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薩摩耶犬

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薩摩耶犬
別名西摩犬
原產國俄罗斯西伯利亚
特徵
體重 雄性 20—30公斤(44—66英磅)
雌性 16—20公斤(35—44英磅)
身高 雄性 51—56 cm(20—22英寸)
雌性 46—51 cm(18—20英寸)
顏色 白色
壽命 12–14年
分類/標準
FCI 標⁠準
家犬Canis lupus familiaris

薩摩耶犬(英語:Samoyed;俄语:Самое́дская соба́ка或Самое́д)是一种带有双层白色厚毛的中型牧羊犬。萨摩耶是一种斯皮茨英语Spitz犬,名字取自西伯利亚地区的萨摩耶族。萨摩耶犬是涅涅茨牧羊莱卡犬英语Nenets Herding Laika的后代,被驯化以辅助牧羊、狩猎、看家以及拉雪橇。

一只两岁半的雄性萨摩耶

萨摩耶犬通常是白色的,也可能会有棕色的品种。其双层厚毛是天然的防水层。萨摩耶犬以被用于在北极南极地区进行探险及有着友善且亲人的性格而闻名。

血统

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萨摩耶犬,约摄于1915年

萨摩耶犬的祖先为涅涅茨牧羊莱卡犬英语Nenets Herding Laika,是一种在西伯利亚北部广泛使用的牧鹿用斯皮茨犬。在当时,被蔑称为萨摩耶族人的涅涅茨人尤其喜爱使用这种犬。[1][2][3]DNA证据确认了萨摩耶是一种基础犬种,其起源早于19世纪现代犬种的出现。[4]一项针对从亚马尔半岛处的涅涅茨人中所取得的近一百年前的两个犬只标本的基因组研究发现,这两个标本与距今2,000年和850年的两个标本有血缘关系,表明这一区域的犬只血统具有延续性。这两只一百年前的犬与萨摩耶品种犬有血缘关系,也表明了古代北极圈犬只的血统在现代的萨摩耶犬身上延续了下来。[5]

启程前往北极的南森·约翰森

During preparation for the Fram expedition to the North Pole in 1893–1896, 33 dogs were purchased from the Nenets people. While 28 of these dogs would go to the North Pole, none of them survived. The remaining dogs, including pups born during the voyage, were left aboard the ship. In April 1893 the bitch had another litter, most of them white. According to Nansen's notes "...all the dogs were strong, tough and excellent at pulling sleds; they worked very well in hunting Polar bears [as well]." These dogs would become the original Samoyeds.[2][6]

Appearance and characteristics

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Samoyed puppy

The AKC Standard requires 45—65英磅(20—29公斤) and 21—23.5英寸(53—60厘米) at the shoulder for males, and 35—50英磅(16—23公斤) and 19—21英寸(48—53厘米) for females.[7] The UK Kennel Club Standard requires 51—56厘米(20—22英寸) for males, and 46—51厘米(18—20英寸) for females.

Samoyed ears are thick and covered with fur, triangular in shape, and erect. They are almost always white but have a light to dark brown tint (known as "biscuit") to a greater or lesser extent. The tint is usually on the ears but can be visible on the whole body.

A two year old adult female Samoyed. The breed is characterized by an alert and happy expression.

The Samoyed tail is one of the breed's distinguishing features. Like the Alaskan Malamute, the tail is carried curled over the back; however, unlike the Alaskan Malamute, the Samoyed tail is held actually touching the back. It is not usually held in a tight curl, or held flag-like; it is usually carried lying over the back and to one side. In cold weather, Samoyeds may sleep with their tails over their noses to provide additional warmth. Almost all Samoyeds will allow their tails to fall when they are relaxed and at ease, as when being stroked or while eating, but will return their tails to a curl when more alert.

Samoyeds have a dense, double layer coat. The topcoat contains long, coarse, and straight guard hairs, which appear white but have a hint of silver coloring. This top layer keeps the undercoat relatively clean and free of debris. The under layer, or undercoat, consists of a dense, soft, and short fur that keeps the dog warm. The undercoat typically sheds heavily once or twice a year, and this seasonal process is sometimes referred to as "blowing coat". This does not mean the Samoyed will shed only during that time however; fine hairs (versus the dense clumps shed during seasonal shedding) will be shed all year round, and have a tendency to stick to cloth and float in the air. The standard Samoyed may come in a mixture of biscuit and white coloring, although pure white and all biscuit dogs are common. Males typically have larger ruffs than females. While this breed is touted as "hypoallergenic", it does shed a fair amount and needs frequent grooming. While the breed may produce fewer allergens, care should be taken for severe allergies.[8]

Shed Samoyed fur is sometimes used as an alternative to wool in knitting, with a texture similar to angora. The fur is sometimes also used for the creation of artificial flies for fly fishing.

Life expectancy for the breed is about 12–13 years.[9]

Temperament

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A Samoyed resting with a teddy bear

Samoyeds' friendly and affable disposition makes them poor guard dogs; an aggressive Samoyed is rare. The breed is characterized by an alert and happy expression which has earned the nicknames "Sammie smile" and "smiley dog".[10] With their tendency to bark, however, they can be diligent watch dogs, barking whenever something approaches their territory. Samoyeds are excellent companions, especially for small children or even other dogs, and they remain playful into old age. According to the Samoyed Club of America, when Samoyeds become bored, they may become destructive or start to dig.[11]

Activities

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An active Samoyed

Samoyeds can compete in dog agility trials, carting, obedience, showmanship, flyball, tracking, mushing and herding events. Herding instincts and trainability can be measured at non-competitive herding tests. Samoyeds exhibiting basic herding instincts can be trained to compete in herding trials.[12]

Health

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A 2024 UK study found a life expectancy of 13.1 years for the breed compared to an average of 12.7 for purebreeds and 12 for crossbreeds.[13]

Samoyed hereditary glomerulopathy

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The breed can be affected by a genetic disease known as Samoyed hereditary glomerulopathy, a kidney disease. The disease is known to be caused by an X-linked recessive faulty allele and therefore the disease is more severe in male Samoyeds.[14] Also known as hereditary nephritis, it is caused by a nonsense mutation in codon 1027 of the COL4A5 gene on the X chromosome (glycine to stop codon), which is similar to Alport's syndrome in humans.

A senior female Samoyed

Carrier females do develop mild symptoms after 2–3 months of age, but mostly[15] do not go on to develop kidney failure. The disease is caused by a defect in the structure of the type-IV collagen fibrils of the glomerular basement membrane. As a consequence, the collagen fibrils of the glomerular basement membrane are unable to form cross-links, so the structural integrity is weakened and the membrane is more susceptible to "wear-and-tear" damage. As the structure of the basement membrane begins to degenerate, plasma proteins are lost in the urine and symptoms begin to appear. Affected males appear healthy for the first three months of life, but then symptoms start to appear and worsen as the disease progresses: the dog becomes lethargic and muscle wastage occurs, as a result of proteinuria. From three months of age onwards, a reduced glomerular filtration rate is detected, indicative of progressive kidney failure.

Clinically, proteinuria is found in both sexes from the age of three to four months; in dogs older than this, kidney failure in combination with more or less pronounced hearing loss occurs swiftly and death at the age of 8 to 15 months is expected. In heterozygous females, the disease develops slowly. The disease can be treated to slow down the development by use of cyclosporine A and ACE inhibitors, but not stopped.[14][16][17][18]

If a carrier female is mated with a healthy stud dog, the female offspring have a 50% chance of being carriers for the disease, and any male offspring have a 50% chance of being affected by the disease. A genetic test is available for this disease.[19]

Other health concerns

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For the Samoyeds several breed-specific hereditary diseases are described in the veterinary literature:

  • Diabetes mellitus similar but not identical to human Type I (insulin deficiency): The disease occurs in middle-aged Samoyeds, the mean age at diagnosis is seven years. The cause is a chronic inflammation of the pancreas and/or autoimmune destruction of the beta cells of islets of Langerhans. Moreover, autoantibodies to insulin were found in affected dogs. Several genetic markers are being discussed as possible causes.[20][21]
  • Progressive retinal atrophy (PRA) caused by a frameshift mutation in the RPRG locus of the X chromosome. The disease leads to a slowly progressive loss of vision, which eventually leads to blindness. The first symptoms appear between two and five years of age. The disease corresponds to the X-linked PRA type 3 in humans.[22][23]
  • Short legs in conjunction with eye abnormalities: a genetic defect at the COL2A1 locus leads to disproportionate dwarfism due to short limbs in connection with cataracts, malformations of the retina and / or retinal detachment, liquefaction of the vitreous and a persistent hyaloid artery. The malformations of the retina are dominant (i.e. they occur in heterozygous dogs); the other symptoms are recessive, so that they are expressed only in homozygous dogs. These conditions have no effect on the expression of the protein opticin.[24][25][26]
  • Pulmonary stenosis occurs more frequently in Samoyeds in comparison with other breeds. The disease can cause shortness of breath, cardiac arrhythmias and rapid fatigue when moving, and increases the risk of congestive heart failure.[27]
  • Hip dysplasia is also a concern for Samoyeds.[28]
  • The breed can also be affected by sebaceous adenitis, an uncommon idiopathic autoimmune skin disease.[29]

See also

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References

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  1. ^ Давай-ка, ненецкая лайка!. kras.mk.ru. 2017-12-20 [2022-02-13] (俄语). 
  2. ^ 2.0 2.1 Presberg, Carole. Herding Dogs of Asia: Russian Siberia. www.bordercolliemuseum.org. 2014 [2022-02-13]. 
  3. ^ Anderson, David G. Siberian Survival: The Nenets and Their Story.. American Anthropologist. 2000, 102 (4): 942–943. ISSN 1548-1433. doi:10.1525/aa.2000.102.4.942 (英语). 
  4. ^ Larson, G. Rethinking dog domestication by integrating genetics, archeology, and biogeography. Proc. Natl. Acad. Sci. U.S.A. 2012, 109 (23): 8878–83. Bibcode:2012PNAS..109.8878L. PMC 3384140可免费查阅. PMID 22615366. doi:10.1073/pnas.1203005109可免费查阅. 
  5. ^ Feuerborn, Tatiana R.; Carmagnini, Alberto; Losey, Robert J.; Nomokonova, Tatiana; Askeyev, Arthur; Askeyev, Igor; Askeyev, Oleg; Antipina, Ekaterina E.; Appelt, Martin; Bachura, Olga P.; Beglane, Fiona; Bradley, Daniel G.; Daly, Kevin G.; Gopalakrishnan, Shyam; Murphy Gregersen, Kristian; Guo, Chunxue; Gusev, Andrei V.; Jones, Carleton; Kosintsev, Pavel A.; Kuzmin, Yaroslav V.; Mattiangeli, Valeria; Perri, Angela R.; Plekhanov, Andrei V.; Ramos-Madrigal, Jazmín; Schmidt, Anne Lisbeth; Shaymuratova, Dilyara; Smith, Oliver; Yavorskaya, Lilia V.; Zhang, Guojie; Willerslev, Eske; Meldgaard, Morten; Gilbert, M. Thomas P.; Larson, Greger; Dalén, Love; Hansen, Anders J.; Sinding, Mikkel-Holger S.; Frantz, Laurent. Modern Siberian dog ancestry was shaped by several thousand years of Eurasian-wide trade and human dispersal. Proceedings of the National Academy of Sciences. 2021, 118 (39): e2100338118. Bibcode:2021PNAS..11800338F. ISSN 0027-8424. PMC 8488619可免费查阅. PMID 34544854. S2CID 237584023. doi:10.1073/pnas.2100338118可免费查阅. 
  6. ^ The Samoyed: Breed Origin and History. samoyedclubofamerica.org. 
  7. ^ Samoyed. AKC. 
  8. ^ Bakalar, Nicholas. The Myth of the Allergy-Free Dog. The New York Times. 11 July 2011 [18 July 2015]. 
  9. ^ Individual Breed Results for Purebred Dog Health Survey. TheKennelClub.org.uk. [26 June 2012]. (原始内容存档于13 August 2013). 
  10. ^ AKC Meet the Breeds: Samoyed. American Kennel Club. [8 October 2011]. (原始内容存档于11 January 2013). 
  11. ^ Bad Habits and Training. 
  12. ^ Hartnagle-Taylor, Jeanne Joy; Taylor, Ty. Stockdog Savvy. Alpine Publications. 2010. ISBN 978-1-57779-106-5. [页码请求]
  13. ^ McMillan, Kirsten M.; Bielby, Jon; Williams, Carys L.; Upjohn, Melissa M.; Casey, Rachel A.; Christley, Robert M. Longevity of companion dog breeds: those at risk from early death. Scientific Reports (Springer Science and Business Media LLC). 2024-02-01, 14 (1): 531. ISSN 2045-2322. PMC 10834484可免费查阅. PMID 38302530. doi:10.1038/s41598-023-50458-w. 
  14. ^ 14.0 14.1 Jansen, B; Tryphonas, L; Wong, J; Thorner, P; Maxie, MG; Valli, VE; Baumal, R; Basrur, PK. Mode of inheritance of Samoyed hereditary glomerulopathy: an animal model for hereditary nephritis in humans. The Journal of Laboratory and Clinical Medicine. 1986, 107 (6): 551–5. PMID 3711721. 
  15. ^ Rawdon, TG. Juvenile nephropathy in a Samoyed bitch. The Journal of Small Animal Practice. 2001, 42 (5): 235–8. PMID 11380016. doi:10.1111/j.1748-5827.2001.tb02027.x. 
  16. ^ Zheng, K; Thorner, PS; Marrano, P; Baumal, R; McInnes, RR. Canine X chromosome-linked hereditary nephritis: a genetic model for human X-linked hereditary nephritis resulting from a single base mutation in the gene encoding the alpha 5 chain of collagen type IV. Proceedings of the National Academy of Sciences of the United States of America. 1994, 91 (9): 3989–93. Bibcode:1994PNAS...91.3989Z. PMC 43708可免费查阅. PMID 8171024. doi:10.1073/pnas.91.9.3989可免费查阅. 
  17. ^ Grodecki, K; Gains, M; Baumal, R; Osmond, D; Cotter, B; Valli, V; Jacobs, R. Treatment of X-linked hereditary nephritis in samoyed dogs with angiotensin converting enzyme (ACE) inhibitor. Journal of Comparative Pathology. 1997, 117 (3): 209–225. PMID 9447482. doi:10.1016/S0021-9975(97)80016-3. 
  18. ^ Chen, D.; Jefferson, B; Harvey, SJ; Zheng, K; Gartley, CJ; Jacobs, RM; Thorner, PS. Cyclosporine A Slows the Progressive Renal Disease of Alport Syndrome (X-Linked Hereditary Nephritis): Results from a Canine Model. Journal of the American Society of Nephrology. 2003, 14 (3): 690–8. PMID 12595505. doi:10.1097/01.ASN.0000046964.15831.16可免费查阅. 
  19. ^ Samoyed Hereditary Glomerulopathy. vetgen.com. Veterinary Genetic Services. [3 February 2013]. (原始内容存档于16 April 2011). 
  20. ^ Kimmel, SE; Ward, CR; Henthorn, PS; Hess, RS. Familial insulin-dependent diabetes mellitus in Samoyed dogs. Journal of the American Animal Hospital Association. 2002, 38 (3): 235–8. PMID 12022409. doi:10.5326/0380235. 
  21. ^ Short, A. D.; Catchpole, B.; Kennedy, L. J.; Barnes, A.; Fretwell, N.; Jones, C.; Thomson, W.; Ollier, W. E.R. Analysis of Candidate Susceptibility Genes in Canine Diabetes. Journal of Heredity. 2007, 98 (5): 518–525. PMID 17611256. doi:10.1093/jhered/esm048可免费查阅. 
  22. ^ Dice, P. F. 2nd. Progressive retinal atrophy in the Samoyed. Modern Veterinary Practice. 1980, 61 (1): 59–60. PMID 7366567. 
  23. ^ Zangerl, B.; Johnson, J. L.; Acland, G. M.; Aguirre, G. D. Independent Origin and Restricted Distribution of RPGR Deletions Causing XLPRA. Journal of Heredity. 2007, 98 (5): 526–530. PMID 17646274. doi:10.1093/jhered/esm060可免费查阅. 
  24. ^ Meyers, VN; Jezyk, PF; Aguirre, GD; Patterson, DF. Short-limbed dwarfism and ocular defects in the Samoyed dog. Journal of the American Veterinary Medical Association. 1983, 183 (9): 975–79. PMID 12002589. 
  25. ^ Acland, Gregory M. Retinal dysplasia in the Samoyed dog is the heterozygous phenotype of the gene (drds) for short limbed dwarfism and ocular defects. Transactions of the American College of Veterinary Ophthalmology. 1991, 22: 44. 
  26. ^ Pellegrini, B; Acland, GM; Ray, J. Cloning and characterization of opticin cDNA: evaluation as a candidate for canine oculo-skeletal dysplasia. Gene. 2002, 282 (1–2): 121–131. PMID 11814684. doi:10.1016/S0378-1119(01)00842-3. 
  27. ^ McCaw, D; Aronson, E. Congenital cardiac disease in dogs. Modern Veterinary Practice. 1984, 65 (7): 509–12. PMID 6749116. 
  28. ^ Martin, SW; Kirby, K; Pennock, PW. Canine hip dysplasia: breed effects. The Canadian Veterinary Journal. 1980, 21 (11): 293–6. PMC 1789813可免费查阅. PMID 7459792. 
  29. ^ Craig, Mark. Clinical refresher: Canine sebaceous adenitis. Companion Animal. 2006, 11 (5): 62–8. doi:10.1111/j.2044-3862.2006.tb00066.x. 

Further reading

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狐狸犬、雪橇犬家族的一員,原是俄國西伯利亞原住民薩摩耶族培育出的犬種。因看起來永遠在笑的樣子又喜歡親近人,而被稱為「微笑天使」、「笑臉天使」、「雪天使」。但同時,他們很容易狼嚎

除了溫和的個性,薩摩耶犬也以忍耐力跟健壯著稱,原是用來拉雪橇跟看守馴鹿,也曾被西方探險家用來從事極地探險的任務。

原有多種毛色,黑、黑白、黑褐,但十九世紀毛皮商人認為白色會最受歡迎而大量輸出白薩摩耶犬到歐美。今日所見的純種薩摩耶犬多是白色系。

性格特征

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聰明,溫和,忠誠,適應性強,機警,充滿活力,樂於服務,友好而富有激情,勇敢而樂於嘗試,真誠而不會去懷疑,進攻性不強。

外部連結

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