Smutsia

African ground pangolin; Temporal range: 9.8–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Late Miocene - Present
	Pangolins from genus Smutsia
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Pholidota
Family:	Manidae
Subfamily:	Smutsiinae; Gray, 1873
Genus:	Smutsia; Gray, 1865
Type species
	Smutsia gigantea; Illiger, 1815
Species
	Smutsia gigantea; Smutsia temminckii;
	Smutsia gigantea Smutsia temminckii
Synonyms
	synonyms of subfamily: Smutsiana (Gray, 1873) ; Smutsiini (Gray, 1873) ;

African ground pangolin (Smutsia) is a genus of African pangolins from subfamily large African pangolins (Smutsiinae), within family Manidae. It is formerly considered a subgenus of Manis.^[2] Its members are the more terrestrial of the African pangolins.^[3]

Taxonomy

Subfamily: Smutsiinae (large African pangolins)
- Genus: Smutsia (African ground pangolin)
  - Smutsia gigantea (Giant pangolin),
  - Smutsia temminckii (Ground pangolin)

Phylogeny

Phylogenetic position of genus Smutsia within family Manidae.^[4]^[5]^[1]

Pholidotamorpha

†Palaeanodonta

Pholidota

†Euromanis

†Eurotamanduidae

Eupholidota

†Eomanoidea

Manoidea

†Patriomanidae

?

†Necromanis

Manidae

Maninae

?

†manidае sp. (DPC 3972 & DPC 4364)

Smutsiinae

Phatagininae

Smutsiinae

Smutsia

	Smutsia gigantea

	†Smutsia sp. (Olteţ River Valley, Romania)

	Smutsia temminckii

sensu stricto

sensu lato

sensu stricto

(Pholidota sensu lato)

References

^ ^a ^b Philippe Gaubert, Agostinho Antunes, Hao Meng, Lin Miao, Stéphane Peigné, Fabienne Justy, Flobert Njiokou, Sylvain Dufour, Emmanuel Danquah, Jayanthi Alahakoon, Erik Verheyen, William T Stanley, Stephen J O’Brien, Warren E Johnson, Shu-Jin Luo (2018) "The Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most Trafficked Mammals on Earth" Journal of Heredity, Volume 109, Issue 4, Pages 347–359
^ Schlitter, D.A. (2005). "Order Pholidota". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 530–531. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ du Toit, Z.; du Plessis, M.; Dalton, D. L.; Jansen, R.; Paul Grobler, J.; Kotzé, A. (2017). "Mitochondrial genomes of African pangolins and insights into evolutionary patterns and phylogeny of the family Manidae". BMC Genomics. 18 (1): 746. doi:10.1186/s12864-017-4140-5. PMC 5609056. PMID 28934931.
^ Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. 16 (4). Heidelberg, Germany: Springer Science+Business Media: 235–305. doi:10.1007/s10914-009-9119-9.
^ Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. doi:10.1080/02724634.2012.694319. ISSN 0272-4634.

[Gaubert2018-1] Philippe Gaubert, Agostinho Antunes, Hao Meng, Lin Miao, Stéphane Peigné, Fabienne Justy, Flobert Njiokou, Sylvain Dufour, Emmanuel Danquah, Jayanthi Alahakoon, Erik Verheyen, William T Stanley, Stephen J O’Brien, Warren E Johnson, Shu-Jin Luo (2018) "The Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most Trafficked Mammals on Earth" Journal of Heredity, Volume 109, Issue 4, Pages 347–359

[Schlitter-2] Schlitter, D.A. (2005). "Order Pholidota". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 530–531. ISBN 978-0-8018-8221-0. OCLC 62265494.

[du_Toit2017-3] u Toit, Z.; du Plessis, M.; Dalton, D. L.; Jansen, R.; Paul Grobler, J.; Kotzé, A. (2017). "Mitochondrial genomes of African pangolins and insights into evolutionary patterns and phylogeny of the family Manidae". BMC Genomics. 18 (1): 746. doi:10.1186/s12864-017-4140-5. PMC 5609056. PMID 28934931.

[web2.utc.edu-4] Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. 16 (4). Heidelberg, Germany: Springer Science+Business Media: 235–305. doi:10.1007/s10914-009-9119-9.

[KondrashovAgadjanian2012-5] Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. doi:10.1080/02724634.2012.694319. ISSN 0272-4634.

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