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Charaxes castor

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This is an old revision of this page, as edited by Mothylator (talk | contribs) at 22:23, 7 October 2021 (Taxonomy: now amended to Related Species: so taxonomic revision (Turlin 2005), and phylogenetics (Out of Africa Again) can be evidenced to support the jasius Group & jasius Subgroup.and). The present address (URL) is a permanent link to this revision, which may differ significantly from the current revision.

Giant emperor
Male
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Nymphalidae
Genus: Charaxes
Species:
C. castor
Binomial name
Charaxes castor
(Cramer, [1775])[1]
Synonyms
  • Papilio castor Cramer, 1775 (not Westwood, 1842)
  • Charaxes castor var. godarti Aurivillius, 1899
  • Charaxes castor f. aginga Stoneham, 1931
  • Charaxes castor godarti f. severus Biederman, 1935
  • Charaxes castor f. flavimarginalis Stoneham, 1936
  • Charaxes castor var. flavifasciatus Butler, 1895
  • Charaxes castor var. orientalis Lanz, 1896
  • Charaxes castor flavifasciatus f. reimeri Rothschild, 1900
  • Charaxes castor flavifasciatus ab. dyscrita van Son, 1979

Charaxes castor, the giant emperor or giant charaxes, is a butterfly of the family Nymphalidae. It is found throughout the Afrotropical realm below the Sahel.[2]

The wingspan is 75–85 mm in males and 85–105 mm in females. The flight period is year-round but more common in late summer to autumn.[3]

Description

Male upperside from Toupé, Ivory Coast
Male underside from the Central African Republic
Female upperside from Maka, Central African Republic

A full description is given by Walter Rothschild and Karl Jordan, 1900 Novitates Zoologicae Volume 7:287-524. [1] page 436 et seq. (for terms see Novitates Zoologicae Volume 5:545-601 [2])

Life history

Larvae feed on Bridelia micrantha, Afzelia quanzensis, Tragia spp., Gymnosporia spp., Maytenus senegalensis, Schotia brachypetala, Bridelia micrantha and Cassia fistula.[2][3]

Subspecies

Listed alphabetically.[2]

  • C. c. arthuri van Someren, 1971 (Pemba island)
  • C. c. castor (Cramer, [1775]) (throughout African range)
  • C. c. comoranus Rothschild, 1903 (Comoros)
  • C. c. flavifasciatus Butler, 1895 (eastern Kenya, northern and eastern Tanzania, Malawi, Mozambique, eastern Zimbabwe, South Africa, Swaziland)

Historical attempts to assemble a cluster of presumably related species into a "Charaxes jasius Group" have not been wholly convincing. More recent taxonomic revision,[4] corroborated by phylogenetic research, allow a more rational grouping congruent with cladistic relationships. Within a well-populated clade of 27 related species sharing a common ancestor approximately 16 mya during the Miocene,[5] 26 are now considered together as the jasius Group.[4] One of the two lineages forms a robust clade of seven species sharing a common ancestor approximately 2-3 mya, i.e. during the Pliocene,[5] and are considered as the jasius subgroup.[4]

The jasius Group (26 Species)

Clade 1: jasius subgroup (7 species):[4]

Clade 2: contains three well-populated subgroups of the jasius Group:[4]

  • brutus subgroup (4 species);
  • pollux subgroup (4 species);
  • eudoxus subgroup (11 species);

The 27th species has shown rapid recent divergence and is treated as a monospecific lactetinctus Group, separated from the jasius Group.[5] Further exploration of the phylogenetic relationships amongst existing Charaxes taxons is required to improve clarity.

References

  1. ^ Afrotropical Butterflies: File H - Charaxinae - Tribe Charaxini
  2. ^ a b c "Charaxes Ochsenheimer, 1816" at Markku Savela's Lepidoptera and Some Other Life Forms
  3. ^ a b Woodhall, Steve (2005). Field Guide to Butterflies of South Africa. Cape Town, South Africa: Struik. ISBN 978-1-86872-724-7.
  4. ^ a b c d e f g Turlin, B. (2005). Bauer & Frankenbach (ed.). Butterflies of the World: Charaxes 1. Vol. 22. Keltern: Goecke & Evers. p. 3. ISBN 3937783156.
  5. ^ a b c "Out of Africa again: A phylogenetic hypothesis of the genus Charaxes (Lepidoptera: Nymphalidae) based on five gene regions". Aduse-Poku, Vingerhoedt, Wahlberg. Molecular Phylogenetics and Evolution (2009) 53;463–478
  • Victor Gurney Logan Van Someren, 1971 Revisional notes on African Charaxes (Lepidoptera: Nymphalidae). Part VII. Bulletin of the British Museum (Natural History) (Entomology)181-226.[3]