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Taxonomy of commonly fossilised invertebrates

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The Ordovician cystoid Echinosphaerites (an extinct echinoderm of the Class Rhombifera) from southern Estonia; approximately 555 cm in diameter.

Although the phylogenetic classification of sub-vertebrate animals (both extinct and extant) remains a work-in-progress, the following taxonomy attempts to be useful by combining both traditional (old) and new (21st-century) paleozoological termonology.

So the paleobiologic systematics which follows is not intended to be all-inclusive or completely comprehensive. For practical reasons and relevancy, the below classification and annotations emphasize invertebrates that (a) are popularly collected as fossils and/or (b) no longer continue alive on this planet. Therefore, as a result, some phyla, classes, and orders of invertebrates are not listed. [1]

If a non-vertebrate animal is mentioned below using its common, vernacular, everyday name, the creature is usually a living, present-day invertebrate. But if, on the other hand, a non-vertebrate is cited below by its scientific, taxonomic genus (in italics), then it is typically an extinct invertebrate, known only from the fossil record. [2]

Invertebrate clades that are (a) very important as fossils (for example, ostracods frequently used as index fossils), and/or (b) very abundant as fossils (for example, crinoids easily found in crinoidal limestone), [3] are highlighted with a bracketed exclamation mark [ ! ].

Invertebrate groups that (a) are now substantially extinct, and/or (b) contain a large proportion of extinct species, are followed by a dashed notation [ -- such as this ]. But invertebrate clades which are now totally -- that is, 100 percent -- extinct are designated with a bracketed dagger/cross [ † ]:


Domain of EUKARYOTA / EUKARYA

(eukaryotes / eukaryans / all cellular organisms bearing a central, organized nucleus with DNA)

Sub-domain of OPISTHOKONTA

(opisthokonts / the animal-related kingdoms / the proto-spongal choanoflagellates, proto-fungal microsporidians, true fungi, and true animals

  • comprises most life forms documented as either living or deceased
    • excludes many molds, all one-celled protists (or protoctists), all algae, and all green plants

Kingdom of ANIMALIA / METAZOA

(metazoans / many-celled true animals / all invertebrates and vertebrates / multi-cellular creatures that grab and ingest their organic food)

Sub-kingdom of PARAZOA

(parazoans / typically-sessile, basal non-eumetazoans / the most-primitive animals / the simplest, colonial, attached, bottom-dwelling, marine invertebrates)

Phylum Archaeocyatha / Archeocyatha / Archaeocyathida / Archeocyathida / Pleospongia [†]

(cone-shaped archaeocyathids/archeocyathids / cup-shaped archaeocyathans/archeocyathans / reef-building pleosponges / calcareous "ancient-cups")

(includes fossil genera such Archaeocyathus, Cambrocyathus, Atikonia, Tumuliolynthus, Kotuyicyathus, Metaldetes, Ajacicyathus and Paranacyathus)

(Archaeocyatha is sometimes classified as a class of Porifera below)

Phylum Porifera / Nuda / Spongia

(quintessential true sponges / marine, colonial, pore-bearing animals / organized collar-flagellates / poriferans; today mostly siliceous) -- half of all documented species of Porifera are fossils and extinct [4]

(Porifera may eventually be broken up into separate phyla)

Phylum Placozoa

Sub-kingdom of EUMETAZOA

(eumetazoans / true metazoans / typically-mobile, multicellular animals)

(Eumetazoa contains most of the living and deceased species of recorded life, including most invertebrates (alive and extinct), as well as all vertebrate animals)

Super-phylum of RADIATA

(radiates / non-bilaterian eumetazoans)

Phylum Cnidaria / Coelenterata

(cnidarians / coelenterates)

Super-phylum of LOPHOTROCHOZOA / PROTOSTOMIA # 1

(lophotrochozoan bilaterians, such as flatworms, ribbon worms, lophophorates, and molluscs)

Phylum Bryozoa / Ectoprocta / Polyzoa

(bryozoans / moss animals) -- half of all documented species of Bryozoa are fossils and extinct [5]

  • Class Stenolaemata / Gymnolaemata [!] (mostly marine, calcareous bryozoans)
    • Order Cheilostomata [!] (living, rimmed-mouthed moss animals)
    • Order Cyclostomata (uncontracted, round-mouthed bryozoans including fossil Stomatopora)
    • Order Cystoporata [†] (extinct, minor group of moss animals)
    • Order Trepostomata [†] [!] (changed-mouthed bryozoans such as extinct Constellaria and Monticulipora)
    • Order Cryptostomata [†] [!] (round hidden-mouthed bryozoans such as Archimedes, Fenestrellina and Rhombopora)
    • Order Ctenostomata [†] (uncommon, comb-mouthed bryozoans)
    • Order Phylactolaemata (living, fresh-water bryozoans)

Phylum Brachiopoda

Rhynchotrema dentatum, a rhynchonellid brachiopod from the Cincinnatian (Upper Ordovician) of southeastern Indiana.

(lampshells, brachiopods or "brachs," not to be confused with the hard-shelled marine mollusks below) -- 99 percent of all documented species of Brachiopoda are now extinct

Phylum Annelida

(segmented worms such as earthworms and leeches)

Phylum Mollusca

(molluscs or mollusks, not to be confused with the hard-shelled marine brachiopods above)

Super-phylum of ECDYSOZOA / PROTOSTOMIA # 2

(ecdysozoans, such as nematodes, horsehair worms, and molting bilaterians / panarthropods))

Phylum Tardigrada

(panarthropodic water bears)

Phylum Onychophora

(panarthropodic velvet worms, including proto-arthropodic fossils of Arthropleura and Aysheaia)

Phylum Arthropoda

(arthropods; jointed legged creatures with an exoskeleton)

Super-phylum of DEUTEROSTOMIA / ENTEROCOELOMATA

(second-mouthed bilaterians called deuterostomians, such as chordates and echinoderms)

Phylum Echinodermata

(echinoderms) -- 72 percent of all documented species of Echinodermata are fossils and extinct [7]

Phylum Hemichordata

(hemichordates such as extant acorn worms) -- Less than half of the documented species of Hemichordata are fossils and extinct

Phylum Chordata

(both invertebrate and vertebrate chordates; animals possessing a notochord)

Invertebrate subphyla
Subphylum Vertebrata

Footnotes

  1. ^ For superb anatomical illustrations and much-more comprehensive information, the aspiring paleozoologist should scan Volume E (Archaeocyatha / Porifera) through Volume V (Graptolithina), published 1953 to 2006 (and continuing), of the Treatise on Invertebrate Paleontology, long-edited by Raymond C. Moore and Roger L. Kaesler (Boulder, Colorado: Geological Society of America; and Lawrence, Kansas: University of Kansas Press). But be warned that some terms therein employed -- such as supersubphylum -- can be unnecessarily wordy or obstruse. Incidentally, revised volumes have been recently published regarding the sponges/archaeocyatha (2004, ISBN 08137 31313) and the brachiopods (2006, ISBN 0813731356).
  2. ^ The names of genera, orders, classes and phyla have been culled from dozens of sources, both current and decades-old. See the International Code of Zoological Nomenclature (ICZN), as well as Volume 1 and Volume 2 of Grzimek's Animal Life Encyclopedia (Farmington Hills, Michigan: Gale Group), edited by zoologists Michael Hutchin, Dennis A. Thorney and Sean F. Craig (2003).
  3. ^ For correspondingly-ancient ecosystems, see the Treatise on Ecology and Paleoecology, Volume 2: Paleoecology, edited for years by Harry S. Ladd (1957 / 1971), and published by both the Geological Society of America (Boulder, Colorado) and the Waverly Press (Washington, D.C.).
  4. ^ The rates of extinction for sponges and other phyla are derived from W. H. Easton, 1960, Invertebrate Paleontology (New York: Harper and Brothers) and various modern sources.
  5. ^ For bryozoans and brachiopods, the same footnote as above.
  6. ^ For bivalves and cephalopods (both mollusks), see the above notation.
  7. ^ For the echinoderms, see the above footnote regarding W. E. Easton, 1960, Invertebrate Paleontology, and other sources.

See also

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