Stipeae

Stipeae
	Celtica gigantea
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Clade:	Commelinids
Order:	Poales
Family:	Poaceae
Clade:	BOP clade
Subfamily:	Pooideae
Supertribe:	Stipodae
Tribe:	Stipeae; Dumort. (1824)
Genera
	28 genera, see text
Synonyms
	supertribe Stipodae L. Liu (1980); subtribe Stipinae Griseb. (1846); Aciachninae Caro (1982); Ortachninae Caro (1982);

The Stipeae are a tribe of grasses within the subfamily Pooidae, with up to 600 described species.^[2]

Description

The defining morphological features of the Stipeae include single-flowered spikelets lacking a rachilla extension, and the lemmas (the external bract) have either a sharp point or a terminal awn (long bristle).^[3]

Genera

The tribe includes 28 genera:^[1]

Many species initially placed into Stipa have now been split off into new genera. Some recent papers have analysed relationships within and between the genera,^[4]^[5]^[6] but a complete analysis has not yet been performed. Stipoid grasses use the C₃ photosynthetic pathway and live in temperate areas worldwide.^[7]

Known fossils date from the late Miocene.^[8]

References

^ ^a ^b Soreng, Robert J.; Peterson, Paul M.; Romaschenko, Konstantin; Davidse, Gerrit; Teisher, Jordan K.; Clark, Lynn G.; Barberá, Patricia; Gillespie, Lynn J.; Zuloaga, Fernando O. (2017). "A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications". Journal of Systematics and Evolution. 55 (4): 259–290. doi:10.1111/jse.12262. ISSN 1674-4918.
^ Barkworth ME, Arriaga MO, Smith JF, Jacobs SWL, Valdes-Reyna J, Bushman BS (2008) Molecules and Morphology in South American Stipeae (Poaceae). Systematic Botany 33(4), 719-731.
^ Cialdella AM, Giussani LM, Aagesen L, Zuloaga FO, Morrone O (2007) A phylogeny of Piptochaetium (Poaceae : Pooideae : Stipeae) and related genera based on a combined analysis including trnL-F, rp116, and morphology. Systematic Botany 32(3), 545-559.
^ Barber, Janet C.; Hames, Kasey A.; Cialdella, Ana Maria; Giussani, Liliana M.; Morrone, Osvaldo (2009). "Phylogenetic relationships of Piptochaetium Presl (Poaceae: Stipeae) and related genera reconstructed from nuclear and chloroplast sequence datasets". Taxon. 58 (2): 375–380. doi:10.1002/tax.582005. ISSN 0040-0262. pdf
^ Jacobs SWL, Bayer R, Everett J, Arriaga MO, Barkworth ME, Sabin-Badereau A, Torres MA, Vazquez FM, Bagnall N (2007) Systematics of the tribe Stipeae (Gramineae) using molecular data. Aliso 23, 349-361.
^ Jacobs SWL, Everett J, Barkworth ME, Hsiao C (2000) Relationships within the Stipoid grasses (Gramineae). In 'Grass Systematics and Evolution.' (Eds SWL Jacobs and J Everett) pp. 75-82. (CSIRO: Melbourne)
^ Romaschenko K, Peterson PM, Soreng RJ, Garcia-Jacas N, Futorna O, Susanna A (2008). "Molecular phylogenetic analysis of the American Stipeae (Poaceae) resolves Jarava sensu lato polyphyletic: evidence for a new genus, Pappostipa". Journal of the Botanical Research Institute of Texas. 2 (1): 165–192. JSTOR 41971613.{{cite journal}}: CS1 maint: multiple names: authors list (link) pdf
^ Thomasson, Joseph R. (2005). "Berriochloa Gabeli and Berrichloa Huletti (Graminae: Stipeae), two new grass species from the late miocene Ash Hollow Formationof Nebraska and Kansas". Journal of Paleontology. 79 (1): 185–199. doi:10.1666/0022-3360(2005)079<0185:BGABHG>2.0.CO;2. ISSN 0022-3360. pdf

External links

Stipa L. sensu lato (sensu lato refers to the tribe) The grass genera of the world website by L. Watson, T.D. Macfarlane, and M.J. Dallwitz

[SorengPeterson2017-1] Soreng, Robert J.; Peterson, Paul M.; Romaschenko, Konstantin; Davidse, Gerrit; Teisher, Jordan K.; Clark, Lynn G.; Barberá, Patricia; Gillespie, Lynn J.; Zuloaga, Fernando O. (2017). "A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications". Journal of Systematics and Evolution. 55 (4): 259–290. doi:10.1111/jse.12262. ISSN 1674-4918.

[2] Barkworth ME, Arriaga MO, Smith JF, Jacobs SWL, Valdes-Reyna J, Bushman BS (2008) Molecules and Morphology in South American Stipeae (Poaceae). Systematic Botany 33(4), 719-731.

[3] Cialdella AM, Giussani LM, Aagesen L, Zuloaga FO, Morrone O (2007) A phylogeny of Piptochaetium (Poaceae : Pooideae : Stipeae) and related genera based on a combined analysis including trnL-F, rp116, and morphology. Systematic Botany 32(3), 545-559.

[4] Barber, Janet C.; Hames, Kasey A.; Cialdella, Ana Maria; Giussani, Liliana M.; Morrone, Osvaldo (2009). "Phylogenetic relationships of Piptochaetium Presl (Poaceae: Stipeae) and related genera reconstructed from nuclear and chloroplast sequence datasets". Taxon. 58 (2): 375–380. doi:10.1002/tax.582005. ISSN 0040-0262. pdf

[5] Jacobs SWL, Bayer R, Everett J, Arriaga MO, Barkworth ME, Sabin-Badereau A, Torres MA, Vazquez FM, Bagnall N (2007) Systematics of the tribe Stipeae (Gramineae) using molecular data. Aliso 23, 349-361.

[6] Jacobs SWL, Everett J, Barkworth ME, Hsiao C (2000) Relationships within the Stipoid grasses (Gramineae). In 'Grass Systematics and Evolution.' (Eds SWL Jacobs and J Everett) pp. 75-82. (CSIRO: Melbourne)

[7] Romaschenko K, Peterson PM, Soreng RJ, Garcia-Jacas N, Futorna O, Susanna A (2008). "Molecular phylogenetic analysis of the American Stipeae (Poaceae) resolves Jarava sensu lato polyphyletic: evidence for a new genus, Pappostipa". Journal of the Botanical Research Institute of Texas. 2 (1): 165–192. JSTOR 41971613.{{cite journal}}: CS1 maint: multiple names: authors list (link) pdf

[8] Thomasson, Joseph R. (2005). "Berriochloa Gabeli and Berrichloa Huletti (Graminae: Stipeae), two new grass species from the late miocene Ash Hollow Formationof Nebraska and Kansas". Journal of Paleontology. 79 (1): 185–199. doi:10.1666/0022-3360(2005)079<0185:BGABHG>2.0.CO;2. ISSN 0022-3360. pdf

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