] as seen here with this female, who displays faded or 'apricot' plumage.

The mallard is a medium-sized waterfowl species although it is often slightly heavier than most other dabbling ducks. It is 50–65 cm (20–26 in) long (of which the body makes up around two-thirds), has a wingspan of 81–98 cm (32–39 in),[24] and weighs 0.72–1.58 kg (1.6–3.5 lb).[25][26] Among standard measurements, the wing chord is 25.7 to 30.6 cm (10.1 to 12.0 in), the bill is 4.4 to 6.1 cm (1.7 to 2.4 in) and the tarsus is 4.1 to 4.8 cm (1.6 to 1.9 in).[27]

The breeding male mallard is unmistakable, with a glossy bottle-green head and white collar which demarcates the head from the purple-tinged brown breast, grey brown wings, and a pale grey belly.[28] The rear of the male is black, with the dark tail having white borders.[29] The bill of the male is a yellowish orange tipped with black, while that of the female is generally darker ranging from black to mottled orange.[30] The female mallard is predominantly mottled with each individual feather showing sharp contrast from buff to very dark brown, a coloration shared by most female dabbling ducks, and has buff cheeks, eyebrow, throat and neck with a darker crown and eye-stripe.[29]

Both male and female mallards have distinct iridescent purple blue speculum feathers edged with white, prominent in flight or at rest, though temporarily shed during the annual summer moult.[31] Upon hatching, the plumage colouring of the duckling is yellow on the underside and face (with streaks by the eyes) and black on the back (with some yellow spots) all the way to the top and back of the head.[32] Its legs and bill are also black.[32] As it nears a month in age, the duckling's plumage will start becoming drab, looking more like the female (though its plumage is more streaked) and its legs will lose their dark grey colouring.[29] Two months after hatching, the fledgling period has ended and the duckling is now a juvenile.[33] Between three and four months of age, the juvenile can finally begin flying, as its wings are fully developed for flight (which can be confirmed by the sight of purple speculum feathers). Its bill will soon lose its dark grey colouring and its sex can finally be distinguished visually by three factors. The bill colouring is yellow in males, black and orange for females.[34] The breast feathers are reddish-brown for males, brown for females.[34] The centre tail feather is curled for males (called a drake feather), straight for females.[34] During the final period of maturity leading up to adulthood (6–10 months of age), the plumage of female juveniles remains the same while the plumage of male juveniles slowly changes to its characteristic colours.[35] This plumage change also applies to adult mallard males when they transition in and out of their non-breeding eclipse plumage at the beginning and the end of the summer moulting period.[35] The adulthood age for mallards is 14 months and the average life expectancy is 3 years, but they can live to twenty.[36] Several species of duck have brown-plumaged females which can be confused with the female mallard.[37] The female gadwall (A. strepera) has an orange-lined bill, white belly, black and white speculum which is seen as a white square on the wings in flight, and is a smaller bird.[29] More similar to the female mallard in North America are the American black duck (A. rubripes), which is notably darker hued in both sexes than the mallard,[38] and the mottled duck (A. fulvigula), which is somewhat darker than the female mallard, with no white edge on the speculum and slightly different bare-part colouration.[38]

In captivity, domestic ducks come in wild-type plumages, white, and other colours.[39] Most of these colour variants are also known in domestic mallards not bred as livestock, but kept as pets, aviary birds, etc., where they are rare but increasing in availability.[39]

A noisy species, the female has a deeper quack stereotypically associated with ducks.[40][41] Male mallards also make a sound which is phonetically similar to that of the female, with a typical quack; although it is a deep and raspy sound which can also sound like breeeeze.[42] When incubating a nest, or when offspring are present, females vocalise differently, making a call which sounds like a truncated version of the usual quack. They will also hiss if the nest or their offspring are threatened or interfered with. When taking off, the wings of a mallard produce a characteristic faint whistling noise.[43]

The mallard is a rare example of both Allen's Rule and Bergmann's Rule in birds.[44] Bergmann's Rule, which states that polar forms tend to be larger than related ones from warmer climates, has numerous examples in birds.[45] Allen's Rule says that appendages like ears tend to be smaller in polar forms to minimize heat loss, and larger in tropical and desert equivalents to facilitate heat diffusion, and that the polar taxa are stockier overall.[46] Examples of this rule in birds are rare, as they lack external ears. However, the bill of ducks is supplied with a few blood vessels to prevent heat loss.[47]

Due to the malleability of the mallard's genetic code, which gives it its vast interbreeding capability, mutations in the genes that decide plumage colour are very common and have resulted in a wide variety of hybrids such as Brewer's duck (mallard × gadwall, Anas strepera).[48]

Distribution and habitat

The mallard is widely distributed across the Northern and Southern Hemispheres; in North America from southern and central Alaska to Mexico, the Hawaiian Islands,[49] across Eurasia,[50] from Iceland[51] and southern Greenland[49] and parts of Morocco (North Africa)[51] in the west, Scandinavia[51] and Britain[51] to the north, and to Siberia,[52] Japan,[53] and South Korea,[53] in the east, south-eastern and south-western Australia[54] and New Zealand[55] in the Southern hemisphere.[24][56][57] It is strongly migratory in the northern parts of its breeding range, and winters farther south.[50] For example, in North America, it winters south to Southern United States and Northern Mexico,[50] but also regularly strays into Central America and the Caribbean between September and May.[58][59]

The mallard inhabits a wide range of habitat and climates, from Arctic tundra to subtropical regions.[60] It is found in both fresh- and salt-water wetlands, including parks, small ponds, rivers, lakes and estuaries, as well as shallow inlets and open sea within sight of the coastline.[61] Water depths of less than 1 metre (3.3 ft) are preferred, birds avoiding areas more than a few metres deep.[62] They are attracted to bodies of water with aquatic vegetation.[41]

Behaviour

Feeding

The mallard is omnivorous and very flexible in its choice of food.[63] Its diet may vary based on several factors, including the stage of the breeding cycle, short-term variations in available food, nutrient availability, and inter and intraspecific competition.[64] The majority of the mallard's diet seems to be made up of gastropods,[65] invertebrates (including beetles, flies, lepidopterans, dragonflies, and caddisflies),[66] crustaceans,[67] worms,[65] many varieties of seeds and plant matter,[65] and roots and tubers.[67] During the breeding season, male birds were recorded to have eaten 37.6% animal matter and 62.4% plant matter, most notably Echinochloa crus-galli, and nonlaying females ate 37.0% animal matter and 63.0% plant matter, while laying females ate 71.9% animal matter and only 28.1% plant matter.[68] Plants generally make up a larger part of the bird's diet, especially during autumn migration and in the winter.[69][70]

It usually feeds by dabbling for plant food or grazing;[71] there are reports of it eating frogs.[71] It usually nests on a river bank, but not always near water. It is highly gregarious outside of the breeding season and forms large flocks, which are known as sords.[72]

Breeding

Egg, Collection Museum Wiesbaden

Mallards usually form pairs (in October and November in the Northern hemisphere) until the female lays eggs at the start of nesting season which is around the beginning of spring.[73] At this time she is left by the male who joins up with other males to await the moulting period which begins in June (in the Northern hemisphere[74]).[75] During the brief time before this, however, the males are still sexually potent and some of them either remain on standby to sire replacement clutches (for female mallards that have lost or abandoned their previous clutch, replacement clutch[76]) or forcibly mate with females that appear to be isolated or unattached regardless of their species and whether or not they have a brood of ducklings.[76][77]

Duckling, one to two days old, is fully capable of swimming

The nesting period can be very stressful for the female since she lays more than half her body weight in eggs.[78] She requires a lot of rest and a feeding/loafing area that is safe from predators. When seeking out a suitable nesting site, the female's preferences are areas that are well concealed, inaccessible to ground predators, or have few predators nearby. This can include nesting sites in urban areas such as roof gardens, enclosed courtyards, and flower boxes on window ledges and balconies more than one story up, which the ducklings cannot leave safely without human intervention. The clutch is 8–13 eggs, which are incubated for 27–28 days to hatching with 50–60 days to fledging.[79][80] The ducklings are precocial and fully capable of swimming as soon as they hatch.[81] However, filial imprinting compels them to instinctively stay near the mother not only for warmth and protection but also to learn about and remember their habitat as well as how and where to forage for food.[82] When ducklings mature into flight-capable juveniles, they learn about and remember their traditional migratory routes (unless they are born and raised in captivity). They may stay with their family group for up to a year, despite being independent and no longer needing protection.[83]

During the breeding season, both male and female mallards can become aggressive, driving off competitors to themselves or their mate by charging at them.[84] Males tend to fight more than females, and attack each other by repeatedly pecking at their rival's chest, ripping out feathers and even skin on rare occasions.

The group of drakes, end up being left out, after the others are paired off with mating partners, sometimes targets an isolated female duck, even one of a different species, and proceeds to chase and peck at her until she weakens, at which point the males take turns copulating with the female.[85] Lebret (1961) calls this behaviour "Attempted Rape Flight" and Stanley Cramp & K.E.L. Simmons (1977) speak of "rape-intent flights".[85] Male mallards also occasionally chase other male ducks of a different species, and even each other, in the same way.[85] In one documented case of "homosexual necrophilia", a male mallard copulated with another male he was chasing after the chased male died upon flying into a glass window.[86] This paper was awarded an Ig Nobel Prize in 2003.[87]

Mallards are opportunistically targeted by brood parasites, occasionally having eggs laid in their nests by redheads, ruddy ducks, lesser scaup, gadwalls, northern shovelers, northern pintails, cinnamon teal, common goldeneyes, and other mallards.[88] These eggs are generally accepted when they resemble the eggs of the host mallard, although the hen may attempt to eject them or even abandon the nest if parasitism occurs during egg laying.[89]

Predators and threats

Mallards of all ages (but especially young ones) and in all locations must contend with a wide diversity of predators including raptors, mustelids, corvids, snakes, raccoons, opossums, skunks, turtles, large fish and felids and canids, including domesticated ones.[90] The most prolific natural predators of adult mallards are red foxes (which most often pick off brooding females) and the faster or larger birds of prey, i.e. peregrine falcons, Aquila eagles or Haliaeetus eagles.[91][92] In North America, adult mallards face no fewer than 15 species of birds of prey, from hen harriers and short-eared owls (both smaller than a mallard) to huge bald and golden eagles, and about a dozen species of mammalian predator, not counting several more avian and mammalian predators who threaten eggs and nestlings.[89]

Mallards are also preyed upon by other waterside apex predators, such as the grey heron (Ardea cinerea),[93] European herring gull (Larus argentatus), the Wels catfish (Silurus glanis) and the Northern pike (Esox lucius).[94] Crows (Corvus sp.) are also known to kill ducklings and adults on occasion.[95] Also, mallards may be attacked by larger anseriformes such as swans (Cygnus sp.) and geese during the breeding season, and are frequently driven off by these birds over territorial disputes. Mute swans (C. olor) have been known to attack mallards if they feel that the ducks pose a threat to their offspring.[96]

Conservation

By Carl Friedrich Deiker (1875)
Several drakes swim in a pond

Unlike many waterfowl, mallards have benefited from human alterations to the world – so much so that they are now considered an invasive species in some regions.[97]

They are a common sight in urban parks, lakes, ponds, and other manmade water features in the regions they inhabit, and are often tolerated or encouraged among human habitat due to their placid nature towards humans and their beautiful and iridescent colours.[31] While most are not domesticated, mallards are so successful at coexisting in human regions that the main conservation risk they pose comes from the loss of genetic diversity among a region's traditional ducks once humans and mallards colonize an area. Mallards are very adaptable, being able to live and even thrive in urban areas which may have supported more localized, sensitive species of waterfowl before development.[98] The release of feral mallards in areas where they are not native sometimes creates problems through interbreeding with indigenous waterfowl.[97][99] These non-migratory mallards interbreed with indigenous wild ducks from local populations of closely related species through genetic pollution by producing fertile offspring.[99] Complete hybridization of various species of wild ducks gene pools could result in the extinction of many indigenous waterfowl.[99] The wild mallard itself is the ancestor of most domestic ducks and its naturally evolved wild gene pool gets genetically polluted in turn by the domesticated and feral populations.[100][101][102]

Over time, a continuum of hybrids ranging between almost typical examples of either species will develop; the speciation process beginning to reverse itself.[103] This has created conservation concerns for relatives of the mallard, such as the Hawaiian duck,[104][105] the A. s. superciliosa subspecies of the Pacific black duck,[104][106] the American black duck,[107][108] the mottled duck,[109][110] Meller's duck,[111] the yellow-billed duck,[103] and the Mexican duck,[104][110] in the latter case even leading to a dispute whether these birds should be considered a species[112] (and thus entitled to more conservation research and funding) or included in the mallard. In the cases mentioned below and above, however, ecological changes and hunting have led to a decline of local species; for example, the New Zealand grey duck population declined drastically due to overhunting in the mid-20th century.[106] Hybrid offspring of Hawaiian ducks seem to be less well-adapted to native habitat, and utilizing them in reintroduction projects apparently reduces success.[104][113] In summary, the problems of mallards "hybridizing away" relatives is more a consequence of local ducks declining than of mallards spreading; allopatric speciation and isolating behaviour have produced today's diversity of mallard-like ducks despite the fact that in most, if not all, of these populations, hybridization must have occurred to some extent.[114]

Regions considered invasive

The last male Mariana mallard
Mallard ducklings following their mother near Boston Harbor, USA

The mallard is considered an invasive species in New Zealand,[24] where it competes with the local New Zealand grey duck, which was overhunted in the past.[115] There, and elsewhere, mallards are spreading with increasing urbanisation and hybridizing with local relatives.[104]

Mallards are also causing severe "genetic pollution" to South Africa's biodiversity by breeding with endemic ducks;[116] although, the Agreement on the Conservation of African-Eurasian Migratory Waterbirds applies to the mallard.[117] The hybrids of mallards and the yellow-billed duck are fertile, capable of producing hybrid offspring.[118] If this continues, only hybrids will occur and in the long term, which will result in the extinction of various indigenous waterfowl.[118][119] The mallard duck can cross breed with 63 other species, posing a severe threat to indigenous waterfowl's genetic integrity.[120] Mallards and their hybrids compete with indigenous birds for resources including nest sites, roosting sites and food.[102]

The availability of mallards, mallard ducklings, and fertilized mallard eggs for public sale and private ownership, either as livestock or as pets, is currently legal in the United States except for the state of Florida, which has currently banned domestic ownership of mallards. This is to prevent hybridisation with the native mottled duck.[121]

The Eastern or Chinese spot-billed duck is currently introgressing into the mallard populations of the Primorsky Krai, possibly due to habitat changes from global warming.[21]

Pacific Islands

The Mariana mallard was a resident allopatric population—in most respects a good species—apparently initially derived from mallard-Pacific black duck hybrids;[122] unfortunately, it became extinct in the late twentieth century.[123]

The Laysan duck is an insular relative of the mallard, with a very small and fluctuating population.[124][125] Mallards sometimes arrive on its island home during migration, and can be expected to occasionally have remained and hybridized with Laysan ducks as long as these species have existed.[126] However, these hybrids are less well adapted to the peculiar ecological conditions of Laysan Island than the local ducks, and thus have lower fitness. These ducks were found throughout the Hawaiian archipelago before 400 AD, after which they suffered a rapid decline during the Polynesian colonization.[127] Now, their range includes only the Laysan Island.[127] It is one of the successfully translocated birds, after having been nearly extinct in the early 20th century.[125]

Relationship with humans

As noted, mallards have had a long and nearly symbiotic relationship with humans. Humans create an urban adaptation of traditional waterways to which mallards are better suited than native ducks, giving them an evolutionary advantage, and have frequently domesticated mallards both as pets and a food species.[128] Mallards have often been ubiquitous in their regions among the ponds, rivers, and streams of human parks, farms, and other man-made waterways – even to the point of visiting water features in human courtyards.[129]

The mallard is depicted in a marginal decoration of the 15th century English illuminated manuscript the Sherborne Missal.[130] Since 1933, the Peabody Hotel in Downtown Memphis, Tennessee has maintained a long tradition of keeping one mallard drake and four mallard hens, called The Peabody Ducks, as a popular hotel attraction and as guests of honour.[131][132] The mallards are provided by a local farmer and friend of the Peabody Hotel and are rotated out and returned to the farm for a new team of mallards every three months.[131] This tradition has also been maintained and observed at the other Peabody Hotels in Little Rock, Arkansas and Orlando, Florida.[133] The children's picture book Make Way for Ducklings, published in 1941 and winner of the 1942 Caldecott Medal for its illustrations,[134] is the story of a pair of mallards who decide to raise their family on an island in the lagoon in Boston Public Garden in Massachusetts.[135] Duck Head, a U.S. clothing brand, uses the image of a mallard's head as its logo.[136]

References

  1. ^ Template:IUCN
  2. ^ Long, John L. (1981). Introduced Birds of the World. Agricultural Protection Board of Western Australia. pp. 21–493.
  3. ^ "Anas platyrhynchos (On-line), Digital Morphology". The University of Texas at Austin. 2004. Retrieved 23 August 2012.
  4. ^ Linnaeus, Carl (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata (in Latin). Stockholm: Laurentius Salvius. p. 125.
  5. ^ Jobling, James A. (30 June 2010). Helm Dictionary of Scientific Bird Names. Bloomsbury Publishing. ISBN 9781408133262.
  6. ^ Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 46, 309. ISBN 978-1-4081-2501-4.
  7. ^ Magnus, PD (2012), Scientific Enquiry and Natural Kinds: from Planets to Mallards, Palgrave Macmillan, ISBN 9781137271259
  8. ^ Ltd, Allied Newspapers. "Ducks of a common feather". Times of Malta. {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  9. ^ "mallard". Oxford English Dictionary (2nd ed.). Oxford University Press. 1989.
  10. ^ "mallard". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
  11. ^ Phillips, John C. (1915). "Experimental studies of hybridization among ducks and pheasants". Journal of Experimental Zoology. 18 (1): 69–112. doi:10.1002/jez.1400180103.
  12. ^ Steadman, David (May 2016). "LATE PLEISTOCENE BIRDS FROM KINGSTON SALTPETER CAVE, SOUTHERN APPALACHIAN MOUNTAINS, GEORGIA" (PDF). Florida Museum of Natural History: 231–248.
  13. ^ Kraus, R.H.S.; Kerstens, H.H.D.; van Hooft, P.; Megens, H.-J.; Elmberg, J.; Tsvey, Arseny; Sartakov, Dmitry; Soloviev, Sergej A.; Crooijmans, Richard P.M.A.; Groenen, Martien A.M.; Ydenberg, Ronald C.; Prins, Herbert H.T. (2012). "Widespread horizontal genomic exchange does not erode species barriers among sympatric ducks". BMC Evolutionary Biology. 12 (45): 45. doi:10.1186/1471-2148-12-45.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  14. ^ "Bird Tawk by Tina Mitchell | Greenwood Wildlife Rehabilitation Center". www.greenwoodwildlife.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  15. ^ Huang, Y.; Li, Y.; Burt, D.W.; Chen, H.; Zhang, Y.; Qian, Wubin; Kim, Heebal; Gan, Shangquan; Zhao, Yiqiang; Li, Jianwen; Yi, Kang; Feng, Huapeng; Zhu, Pengyang; Li, Bo; Liu, Qiuyue; Fairley, Suan; Magor, Katharine E; Du, Zhenlin; Hu, Xiaoxiang; Goodman, Laurie; Tafer, Hakim; Vignal, Alain; Lee, Taeheon; Kim, Kyu-Won; Sheng, Zheya; An, Yang; Searle, Steve; Herrero, Javier; Groenen, Martien A.M.; et al. (2013). "The duck genome and transcriptome provide insight into an avian influenza virus reservoir species". Nature Genetics. 45 (7). Nature Publishing: 776–783. doi:10.1038/ng.2657. PMC 4003391. PMID 23749191.
  16. ^ Johnson, Kevin P.; Sorenson, M.D. (1999). "Phylogeny and biogeography of dabbling ducks (genus Anas): a comparison of molecular and morphological evidence" (PDF). The Auk. 116 (3): 792–805. doi:10.2307/4089339.
  17. ^ a b Kulikova, Irina V.; Drovetski, S.V.; Gibson, D.D.; Harrigan, R.J.; Rohwer, S.; Sorenson, Michael D.; Winker, K.; Zhuravlev, Yury N.; McCracken, Kevin G. (2005). "Phylogeography of the Mallard (Anas platyrhynchos): hybridization, dispersal, and lineage sorting contribute to complex geographic structure". The Auk. 122 (3): 949–965. doi:10.1642/0004-8038(2005)122[0949:POTMAP]2.0.CO;2. (Erratum: The Auk 122 (4): 1309, doi:10.1642/0004-8038(2005)122[1309:POTMAP2.0.CO;2].)
  18. ^ Delacour, Jean (1964). The Waterfowl of the World. Country Life.
  19. ^ Kraus, R.H.S.; Zeddeman, A.; van Hooft, P.; Sartakov, D.; Soloviev, S.A.; Ydenberg, Ronald C.; Prins, Herbert H.T. (2011). "Evolution and connectivity in the world-wide migration system of the mallard: Inferences from mitochondrial DNA". BMC Genetics. 12 (99): 99. doi:10.1186/1471-2156-12-99. PMC 3258206. PMID 22093799.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  20. ^ Kraus, R.H.S.; van Hooft, P.; Megens, H.-J.; Tsvey, A.; Fokin, S.Y.; Ydenberg, Ronald C.; Prins, Herbert H.T. (2013). "Global lack of flyway structure in a cosmopolitan bird revealed by a genome wide survey of single nucleotide polymorphisms". Molecular Ecology. 22 (1) (published January 2013): 41–55. doi:10.1111/mec.12098. PMID 23110616.
  21. ^ a b Kulikova, Irina V.; Zhuravlev, Yury N.; McCracken, Kevin G. (2004). "Asymmetric hybridization and sex-biased gene flow between Eastern Spot-billed Ducks (Anas zonorhyncha) and Mallards (A. platyrhynchos) in the Russian Far East". The Auk. 121 (3): 930–949. doi:10.1642/0004-8038(2004)121[0930:AHASGF]2.0.CO;2.
  22. ^ a b Ogilvie, M. A.; Young, Steve (2002). Wildfowl of the World. New Holland Publishers. ISBN 9781843303282.
  23. ^ "Everglades News | Mallards". American Bird. 3 June 2015. {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  24. ^ a b c Cramp 1977, p. 505
  25. ^ "Mallard". All About Birds. Cornell Lab of Ornithology.
  26. ^ Dunning, John B. Jr., ed. (1992). CRC Handbook of Avian Body Masses. CRC Press. ISBN 978-0-8493-4258-5.
  27. ^ Madge, Steve (1992). Waterfowl: An Identification Guide to the Ducks, Geese, and Swans of the World. Houghton Mifflin Harcourt. ISBN 978-0-395-46726-8.
  28. ^ Ogilvie, M. A.; Young, Steve (2002). Wildfowl of the World. New Holland Publishers. ISBN 9781843303282.
  29. ^ a b c d Cramp 1977, p. 506
  30. ^ Jiguet, Frédéric; Audevard, Aurélien (21 March 2017). Birds of Europe, North Africa, and the Middle East: A Photographic Guide. Princeton University Press. ISBN 9780691172439.
  31. ^ a b Fergus, Charles; Hansen, Amelia (2000). Wildlife of Pennsylvania and the Northeast. Stackpole Books. ISBN 9780811728997.
  32. ^ a b Lancaster, John Frederick (17 December 2013). The inheritance of plumage colour in the common duck (Anas platyrhynchos linné). Springer. ISBN 9789401768344.
  33. ^ Station, Delta Waterfowl and Wetlands Research (1984). Annual Report. The Station.
  34. ^ a b c Moulton, Judy (7 November 2014). Daisy and Ducky Mallard. Xlibris Corporation. ISBN 9781503511910.
  35. ^ a b Vinicombe, Keith (27 March 2014). The Helm Guide to Bird Identification. Bloomsbury Publishing. ISBN 9781472905543.
  36. ^ Robinson, R.A. (2005). "Mallard Anas platyrhynchos". BirdFacts: profiles of birds occurring in Britain & Ireland (BTO Research Report 407). Thetford: BTO. Retrieved 10 February 2015.
  37. ^ Moss, Stephen; Cottridge, David (2000). Attracting Birds to Your Garden. New Holland Publishers. ISBN 9781859740057.
  38. ^ a b Kaufman, Kenn (2005). Kaufman Field Guide to Birds of North America. Houghton Mifflin Harcourt. ISBN 0618574239.
  39. ^ a b Hicks, James Stephen (1923). The Encyclopaedia of Poultry. Waverley Book Company.
  40. ^ Rogers, D. (2001). "Anas platyrhynchos (On-line)". Animal Diversity Web. Museum of Zoology, University of Michigan. Retrieved 8 December 2006.
  41. ^ a b Cramp 1977, p. 507
  42. ^ Carver, Heather (2011). The Duck Bible. Lulu.com. ISBN 9780557901562.
  43. ^ "Mallard description". science.nature.nps.gov. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  44. ^ Salewski, Volker; Hochachka, Wesley M.; Fiedler, Wolfgang (September 2009). "Global warming and Bergmann's rule: do central European passerines adjust their body size to rising temperatures?". Oecologia. 162 (1): 247–260. doi:10.1007/s00442-009-1446-2. ISSN 0029-8549. PMC 2776161. PMID 19722109.
  45. ^ Shelomi, Matan; Zeuss, Dirk (2017). "Bergmann's and Allen's Rules in Native European and Mediterranean Phasmatodea". Frontiers in Ecology and Evolution. 5. doi:10.3389/fevo.2017.00025. ISSN 2296-701X.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  46. ^ Bidau, Claudio J.; Martí, Dardo A. (August 2008). "A test of Allen's rule in ectotherms: the case of two south American Melanopline Grasshoppers (Orthoptera: Acrididae) with partially overlapping geographic ranges". Neotropical Entomology. 37 (4): 370–380. doi:10.1590/S1519-566X2008000400004. ISSN 1519-566X.
  47. ^ Ducks Unlimited Magazine. Vol. 67–68. Ducks Unlimited, Incorporated. 2003. p. 62.{{cite book}}: CS1 maint: date and year (link)
  48. ^ "Brewer's Duck". audubon.org. National Audubon Society. Retrieved 19 May 2012.
  49. ^ a b Madge, Steve (30 September 2010). Wildfowl. A&C Black. ISBN 9781408138953.
  50. ^ a b c "Mallard". www.allaboutbirds.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  51. ^ a b c d Finlayson, Clive (30 September 2010). Birds of the Strait of Gibraltar. Bloomsbury Publishing. ISBN 9781408136942.
  52. ^ Prins, Herbert H. T.; Namgail, Tsewang (6 April 2017). Bird Migration across the Himalayas: Wetland Functioning amidst Mountains and Glaciers. Cambridge University Press. ISBN 9781107114715.
  53. ^ a b Yamaguchi, Noriyuki; Hiraoka, Emiko; Fujita, Masaki; Hijikata, Naoya; Ueta, Mutsuyuki; Takagi, Kentaro; Konno, Satoshi; Okuyama, Miwa; Watanabe, Yuki (September 2008). "Spring migration routes of mallards (Anas platyrhynchos) that winter in Japan, determined from satellite telemetry". Zoological Science. 25 (9): 875–881. doi:10.2108/zsj.25.875. ISSN 0289-0003. PMID 19267595.
  54. ^ "Mallard (Anas platyrhynchos)". www.ozanimals.com. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  55. ^ "Northern Mallard | BIRDS in BACKYARDS". www.birdsinbackyards.net. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  56. ^ "Mallard (Anas platyrhynchos) – BirdLife species factsheet". datazone.birdlife.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  57. ^ "Anas platyrhynchos (Common Mallard, Mallard, Northern Mallard)". www.iucnredlist.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  58. ^ Herrera, Néstor; Rivera, Roberto; Ibarra Portillo, Ricardo; Rodríguez, Wilfredo (2006). "Nuevos registros para la avifauna de El Salvador" (PDF). Boletín de la Sociedad Antioqueña de Ornitología (in Spanish). 16 (2): 1–19. {{cite journal}}: Unknown parameter |trans_title= ignored (|trans-title= suggested) (help)
  59. ^ "Mallard: Waterfowl ID". www.ducks.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  60. ^ "Mallard (Anas platyrhynchos)". www.yerevanzoo.am. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  61. ^ Burton, Maurice; Burton, Robert (2002). International Wildlife Encyclopedia: Leopard – marten. Marshall Cavendish. ISBN 9780761472773.
  62. ^ "Mallard Duck • Elmwood Park Zoo | Elmwood Park Zoo | www.elmwoodparkzoo.org". www.elmwoodparkzoo.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  63. ^ van Toor, Mariëlle L.; Hedenström, Anders; Waldenström, Jonas; Fiedler, Wolfgang; Holland, Richard A.; Thorup, Kasper; Wikelski, Martin (30 August 2013). "Flexibility of Continental Navigation and Migration in European Mallards". PLoS ONE. 8 (8). doi:10.1371/journal.pone.0072629. ISSN 1932-6203. PMC 3758317. PMID 24023629.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  64. ^ Krapu, Gary L.; Reinecke, Kenneth J. (1992). "Foraging ecology and nutrition". In Batt, Bruce D.J.; Afton, Alan D.; Anderson, Michael G.; Ankney, C. Davison; Johnson, Douglas H.; Kadlec, John A.; Krapu, Gary L. (eds.). Ecology and Management of Breeding Waterfowl. Minneapolis: University of Minnesota Press. pp. 1–30 (10). ISBN 978-0-8166-2001-2.
  65. ^ a b c Baldassarre, Guy A. (2014). Ducks, Geese, and Swans of North America. JHU Press. ISBN 9781421407517.
  66. ^ Eldridge, Jan (1990). "Waterfowl Management Handbook" (PDF). Fish and Wildlife Leaflet.
  67. ^ a b Rappole, John H. (6 February 2012). Wildlife of the Mid-Atlantic: A Complete Reference Manual. University of Pennsylvania Press. ISBN 0812222016.
  68. ^ Swanson, George A.; Meyer, Mavis I.; Adomaitis, Vyto A. (1985). "Foods consumed by breeding mallards on wetlands of south-central North Dakota". Journal of Wildlife Management. 49 (1): 197–203. doi:10.2307/3801871. JSTOR 3801871.
  69. ^ Gruenhagen, Ned M.; Fredrickson, Leigh H. (1990). "Food use by migratory female mallards in northwest Missouri". Journal of Wildlife Management. 54 (4): 622–626. doi:10.2307/3809359. JSTOR 3809359.
  70. ^ Combs, Daniel L.; Fredrickson, Leigh H. (1990). "Foods used by male mallards wintering in southeastern Missouri". Journal of Wildlife Management. 60 (3): 603–610. doi:10.2307/3802078. JSTOR 3802078.
  71. ^ a b Sandilands, Al (1 January 2011). Birds of Ontario: Habitat Requirements, Limiting Factors, and Status: Volume 1–Nonpasserines: Loons through Cranes. UBC Press. ISBN 9780774859431.
  72. ^ Ross, Terry. "Group Name for Birds: A Partial List". Baltimore Bird Club. Retrieved 3 June 2007.
  73. ^ The Encyclopedia of Birds. Parragon. 2005. ISBN 9781405498517.
  74. ^ Ginn, H. B.; Melville, Dorothy Sutherland (1983). Moult in birds. British Trust for Ornithology.
  75. ^ Boere, G. C.; Galbraith, Colin A.; Stroud, David A. (2006). Waterbirds Around the World: A Global Overview of the Conservation, Management and Research of the World's Waterbird Flyways. The Stationery Office. ISBN 9780114973339.
  76. ^ a b Boere, G. C.; Galbraith, Colin A.; Stroud, David A. (2006). Waterbirds Around the World: A Global Overview of the Conservation, Management and Research of the World's Waterbird Flyways. The Stationery Office. ISBN 9780114973339.
  77. ^ Feinstein, Julie (2011). Field Guide to Urban Wildlife. Stackpole Books. ISBN 9780811705851.
  78. ^ Dekker, Dick (1 January 1980). Naturalist Painter: An Artist's Observations of Western Wildlife. Western Producer Prairie Books. ISBN 9780888330659.
  79. ^ Burton, Maurice; Burton, Robert (2002). International Wildlife Encyclopedia: Leopard – marten. Marshall Cavendish. ISBN 9780761472773. Retrieved 16 June 2016.
  80. ^ DK; International, BirdLife (1 March 2011). The Illustrated Encyclopedia of Birds. Dorling Kindersley Limited. ISBN 9781405336161.
  81. ^ Al-Obaidi, Faris; Al-Shadeedi, Shahrazad (July 2016). "Comparison Study of Egg Morphology, Component and Chemical Composition of Mallard Duck and Domestic Peking Duck" (PDF). The Journal of Bio Innovation: 555–562.
  82. ^ Townsley, Frank (10 March 2016). British Columbia: Graced by Nature's Palette. FriesenPress. ISBN 9781460277737.
  83. ^ "Pictures of Mallards". The Spruce. {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  84. ^ Kear, Janet (30 November 2010). Man and Wildfowl. Bloomsbury Publishing. ISBN 9781408137604.
  85. ^ a b c Moeliker, Cornelis (2001). "The first case of homosexual necrophilia in the mallard Anas platyrhynchos (Aves:Anatidae)" (PDF). Deinsea 8: 243–248.
  86. ^ Moeliker, C. W. (2001). "The first case of homosexual necrophilia in the Mallard Anas platyrhynchos (Aves: Anatidae)" (PDF). Deinsea. 8 (243–247).
  87. ^ MacLeod, Donald (8 March 2005). "Necrophilia among ducks ruffles research feathers". The Guardian. Retrieved 9 December 2006.
  88. ^ Baldassarre, Guy A. (2014). Ducks, Geese, and Swans of North America. JHU Press. ISBN 9781421407517.
  89. ^ a b Drilling, Nancy; Titman, Roger; McKinney, Frank (2002). Poole, A. (ed.). "Mallard (Anas platyrhynchos)". The Birds of North America Online. Ithaca: Cornell Lab of Ornithology. doi:10.2173/bna.658. Retrieved 1 February 2011.
  90. ^ "Anas platyrhynchos". US Forest Service. Retrieved 23 August 2012.
  91. ^ "Impact of Red Fox Predation on the Sex Ratio of Prairie Mallards". USGS Northern Prairie Wildlife Research Center. 3 August 2006. Retrieved 23 August 2012.
  92. ^ "Geese Ducks and Swans: Anatidae – Mallard (Anas Platyrhynchos): Species Accounts – Mallards, Eat, Male, and Female". JRank. Retrieved 7 September 2014.
  93. ^ Marquiss, M.; Leitch, A. F. (1 October 1990). "The diet of Grey Herons Ardea cinerea breeding at Loch Leven, Scotland, and the importance of their predation on ducklings". Ibis. 132 (4): 535–549. doi:10.1111/j.1474-919X.1990.tb00277.x. ISSN 1474-919X.
  94. ^ Reader's Digest Scenic wonders of Canada: an illustrated guide to our natural splendors. Reader's Digest Association (Canada). 1976. ISBN 9780888500496.
  95. ^ Adams, Mary (October 1995). Ecosystem Matters: Activity and Resource Guide for Environmental Educators. DIANE Publishing. ISBN 9780788124532.
  96. ^ Fray, Rob; Davies, Roger; Gamble, Dave; Harrop, Andrew; Lister, Steve (30 June 2010). The Birds of Leicestershire and Rutland. Bloomsbury Publishing. ISBN 9781408133118.
  97. ^ a b Mooney, H. A.; Cleland, E. E. (8 May 2001). "The evolutionary impact of invasive species". Proceedings of the National Academy of Sciences. 98 (10): 5446–5451. doi:10.1073/pnas.091093398. ISSN 0027-8424. PMC 33232. PMID 11344292.
  98. ^ Leedy, Daniel L.; Adams, Lowell W. (1984). A Guide to Urban Wildlife Management. National Institute for Urban Wildlife.
  99. ^ a b c Uyehara, Kimberly; Engilis, Andrew; Reynolds, Michelle. Hendley, James (ed.). "Hawaiian Duck's Future Threatened by Feral Mallards" (PDF). {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  100. ^ "Mottled Ducks : The Problem : Hybridization; Florida Fish and Wildlife Conservation Commission". Florida Fish and Wildlife Conservation Commission. Archived from the original on 20 April 2006. Retrieved 23 August 2012.[failed verification]
  101. ^ Bowers, Frank (May 2002). "Environmental assessment for control of free-ranging resident mallards in Florida". US Fish and Wildlife Service. Retrieved 23 August 2012.
  102. ^ a b "Invasive Alien Bird Species Pose A Threat, Kruger National Park, Siyabona Africa Travel (Pty) Ltd – South Africa Safari Travel Specialist". Krugerpark.co.za. Retrieved 23 August 2012.
  103. ^ a b Rhymer, Judith M. (2006). "Extinction by hybridization and introgression in anatine ducks" (PDF). Acta Zoologica Sinica. 52 (Supplement): 583–585.
  104. ^ a b c d e Rhymer, Judith M.; Simberloff, Daniel (1996). "Extinction by hybridization and introgression". Annual Review of Ecology and Systematics. 27: 83–109. doi:10.1146/annurev.ecolsys.27.1.83.
  105. ^ Griffin, C.R.; Shallenberger, F.J.; Fefer, S.I. (1989). "Hawaii's endangered waterbirds: a resource management challenge". In Sharitz, R.R.; Gibbons, I.W. (eds.). Proceedings of Freshwater Wetlands and Wildlife Symposium. Aiken, South Carolina: Savannah River Ecology Lab. pp. 155–169.
  106. ^ a b Williams, Murray; Basse, Britta (2006). "Indigenous gray ducks, Anas superciliosa, and introduced mallards, A. platyrhynchos, in New Zealand: processes and outcome of a deliberate encounter" (PDF). Acta Zoologica Sinica. 52 (Supplement): 579–582.
  107. ^ Avise, John C.; Ankney, C. Davison; Nelson, William S. (1990). "Mitochondrial gene trees and the evolutionary relationship of Mallard and Black Ducks" (PDF). Evolution. 44 (4): 1109–1119. doi:10.2307/2409570.
  108. ^ Mank, Judith E.; Carlson, John E.; Brittingham, Margaret C. (2004). "A century of hybridization: decreasing genetic distance between American black ducks and mallards". Conservation Genetics. 5 (3): 395–403. doi:10.1023/B:COGE.0000031139.55389.b1.
  109. ^ Mazourek, J.C.; Gray, P.N. (1994). "The Florida duck or the mallard?". Florida Wildlife. 48 (3): 29–31. Archived from the original (DOC) on 10 August 2007. {{cite journal}}: Unknown parameter |deadurl= ignored (|url-status= suggested) (help)
  110. ^ a b McCracken, Kevin G.; Johnson, William P.; Sheldon, Frederick H. (2001). "Molecular population genetics, phylogeography, and conservation biology of the mottled duck (Anas fulvigula)". Conservation Genetics. 2 (2): 87–102. doi:10.1023/A:1011858312115.
  111. ^ Young, H. Glyn; Rhymer, Judith M. (1998). "Meller's duck: A threatened species receives recognition at last". Biodiversity and Conservation. 7 (10): 1313–1323. doi:10.1023/A:1008843815676.
  112. ^ American Ornithologists' Union (1983). Check-list of North American Birds (6th ed.). Washington, DC: American Ornithologists' Union.
  113. ^ Kirby, Ronald E.; Sargeant, Glen A.; Shutler, Dave (2004). "Haldane's rule and American black duck × mallard hybridization". Canadian Journal of Zoology. 82 (11): 1827–1831. doi:10.1139/z04-169.
  114. ^ Tubaro, Pablo L.; Lijtmaer, Dario A. (1 October 2002). "Hybridization patterns and the evolution of reproductive isolation in ducks". Biological Journal of the Linnean Society. 77 (2): 193–200. doi:10.1046/j.1095-8312.2002.00096.x. ISSN 0024-4066.
  115. ^ Woodhouse, Graeme. "TerraNature | New Zealand ecology – Grey duck (Anas superciliosa superciliosa)". www.terranature.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  116. ^ "Kruger Park Times | Alien Bird Species Pose A Threat | Online News Publication..." www.krugerpark.co.za. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  117. ^ "AGREEMENT on the Conservation of African-Eurasian Migratory Waterbirds" (PDF). Official Journal of the European Union. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  118. ^ a b "Marina da Gama". www.mdga.co.za. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  119. ^ "Waterfowl Hybrids". www.ducks.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  120. ^ "Those Mighty Mallards Can Bust the Speed Limit". San Quentin News. 1 June 2010. {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  121. ^ "Mallard Possession Rule". Florida Fish and Wildlife Conservation Commission. Retrieved 10 February 2015.
  122. ^ Yamashina, Y. (1948). "Notes on the Marianas mallard". Pacific Science. 2: 121–124.
  123. ^ Madge, Steve; Burn, Hilary (2010) [1989]. Wildfowl. London: Christopher Helm Publ. p. 211. ISBN 1408138956.
  124. ^ BROWNE, ROBERT; GRIFFIN, CURTICE; CHANG, PAUL; HUBLEY, MARK; MARTIN, AMY (1993). "GENETIC DIVERGENCE AMONG POPULATIONS OF THE HAWAIIAN DUCK, LAYSAN DUCK, AND MALLARD". The Auk. 110 (1). American Ornithological Society: 49–56. JSTOR 4088230.
  125. ^ a b "Anas laysanensis (Laysan Duck, Laysan Teal)". www.iucnredlist.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  126. ^ "Recovery Strategy – Laysan Duck Revised Recovery Plan". www.fws.gov. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  127. ^ a b "Anas laysanensis (Laysan duck)". Animal Diversity Web. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  128. ^ "Anas platyrhynchos, Domestic Duck; DigiMorph Staff – The University of Texas at Austin". Digimorph.org. Retrieved 23 December 2012.
  129. ^ Channel Improvements, Columbia and Lower Willamette River Federal Navigation Channel, (OR,WA): Environmental Impact Statement. 1999.
  130. ^ Clark, Kenneth (1977). Animals and Men. London: Thames and Hudson. p. 107. ISBN 0-500-23257-1.
  131. ^ a b "Peabody Hotel Ducks | The Peabody Memphis | Memphis, Tennessee". www.peabodymemphis.com. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  132. ^ Chozick, Amy (24 December 2013). "Hotel Ducks Gone Wild May End Up in Hunters' Sights". The New York Times. ISSN 0362-4331. {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  133. ^ "Peabody Ducks". The Peabody Memphis Tennessee Hotels. Retrieved 19 May 2012.
  134. ^ "Make Way For Celebration: These Ducklings Are Turning 75". NPR.org. {{cite web}}: Cite has empty unknown parameter: |dead-url= (help)
  135. ^ "Make Way for Ducklings | Boston Discovery Guide". {{cite news}}: Cite has empty unknown parameter: |dead-url= (help)
  136. ^ "Our Story". Duck Head International LLC. Retrieved 15 February 2015.

Bibliography