Portal:Amphibians
The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
Young amphibians generally undergo metamorphosis from an aquatic larval form with gills to an air-breathing adult form with lungs. Amphibians use their skin as a secondary respiratory interface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require access to water bodies to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators to habitat conditions; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from tetrapodomorph sarcopterygians (lobe-finned fish with articulated limb-like fins) that evolved primitive lungs, which were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (predecessors of mammals). The origin of modern lissamphibians, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondyls. A fourth group of lissamphibians, the Albanerpetontidae, became extinct around 2 million years ago. (Full article...)
Selected frog article
The Hemiphractidae are a family of frogs from South and Central America. Previously, this group had been classified as a subfamily (Hemiphractinae) under family Hylidae. More recent research classifies these genera into their own family, or sometimes into three separate families: Amphignathodontidae (Flectonotus and Gastrotheca), Cryptobatrachidae (Cryptobatrachus and Stefania), and Hemiphractidae (Hemiphractus and Fritziana). An active question still exists as to which of these groupings is more accurate. (Full article...)
Selected salamander article
The Salamandroidea are a suborder of salamanders, referred to as advanced salamanders. The members of the suborder are found worldwide except for Antarctica, sub-Saharan Africa, and Oceania. They differ from suborder Cryptobranchoidea as the angular and prearticular bones in their lower jaws are fused, their trunk ribs are bicapitate, and all members use internal fertilization. The female is fertilized by means of a spermatophore, a sperm-containing cap placed by the male in her cloaca. The sperm is stored in spermathecae on the roof of the cloaca until it is needed at the time of oviposition.
The earliest known salamandroid fossils remain contested. Some studies suggest that the earliest salamandroids are represented by specimens of the species Beiyanerpeton jianpingensis and Qinglongtriton gangouensis from the Tiaojishan Formation of Inner Mongolia, China, dated to the Late Jurassic about 157 million years ago. Alternative analyses suggest that Beiyanerpeton jianpingensis and Qinglongtriton gangouensis are stem salamanders and that the oldest known certain salamandroid is Valdotriton gracilis from the Early Cretaceous of Spain, about 127 Ma. (Full article...)
List of selected salamander articles
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Did you know? –
- ... that although it was first classified as a reptile, the extinct genus Batropetes (restoration pictured) is now known to be a microsaur amphibian?
- ... that golden frogs are often mistaken for bronzed frogs?
- ... that the largest amphibian in the world is the Chinese giant salamander (pictured)?
- ...that the Eastern Newt changes its skin colour from bright red to olive green when it becomes an adult, and is known as the red eft before adulthood?
- ... that the East Bay Vivarium is the oldest and largest retail vivarium in the United States?
Selected amphibian type
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela from the group Caudata. Urodela is a scientific Latin term based on the Ancient Greek οὐρά δήλη: ourà dēlē "conspicuous tail". Caudata is the Latin for "tailed ones", from cauda: "tail".
Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders never have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults.
This group of amphibians is capable of regenerating lost limbs as well as other damaged parts of their bodies. Researchers hope to reverse engineer the regenerative processes for potential human medical applications, such as brain and spinal cord injury treatment or preventing harmful scarring during heart surgery recovery. The remarkable ability of salamanders to regenerate is not just limited to limbs but extends to vital organs such as the heart, jaw, and parts of the spinal cord, showing their uniqueness compared to different types of vertebrates. This ability is most remarkable for occurring without any type of scarring. This has made salamanders an invaluable model organism in scientific research aimed at understanding and achieving regenerative processes for medical advancements in human and animal biology. (Full article...)
Selected images
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Photograph credit: Charles J. SharpMantella baroni, commonly known as Baron's mantella, variegated golden frog or Madagascar poison frog, is a species of frog in the family Mantellidae. It is native to Madagascar and is seen here in Ranomafana National Park in the southeastern part of the country. The species was described in 1888 by George Albert Boulenger, who named it after its collector, Richard Baron. It has been classified as a least-concern species by the International Union for Conservation of Nature due to its relatively wide distribution, but is threatened by habitat loss. The frog's diet includes mites, which cause it to accumulate a high alkaloid concentration in its skin, making it toxic. The bright colours may serve as a warning sign to potential predators of the frog's toxicity. -
Photograph: JJ HarrisonThe southern brown tree frog (Litoria ewingii) is a species of tree frog native to Australia found in a wide range of habitats. Reaching 45 millimetres (1.8 in), this species is generally brown, but green and green-striped morphs are also recorded. -
Photo credit: Carey James BalboaThe Red-eyed Tree Frog (Agalychnis callidryas) is a small-sized tree frog, reaching lengths of about 5 to 7 centimeters (3 inches), native to Neotropical rainforests in Central America. They are not poisonous and rely on camouflage to protect them. During the day, they remain motionless with their colorful parts hidden. Thus, they appear almost completely green, and well hidden among the foliage. -
Photograph credit: Charles James SharpThe marsh frog (Pelophylax ridibundus) is a species of amphibian native to Europe and parts of Asia. It is the largest type of frog in most of its range, growing to a snout-to-vent length of around 100 mm (4 in); tadpoles can reach up to 190 mm (7.5 in) in length, but this usually occurs in places with long winters where the tadpole has time to grow. Marsh frogs hibernate during the winter, either underwater or in burrows, and are able to use the Earth's magnetic field to locate breeding ponds. This marsh frog was photographed in Kampinos National Park, Poland. -
Photo credit: Patrick CoinA terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin. -
Illustration credit: Ernst HaeckelA frog is an amphibian characterized by long hind legs, a short body, webbed digits, protruding eyes and the absence of a tail. Frogs are most noticeable through their call, which can be widely heard during the mating season. It is estimated that up to 20% of amphibian species may care for their young in one way or another, and there is a great diversity of parental behaviours. For example, frogs in the Gastrotheca genus (upper left) carry their eggs in a pouch, and females of the Eleutherodactylus lineatus species (center left) carry their young on their back. -
Common toads in amplexusPhoto: Bernie KohlTwo common toads (Bufo bufo) in amplexus, a form of pseudocopulation found in amphibians and limulids. A male common toad grasps a female with his front legs as part of the mating process. Common toads stay in amplexus for several days. As the female lays a long, double string of small black eggs, the male fertilises them with his sperm; the gelatinous egg strings, which may contain 3000 to 6000 eggs and be 3 to 4.5 metres (10 to 15 ft) in length, are later tangled in plant stalks. -
Photograph credit: Petar MiloševićThe agile frog (Rana dalmatina) is a species of true frog in the family Ranidae. Native to central and southern Europe, its brownish dappled colouring helps to conceal it among the leaf litter on the forest floor. This frog, about 5 cm (2.0 in) long, was photographed in the Golovec Forest in Slovenia. -
The edible frog (Pelophylax kl. esculentus) is a fertile hybrid of the pool and marsh frogs which is commonly found in Europe. The species is commonly used in food, including the French delicacy frog legs. -
The fire salamander (Salamandra salamandra) is a European salamander species with a long lifespan. These nocturnal animals generally eat various insects, spiders, earthworms and slugs, but they also occasionally eat newts and young frogs. -
Photo: QuartlDendrobates azureus, commonly known as the blue poison dart frog or okopipi, is a poison dart frog found in the forests surrounded by the Sipaliwini Savanna in southern Suriname. Authorities have recently treated it as a variant of Dendrobates tinctorius rather than a distinct species as before. -
Photo credit: FroggydarbA specimen from the "hybrid zone" of the Leaf Green Tree Frog (Litoria phyllochroa) and the Southern Leaf Green Tree Frog (L. nudidigitus), showing physical characteristics of both species. These small stream-dwelling frogs (averaging only 40mm in length), are native to eastern Australia and occur together south of Sydney. The two species are differentiated only by distribution, call and slight differences in flank markings. -
Photograph: Brian GratwickeRabbs' fringe-limbed treefrog (Ecnomiohyla rabborum) is a large species of frog originally found in the forest canopies of central Panama. Only discovered in 2005, the species is thought to be extinct in the wild; only one specimen, a male at the Atlanta Botanical Garden, survives. -
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name. -
Photo: Benny TrappAn Italian tree frog (Hyla intermedia) with an inflated vocal sac. Vocal sacs, flexible membranes of skin possessed by most male frogs, are generally used to amplify mating or advertisement calls. However, some species in the family Rhinodermatidae may also use the sac to store or transport their tadpoles.
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Selected toad article
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Nectophryne, or African tree toads, is a small genus of true toads with only two species. They are native to West and Central Africa: Nigeria, Cameroon, Gabon, northeastern Congo, Bioko and Equatorial Guinea. Nectophryne afra uses small bodies of water to lay its eggs which are then guarded by the male. (Full article...)
Selected caecilian article
Nectocaecilia petersii is a species of semiaquatic amphibian in the family Typhlonectidae. It belongs to the monotypic genus Nectocaecilia. This species is found in Venezuela and Brazil, and possibly Colombia and Peru. Its natural habitats are subtropical or tropical moist lowland forests and rivers, where it tends to live by the banks of streams. It lives in eel burrows and the roots of floating vegetation. It is threatened by habitat loss. This species has a number of distinct traits that illustrate that it is a burrowing vertebrate, including its very small eyes and long, thin body. (Full article...)
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