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Cretacladoides

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Cretacladoides
Temporal range: Valanginian
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Chondrichthyes
Genus: Cretacladoides
Feichtinger et al. 2018
Type species
Cretacladoides ogiveformis
Feichtinger et al., 2018
Species
  • C. ogiveformis (Feichtinger et al., 2018)
  • C. noricum (Feichtinger et al., 2018)

Cretacladoides was a possible genus of falcatid chondrichthyan found in France and Austria. It consists of two species, C. ogiveformis and C. noricum.[1] Assuming a falcatid identity, it is the most recent member of the family, which otherwise died out at the end of the Carboniferous.[1]

History of Discovery

The teeth of Cretacladoides were discovered on a field trip by Alexander Lukeneder in 2012. The limestone in which it was found yielded a diverse tooth assemblage, and was dissolved in acetic acid so they could be extracted. In total, 7 kg of limestone contained roughly 88 diagnosable teeth, including 41 teeth that undoubtedly belonged to elasmobranchs. While some of these had already been described, the others were mounted on stubs, coated with gold and scanned with an electron microscope at the Palaeontological Department of the University of Vienna.[1]

The generic name of Cretacladoides is a combination of "Cretaceous," "cladodont," and the Greek word "oides," which means "similar," alluding to its age and similarities to the teeth of Palaeozoic cladodontomorph chondrichthyans.[1]

Description

The teeth of Cretacladoides are multicuspid, with a cladodont-like crown, and measure less than 1mm in width and height. The main cusp is triangular, and flanked by either two or three pairs of lateral cusplets. The first pair of cusplets is very small, while the second reaches three quarters of the main cusp's height. The third pair, when present, is roughly half the size of the second pair. The cutting edges are apparently continuous between the apex of the main cusps and the cusplets. In profile view, all cusps are inclined lingually. The labial crown face possesses either an ogive-shaped enameloid structure on the main cusp, or two converging ridges.[1]

Viewed from the base, the root is D-shaped and of hemiaulacorhize type, possessing a small labial budge beneath the main cusp. From a lingual view, the root face has a well-developed protuberance below the main cusp. One pair of foramina opens laterally to this protuberance, while the second pair is below a notch which separates the second and third lateral cusplets. The aboral root face possesses two foramina, one central to the root face and the other in line with the main cusp lingually.[1]

Late Survival

Assuming that Cretacladoides is a member of the Falcatidae, a family of symmoriiform chondrichthyans that otherwise became extinct in the Carboniferous, it implies the existence of a ghost lineage.[1] One explanation put forward by its describers is that it occupied deep-sea habitats, which allowed its line to survive the Permian–Triassic extinction event and leave very few fossils.[1] It was described in conjunction with a possible ctenacanthid, Natarapax trivortex, which may indicate an additional ghost lineage in France.[1]

References

  1. ^ a b c d e f g h i Iris Feichtinger, Andrea Engelbrecht, Alexander Lukeneder & Jürgen Kriwet (2020) New chondrichthyans characterised by cladodont-like tooth morphologies from the Early Cretaceous of Austria, with remarks on the microstructural diversity of enameloid, Historical Biology, 32:6, 823-836, DOI: 10.1080/08912963.2018.1539971