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A conserved protein binding domain of approximately 80 amino-acid alpha crystallins are knows as small heat shock proteins (sHSP)^[1]

According to Marvin et al sHSPs independently express not only in heat shock response but also have developmental roles in embryonic or juvenile stages of mammals, teleost fish and some lower vertebral gemones.^[2]

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^ Lahvic, Jamie L.; Ji, Yongchang; Marin, Paloma; Zuflacht, Jonah P.; Springel, Mark W.; Wosen, Jonathan E.; Davis, Leigh; Hutson, Lara D.; Amack, Jeffrey D. (2013-12-15). "Small heat shock proteins are necessary for heart migration and laterality determination in zebrafish". Developmental biology. 384 (2): 166–180. doi:10.1016/j.ydbio.2013.10.009. ISSN 0012-1606. PMC 3924900. PMID 24140541.{{cite journal}}: CS1 maint: PMC format (link)
^ Marvin, Martha; O'Rourke, Devon; Kurihara, Tomoki; Juliano, Courtney E.; Harrison, Krista L.; Hutson, Lara D. (February 2008). "Developmental expression patterns of the zebrafish small heat shock proteins". Developmental Dynamics: An Official Publication of the American Association of Anatomists. 237 (2): 454–463. doi:10.1002/dvdy.21414. ISSN 1058-8388. PMID 18161059.

[1] Lahvic, Jamie L.; Ji, Yongchang; Marin, Paloma; Zuflacht, Jonah P.; Springel, Mark W.; Wosen, Jonathan E.; Davis, Leigh; Hutson, Lara D.; Amack, Jeffrey D. (2013-12-15). "Small heat shock proteins are necessary for heart migration and laterality determination in zebrafish". Developmental biology. 384 (2): 166–180. doi:10.1016/j.ydbio.2013.10.009. ISSN 0012-1606. PMC 3924900. PMID 24140541.{{cite journal}}: CS1 maint: PMC format (link)

[2] Marvin, Martha; O'Rourke, Devon; Kurihara, Tomoki; Juliano, Courtney E.; Harrison, Krista L.; Hutson, Lara D. (February 2008). "Developmental expression patterns of the zebrafish small heat shock proteins". Developmental Dynamics: An Official Publication of the American Association of Anatomists. 237 (2): 454–463. doi:10.1002/dvdy.21414. ISSN 1058-8388. PMID 18161059.

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