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An operant conditioning chamber (also known as a Skinner Box) is a device used to study animal behavior, especially operant conditioning. It was invented by Skinner while he was a graduate student at Harvard University, where he received the doctorate in 1931. As used by Skinner, the box had a lever (for rats) or a disk in one wall (for pigeons). A press on this "manipulandum" could deliver food to the animal through an opening in the wall, and responses reinforced in this way increased in frequency. By controlling this reinforcement together with discriminative stimuli such as lights and tones, or punishments such as electric shocks, experimenters have used the operant box to study a wide variety of topics, including schedules of reinforcement, discriminative control, delayed response ("memory"), punishment, and so on. By channeling research in these directions, the operant conditioning chamber has had a huge influence on course of research in animal learning and its applications. It enabled great progress on problems that could be studied by the measuring the rate, probability, or force of a simple, repeatable response. However, it discouraged the study of behavioral processes not easily conceptualized in such terms - spatial learning, in particular, which is now studied in quite different ways, for example by the use of the water maze..